Rosaceae

Argentina Hill.

Last edited by Pieter B. Pelser, 4 August 2019

Photos 
  1. Argentina papuana (Focke) Soják, Thaiszia 20 (2010) 95; --Potentilla papuana Focke, Abh. Naturw. Ver. Bremen 13 (1895) 162; --Kalkm., Fl. Males. ser. 1, 11 (1993) 294; --Type: from New Guinea. New Guinea, Sulawesi, Philippines. LUZON: Benguet. Alpine or subalpine grasslands.
  2. Argentina parvula (Stapf) Soják, Thaiszia 20 (2010) 95; --Potentilla parvula Staff, Hook. Icon. Plant. 4, 3 (1894) pl.2294; --Kalkm., Fl. Males. ser. 1, 11 (1993) 294. --Argentina philippinensis (Merr.) Soják, Thaiszia 20 (2010) 95; --Potentilla philippinensis Merr., PJS 29 (1926) 480; --Type: Clemens 5006 (UC, holo), Luzon: Benguet, Mt Pulag, c. 2800m, seen in only one place on an old trail in the summit grasslands, associated with Schoenus (Cyperaceae), 24 to 27-Feb-1925. Borneo, New Guinea, Sulawesi, Philippines. LUZON: Benguet. Subalpine grassland, c. 2800m.

Eriobotrya Lindl. = Rhaphiolepis

Photinia Lindl.

Last edited by Pieter B. Pelser, 25 November 2019

Photos
  1. Photinia serratifolia (Desf.) Kalkm., Blumea 21 (1973) 424; Fl. Males. ser. 1, 11 (1993) 315; --Crataegus serratifolia Desf., Tabl. École Bot., ed. 3 (Cat. Pl. Horti Paris.) (1829) 408. --Photinia serrulata Lindl., Trans. Linn. Soc. 13 (1821) 103; --Merr., EPFP 2 (1923) 226. India, China, Japan, Taiwan, Philippines, ?Sumatra. LUZON, MINDANAO. Mossy forests, c. 2500m. Photos

Potentilla L.

Last edited by Pieter B. Pelser, 4 August 2019

Photos
  1. Potentilla indica (Andrews) Th.Wolf in Asch. & Graebn., Syn. Mitt.-Eur. Fl. 6, 1 (1904) 661; --Kalkm., Blumea 16 (1968) 344; Fl. Males. ser. 1, 11 (1993) 291; --Duchesnea indica (Andrews.) Focke in Engl. & Prantl, Nat. Pflanzenfam. 3, 3 (1888) 33; --Fragaria indica Andrews, Bot. Repos. 7 (1807) t. 479. Afghanistan, Pakistan, India, Sri Lanka, China, Japan, Java, Korea, Laos, Myanmar, Nepal, Taiwan, Thailand, Tibet, Vietnam, Lesser Sunda Isls, Philippines. Mostly in disturbed habitats: roadside, plantations, rarely in forest, rather often in damp and waste places, 700-2300m. Photos

Prunus L.

Last edited by Pieter B. Pelser, 1 June 2018

Photos
  1. Prunus arborea (Blume) Kalkm., Blumea 13 (1965) 90; Fl. Males. ser. 1, 11 (1993) 329; --Polydontia arborea Blume, Bijdr. Fl. Ned. Ind. 17 (1826) 1105. Photos
  2. Prunus clementis (Merr.) Kalkm., Blumea 13 (1965) 70; Fl. Males. ser. 1, 11 (1993) 334; --Pygeum clementis Merr., PJS 3 c (1908) Bot. 227; EPFP 2 (1923) 232; --Type: Clemens 760 (lecto); Clemens 990; both Mindanao. --Pygeum apoense Elmer, LPB 5 (1913) 1623; --Merr., EPFP 2 (1923) 232; --Type: Elmer 11729, Mindanao. Philippines, Sulawesi. MINDANAO. Primary and secondary forests.
  3. Prunus fragrans (Elmer) Kalkm., Blumea 13 (1965) 74; Fl. Males. ser. 1, 11 (1993) 335; --Pygeum fragrans Elmer, LPB 2 (1908) 475; --Merr., EPFP 2 (1923) 232; --Type: Elmer 7504 (lecto), Luzon. --Pygeum coccineum (Elmer) Elmer, LPB 5 (1913) 1621; --Merr., EPFP 2 (1923) 232; --Parinarium coccineum Elmer; --Type: Elmer 9787, Negros. --Pygeum megaphyllum Merr. ex Elmer, LPB 5 (1913) 1621; --Merr., EPFP 2 (1923) 233; --Type: BS 14923 Ramos, Luzon. Philippines, ?Taiwan, Philippines. CAMIGUIN DE BABUYANES, LUZON, NEGROS, PANAY. Primary and secondary forests, (150-)600-2100(-2500)m. Photos
  4. Prunus grisea (Blume) Kalkm., Blumea 13 (1965) 58; Fl. Males. ser. 1, 11 (1993) 338; --Pygeum griseum Blume, Mél. Bot. nr. 2 (1855) 11; --Muell. in Walp. Ann. 4 (1857) 642; --Type: from Java. Photos
  5. Prunus javanica (Teijsm. & Binn.) Miq., Fl. Ind. Bat. 1, 1 (1855) 365; --Kalkm., Blumea 13 (1965) 47; Fl. Males. ser. 1, 11 (1993) 328; --Cerasus javanica Teijsm. & Binn., Natuurk. Tijdschr. Ned. Indië 2 (1851) 309; --Type: from Java. --Prunus junghuhnianus Miq., Pl. Jungh. (1855) 402; --Merr., EPFP 2 (1923) 234; --Type: from Java. Andaman Isls, Myanmar, Thailand, Vietnam, Sumatra, Peninsular Malaysia, Java, Lesser Sunda Isls, Borneo, Sulawesi, Moluccas, New Guinea, Philippines. PALAWAN. Photos
  6. Prunus kinabaluensis Kalkm., Blumea 13 (1965) 64; Fl. Males. ser. 1, 11 (1993) 339; --Type: from Sabah. Borneo (Sabah, Ranau District), Philippines. LUZON. Forests, 1300-2400m.
  7. Prunus marsupialis Kalkm., Blumea 13 (1965) 71; Fl. Males. ser. 1, 11 (1993) 341l; --Pygeum glandulosum Merr., PJS 3 c (1908) Bot. 226; EPFP 2 (1923) 233; --Type: R.S. Williams 642 (lecto); 25 syntypes, all Philippines. --Pygeum pubescens Merr., PJS 9 c (1914) Bot. 359; EPFP 2 (1923) 234; --Type: Wenzel 333 (lecto), Philippines. Both the epithet glandulosum and pubescens are not available in Prunus. Endemic to the Philippines. LUZON, MINDORO, PANAY, NEGROS, BOHOL, LEYTE, SAMAR. Forests and forest edges, 0-c. 1100m. Photos
  8. Prunus pulgarensis (Elmer) Kalkm., Blumea 13 (1965) 67; Fl. Males. ser. 1, 11 (1993) 344; --Pygeum pulgarense Elmer, LPB 5 (1913) 1626; --Merr., EPFP 2 (1923) 234; --Type: Elmer 13200, Palawan. --Pygeum monticolum Merr., PJS 10 c (1915) Bot. 312; EPFP 2 (1923) 233; --Type: Whitford 1203, Luzon. Endemic to the Philippines. LUZON, PALAWAN. Montane forests, c. 1000-1200m. Note by L.L.Co: Known only from four collections, no new material of this species has been collected since 1965. Conservation status: Vulnerable (DAO 2017-11).
  9. Prunus rubiginosa (Elmer) Kalkm., Blumea 13 (1965) 72; Fl. Males. ser. 1, 11 (1993) 345; --Pygeum rubiginosum Elmer, LPB 5 (1913) 1624; --Merr., EPFP 2 (1923) 234; --Type: Elmer 11857 (holo); 14067, both Mindanao. --Pygeum elmerianum Koehne in Engl., Bot. Jahrb. 51 (1913) 206; --Merr., EPFP 2 (1923) 232; --Type: Elmer 12210, Sibuyan. Endemic to the Philippines. LUZON, MINDORO, SIBUYAN, MINDANAO. Forests, 250-1200m. Note by L.L.Cp: Known only from 1 or 2 specimens in each island. Conservation status: Vulnerable (DAO 2017-11). Photos
  10. Prunus subglabra (Merr.) Kalkm., Blumea 13 (1965) 87; Fl. Males. ser. 1, 11 (1993) 347; --Pygeum subglabrum Merr., PJS 30 (1926) 395; --Type: BS 45014 Ramos & Edaño (UC, holo; L, iso), Luzon: Benguet, Mt Pulag, mossy forest, c. 2700m, 27-Feb-1925. Endemic to the Philippines. LUZON: Benguet (Mt Pulag; Mt Tabayoc). Mossy forests, 2400-2700m. Conservation status: Vulnerable (DAO 2017-11).

 Possibly present:

  1. Prunus spicata Kalkm., Blumea 13 (1965) 69; Fl. Males. ser. 1, 11 (1993) 346; --Type: from Sabah. ?LUZON, ?MINDORO. A doubtful sterile specimen seen by Kalkman from Mindoro.

Incertae sedis:

  1. Prunus C. --Kalkm., Fl. Males. ser. 1, 11 (1993) 350. --Prunus luzoniensis Merr. & Quisumb. in sched., based on PNH 4337 Celestino. Endemic to the Philippines. LUZON: Benguet (Mt Pulag). Partly open field, close to stream, 2000m. Up to now the only Malesian specimen belonging to subgenus Padus.
  2. Prunus ramiflorum Merr., PJS 9 c (1914) Bot. 447; EPFP 2 (1923) 234; --Type: Vanoverbergh 2820, Luzon: Mountain Province, Bauco, in forests, c. 1500m, 4-Apr-1913. LUZON: Mountain Province. Forests, 1400m. Not treated in Kalkman’s Flora Malesiana revision. But according to Merrill (1914), “a characteristic species, recognizable by its eglandular leaves which are comparatively small and few-nerved, and its rather congested, paniculate, short inflorescences which are borne on the larger branches below the leaves, not on the ultimate branchlets”

Cultivated, not naturalized:

  1. Prunus armeniaca L., Sp. Pl. (1753) 474. LUZON. Occasionally cultivated in the Cordillera Highlands of N Luzon. Apricot. Photos
  2. Prunus persica (L.) Batsch., Beitr. Entw. Pragm. Gesch. 1 (1801) 30; --Amygdalus persica L., Sp. Pl. 1 (1753) 472. China. Cultivation spread from there, now cultivated wherever the climate is suitable; in Malesia rarely cultivated in montane areas; in the Philippines, grown to a limited scale in the Cordillera highlands. Peach. Photos

 Pygeum Gaertn. = Prunus

Pyracantha M.Roem.

Last edited by Pieter B. Pelser, 27 February 2021

Photos
  1. Pyracantha crenulata (D.Don) M.Roem., Fam. Nat. Syn. Monogr. 3 (Rosifl.) (1847) 220; --Feng et al., Phytotaxa 478 (2021) 244; --Mespilus crenulata D.Don, Prodr. Fl. Nepal. (1825) 238. China, India, Myanmar, Nepal, Pakistan, Tibet, Philippines. Note by P.B. Pelser (27-Feb-21): This species is listed for the Philippines by Feng et al. (2021), but no specimens were cited.

Cultivated, not naturalized:

  1. Pyracantha angustifolia (Franch.) C.K.Schneid., Ill. Handb. Laubholzk. 1 (1906) 761; --Cotoneaster angustifolius Franch., Pl. Delav. (1889) 221; --Type: from China. China. LUZON. Known in Luzon from the Cordillera Highlands, Baguio-Bontoc Road, based on PNH 20368 Mendoza and Steenis 17923, collected from the same locality and date, apparently from cultivated plants, as well as PNH 39575 Steiner, probably collected in the same general locality. According to Kalkman, these specimens represent a glabrous form of Pyracantha angustifolia which is wild in Yunnan and is sometimes planted as an ornamental in Europe and N America

Rhaphiolepis Lindl.

Last edited by Pieter B. Pelser, 5 August 2020

Photos
  1. Rhaphiolepis bibas (Lour.) Galasso & Banfi, Ital. Botanist 9 (2020) 66; --Liu et al., PhytoKeys 154 (2020) 26; --Crataegus bibas Lour., Fl. Cochinch. 1 (1790) 319. --Eriobotrya japonica (Thunb.) Lindl., Trans. Linn. Soc. 13 (1822) 102; --Mespilus japonicus Thunb., Fl. Japon. (1784) 206; --Type: from Japan. China, Japan. LUZON: Mountain Province. China, there and in Japan cultivated for many years. Now throughout the tropics and subtropics cultivated as a fruit tree. In the Philippines, grown in Mountain Province, particularly common in the vicinity of Sagada, 1400-1500m. Loquat. Not naturalized? Photos
  2. Rhaphiolepis oblongifolia (Merr. & Rolfe) B.B.Liu & J.Wen, Front. Plant Sci. 10-1731 (2020) 11; --Liu et al., PhytoKeys 154 (2020) 41; --Eriobotrya oblongifolia Merr. & Rolfe, PJS 3 c (1908) Bot. 102; --Merr., EPFP 2 (1923) 226; --Type: FB 4680 Mearns & Hutchinson (NY, lecto; US, isolecto), Mindanao: Misamis, Mt Malindang, May-1906. Endemic to the Philippines. MINDANAO: Misamis Occidental.
  3. Rhaphiolepis philippinensis (S.Vidal) Kalkm., Blumea 21 (1973) 434; --Liu et al., PhytoKeys 154 (2020) 42; Fl. Males. ser. 1, 11 (1993) 318; --Pyrus philippinensis (S.Vidal) M.F.Fay & Christenh., Global Fl. 4 (2018) 115; --Eriobotrya philippinensis S.Vidal, Rev. Pl. Vasc. Filip. (1886) 123; --Merr., EPFP 2 (1923) 226; --Type: Vidal 1350 (K, lecto; MA, isolecto), Luzon: Zambales, Santa Cruz. --Eriobotrya luzonensis (Merr.) Nakai, J. Arnold Arb. 5 (1924) 69; --Merr., EPFP 4 (1925) 242; --Photinia luzonensis Merr., Govt. Lab. Publ. (Philipp.) 17 (1904) 18; EPFP 2 (1923) 226; --Type: Merrill 3223 (NY, lecto; K, P), Luzon: Bataan, Mt Mariveles, Oct-1903. --Eriobotrya acuminatissima Nakai, J. Arnold Arb. 5 (1924) 71; --Merr., EPFP 4 (1925) 242; --Type: BS 35622 Martelino & Edaño (A, lecto; BM, L, P, isolecto), Panay: Capiz, Mt Salibongbong, Jun-1919. Borneo, Philippines. LUZON: Bataan, Zambales, PANAY. In forests, 300-2600m. Also on ultramafics near sea level. Photos

Rosa L.

Last edited by Pieter B. Pelser, 18 August 2017

Photos
  1. Rosa luciae Franch. & Rochebr. ex Crépin, Bull. Soc. Roy. Bot. Belg. 10 (1871) 324; --Kalkm., Blumea 21 (1973) 284; Fl. Males. ser. 1, 11 (199) 305; --Type: from Japan. --Rosa philippinensis Merr., PJS 17 (1920) 260, incl. var. depauperata Merr.; EPFP 2 (1923) 231; --Type: Elmer 5794. Japan, Korea, Ryukyu Isls, E China, Taiwan, N Philippines. LUZON. Thickets, 1200-1700m. Conservation status: Other Threatened Species (DAO 2017-11). Photos
  2. Rosa transmorrisonensis Hayata, Icon. Pl. Formos. 3 (1913) 97; --Kalkm., Blumea 21 (1973) 284; Fl. Males. Ser. 1, 11 (199) 305; --Type: from Taiwan. --Rosa luzoniensis Merr., PJS 17 (1920) 259, including var. parvifolia Merr.; --Type: BS 31876 (syn); Mearns 4300 (syn); BS 8336 McGregor (syn), Luzon: Benguet, Pauai. Taiwan, N Philippines. LUZON: Mountain Province, Benguet. Thickets, c. 1200m and higher. Conservation status: Other Threatened Species (DAO 2017-11).

Rubus L.

Last edited by Pieter B. Pelser, 21 November 2021

Photos
  1. Rubus benguetensis Elmer, LPB 1 (1908) 296; LPB 2 (1908) 449; --Focke, Bibl. Bot. 17, 72 (1910) 66; --Merr., EPFP 2 (1923) 227; --Kalkm., Blumea 29 (1984) 337; Fl. Males. ser. 1, 11 (1992) 268; --Type: Elmer 8383 (K, L, MO, NY, US, iso), Luzon: Benguet, Baguio, in rock crevices of limestone outcroppings in the pine region at 4500ft, Mar-1907. Borneo, Philippines, Sulawesi. LUZON: Benguet, Isabela. Montane thickets, on limestone at c. 1300m in Benguet. Photos
  2. Rubus clementis Merr., PJS 3 c (1908) 139; --Kalkm., Blumea 32 (1987) 325; Fl. Males. ser. 1, 11 (1992) 253; --Rubus lucens Focke ssp. clementis (Merr.) Focke, Bibl. Bot. 72 (1911) 213; --Merr., EPFP 2 (1923) 228; --Type: M.S. Clemens 740 (PNH, holo, lost; K, US and as Clemens s.n. in G, iso), Mindanao: Lanao del Sur, Camp Keithley. Sumatra, Borneo, Sulawesi, Philippines. MINDANAO: Lanao del Sur, Bukidnon.
  3. Rubus copelandii Merr., PJS 1 Suppl. 3 (1906) 194, as ‘copelandi’; EPFP 2 (1923) 227; --Zandee & Kalkm., Blumea 27 (1981) 98; --Kalkm., Fl. Males. ser. 1, 11 (1992) 259; --Type: Merrill 4810 (NY, US, iso), Luzon: Benguet, Pauai. Endemic to the Philippines. LUZON: Mountain Province, Benguet. More or less open spots in mossy forest, forest edges and thickets, 1700-2450m. Photos
  4. Rubus cumingii Kuntze, Meth. Speciesbeschr. (1879) 72; --Focke, Bibl. Bot. 17, 72 (1910) 58; --Merr., EPFP 2 (1923) 227; --Kalkm., Blumea 29 (1984) 332; Fl. Males. ser. 1, 11 (1992) 271; --Type: Cuming s.n. (B, holo), Philippines. Endemic to the Philippines. Known only from the type which has no indicated definite locality on the label; according to Focke, this comes from Luzon.
  5. Rubus edanoi Merr., PJS 13 c (1918) Bot. 14; EPFP 2 (1923) 227, as ‘edanoii’; --Type: BS 26505 Ramos & Edaño (US, iso), Luzon: Nueva Ecija, Mt Umingan. Mossy forest, c. 300m, 6-Sep-1916. Endemic to the Philippines. LUZON: Nueva Ecija. Known only from the type. According to Merrill (1918), this is allied to Rubus cumingii Kuntze, from which it is distinguished at once, among other characters, by its sharply acuminate leaves.
  6. Rubus ellipticus Sm. in Rees, Cyclop. (1815) 30; --Merr., EPFP 2 (1923) 227; --Zandee & Kalkm., Blumea 27 (1981) 109; --Kalkm., Fl. Males. ser. 1, 11 (1992) 260; --Type: from Nepal. Sri Lanka, India, China, Nepal, Vietnam, Philippines. LUZON: Benguet. Montane oak forest, pine forest and in secondary growth, 1000-2400m. Photos
  7. Rubus fraxinifolius Poir. in Lam., Encycl. Méth. 6 (1806) 242; --Merr., EPFP 2 (1923) 227; --Zandee & Kalkm., Blumea 27 (1981) 87; --Kalkm., Fl. Males. ser. 1, 11 (1992) 261; --Type: from Java. --Rubus fraxinifolius Poir. var. haightii Elmer, LPB 2 (1908) 461; --Type: a Mearns specimen from Pauai (Mr. Haight’s Place), Luzon. This may be BS 4459 Mearns, cited by Merrill under Rubus merrillii. --Rubus merrillii Focke, Bibl. Bot. 72 (1911) 153; --Merr., EPFP 2 (1923) 228; --Type: BS 862 Merrill or BS  6637, Luzon: Benguet, Pauai. Taiwan, Java, Lesser Sunda Isls, Borneo, Philippines, Sulawesi, Moluccas, New Guinea, Solomon Isls, Bismarck Arch. LUZON, MINDANAO. N Luzon to Mindanao. More or less open sites such as forest edges and gaps, along trails, streambanks, abandoned cultivations, etc, 0-2500m. Photos
  8. Rubus heterosepalus Merr., PJS 20 (1922) 387; EPFP 2 (1923) 228; --Kalkm., Blumea 29 (1984) 344; Fl. Males. ser. 1, 11 (1992) 273; --Type: BS 37609 Ramos & Edaño (PNH, holo, ?lost; A, K, L, US, iso), Luzon: Mountain Province, Mt  Polis, mossy forest, 1800m, Feb-1920. Endemic to the Philippines. LUZON: Mountain Province, Ifugao. Mossy forest and associated thickets, 1800-1900m. Conservation status: Other Threatened Species (DAO 2017-11).
  9. Rubus luzoniensis Merr., PJS 1 Suppl. 1 (1906) 195; --Elmer, LPB 2 (1908) 452; --Focke, Bibl. Bot. 17, 72 (1910) 59; --Merr., EPFP 2 (1923) 228; --Kalkm., Blumea 29 (1984) 328; Fl. Males. ser. 1, 11 (1992) 274; --Type: Merrill 4596 (PNH, holo, lost; BO, K, L, NY, iso), Luzon: Mountain Province, Mt Data, Nov-1905. Endemic to the Philippines. LUZON: Ifugao, Mountain Province, Benguet. Mossy forest and associated clearings, c. 2000- 2500m. Photos
  10. Rubus mearnsii Elmer, LPB 2 (1908) 448; --Merr., PJS 5 c (1910) Bot. 353; EPFP 2 (1923) 228; --Focke, Bibl. Bot. 17, 72 (1910) 74; --Kalkm., Blumea 29 (1984) 341; Fl. Males. ser. 1, 11 (1992) 275; --Type: BS 4304 Mearns (?lost), Luzon: Benguet, Pauai near Mr Haights’ place, Jul-1907. Endemic to the Philippines. LUZON: Mountain Province, Benguet. Reported from mossy forest, but seen along roadside clearings, c. 2200-2400m. Kalkman (1984) hints at some possibility that this taxon might be conspecific with Rubus benguetensis Elmer.  
  11. Rubus moluccanus L., Sp. Pl. (1753) 1197; --Kalkm., Blumea 29 (1984) 346; Fl. Males. ser. 1, 11 (1992) 275; --Type: Rumph. Herb. Amboin. 5 (1747) 88, pl. 47, Fig. 2 (lecto).
  12. Rubus niveus Thunb., Diss. Rubo (1813) 9; --Merr., EPFP 2 (1923) 229; --Zandee & Kalkm., Blumea 27 (1981) 107; --Kalkm., Fl. Males. ser. 1, 11 (1992) 264; --Type: from Java. Sri Lanka, India, Vietnam, Taiwan, Sumatra, Java, Lesser Sunda Isls, Philippines, Sulawesi.
  13. Rubus pectinellus Maxim., Mél. Biol. Acad. St. Pétersb. 8 (1872) 324 (Bull. Acad. St. Pétersb. 17); --Merr., EPFP 2 (1923) 229; --Zandee & Kalkm., Blumea 27 (1981) 78; --Kalkm., Fl. Males. ser. 1, 11 (1992) 284; --Type: from Japan. China, Taiwan, Japan, Philippines. LUZON: Ilocos Norte, Abra, Mountain Province, Ifugao, Benguet, Quezon, MINDORO, MINDANAO: Bukidnon, Davao. Primary and secondary forests, also in mossy forest, c. 750-2750m. Photos
  14. Rubus perfulvus Merr., PJS 20 (1922) 386; EPFP 2 (1923) 229; --Kalkm., Blumea 29 (1984) 374; Fl. Males. ser. 1, 11 (1992) 280; --Type: BS 38566 Ramos & Edaño, Mindanao: Bukidnon, Mt Lipa. Endemic to the Philippines. MINDANAO: Bukidnon. Only known from the type collection (Mt Lipa).
  15. Rubus pirifolius Sm., Pl. Icon. Hact. Ined. 3 (1791) t. 61; --Focke, Bibl. Bot. 17, 72 (1910) 64; --Merr., EPFP 2 (1923) 229; --Backer & Bakh.f., Fl. Java 1 (1964) 516; --Kalkm., Blumea 29 (1984) 333; Fl. Males. ser. 1, 11 (1992) 280; --Type: from Java. --Rubus philippinensis Focke in Elmer & Focke, LPB 5 (1913) 1617; --Focke, Bibl. Bot. 17, 83 (1914) 28; --Merr., EPFP 2 (1923) 229; --Type: Elmer 13606 (PNH, holo, lost; A, BISH, BM, BO, GH, K, L, NY, P, U, US, iso), Mindanao: Agusan del Norte, Cabadbaran, Mt Urdaneta, moist stony soil of dense woods at 3750ft or between Duros & Cawilanan peaks, Aug-1912. --Rubus brevipetalus Elmer, LPB 2 (1908) 450; --Type: Elmer 10099 (PNH, holo, lost; A, BISH, BM, BO, K, L, NY, US, iso), Negros: Negros Oriental, Cuernos Mtns, in humid woods at 4000ft, May-1908. Thailand, Laos, Vietnam, China, Sumatra, Java, Lesser Sunda Isls, Borneo, Philippines, Sulawesi. NEGROS, LEYTE, MINDANAO: Zamboanga, Agusan del Norte, Davao. 900-1200m, forest and forest edges. Photos
  16. Rubus rolfei S.Vidal, Phan. Cuming. (1885) 171; --Merr., EPFP 2 (1923) 281; --Kalkm., Blumea 29 (1984) 371; Fl. Males. ser. 1, 11 (1992) 281; --Type: Cuming 808 (lecto); Vidal 294, both from Luzon. --Rubus elmeri Focke, Bibl. Bot. 72 (1910) 112; --Merr., EPFP 2 (1923) 227; --Type: Merrill 4651, BS 4305 Mearns, Luzon. Taiwan, Philippines. LUZON, MINDORO, NEGROS. Forests and more open places like clearings at higher elevation, (900-)1500-2700m. Photos
  17. Rubus rosifolius Sm., Pl. Icon. Hact. Ined. 3 (1791) t. 60, as 'rosaefolius'; --Merr., EPFP 2 (1923) 230; --Zandee & Kalkm., Blumea 27 (1981) 81; --Kalkm., Fl. Males. ser. 1, 11 (1992) 265; --Type: from Mauritius. --Rubus tagallus Cham. & Schltdl., Linnaea 2 (1827) 9; --Merr., EPFP 2 (1923) 230; --Type: Chamisso s.n. (LE, holo; B, iso), Luzon. --Rubus comintanus Blanco, Fl. Filip. ed. 2 (1845) 298; --Type: Merrill Sp. Blancoanae 721 (L, neo), Luzon: Rizal, Montalban, Dec-1914. --Rubus apoensis Elmer, LPB 5 (1913) 1618; --Type: Elmer 10464 (L, holo; NY, US, iso), Mindanao: Davao, Mt Apo, Todaya, 4000ft, May-1909. India, Cambodia, Vietnam, ?Japan, Taiwan, Philippines, Borneo, Sulawesi, Java, Lesser Sunda Isls, New Guinea, New Caledonia, Australia. LUZON: Ilocos Norte, Mountain Province, Benguet, Ifugao, Aurora, Pampanga, Bataan, Bulacan, Rizal, Laguna, Quezon, Albay, Sorsogon, MINDORO, NEGROS, MINDANAO: Misamis Occidental, Bukidnon, Davao. In open secondary places like clearings, forest edges, roadsides, landslides, grasslands, riverbanks, fallow gardens, also in shrubberies and rarely in the undergrowth of the lighter types of forest, from sea-level up to 2000m, rarely higher. Photos
  18. Rubus sorsogonensis Elmer, LPB 10 (1939) 3777; --Kalkm., Blumea 29 (1984) 339; Fl. Males. ser. 1, 11 (1992) 282; --Type: Elmer 14607 (A, BISH, BM, BO, K, L, MO, NY, P, U, iso), Luzon: Sorsogon, Irosin, Mt Bulusan, climbing upon trees and sprawling over their tops in wet earth of deep woods at 1500ft, Oct-1915. Endemic to the Philippines. LUZON: Sorsogon. Only known from the type on Mt Bulusan. 450m.

 Incertae sedis:

  1. Rubus mindanaensis Focke, Ann. Cons. Jard. Bot. Genève 20 (1917) 104. Endemic to the Philippines. LUZON. Considered by Focke to be related to Rubus niveus which, according to the material seen, is in the Philippines restricted to Luzon; --Type: Warburg 14483 (Kalkman 1992).
  2. Rubus vanoverberghii Merr., PJS 9 c (1914) Bot. 448; EPFP 2 (1923) 230; --Type: Vanoverbergh 2683 (PNH, holo, lost; K, US, iso), Luzon: Mountain Province. Endemic to the Philippines. LUZON: Mountain Province. Possibly represents Rubus glomeratus Blume in the Philippines (Kalkman 1984).
  3. Rubus zambalensis Elmer, LPB 2 (1908) 451; --Merr., EPFP 2 (1923) 231; --Kalkm., Blumea 29 (1984) 385; --Type: FB 8085 Curran & Merritt, Luzon: Zambales. Endemic to the Philippines. LUZON: Zambales. The correct disposition of this name remains enigmatic. The type specimen obviously had only young flower buds and Elmer described them as placed in small glomerules and as subtended by a 4-lobed involucrum, which seems rather strange (Kalkman 1984).

Stranvaesia Lindl.

Last edited by Pieter B. Pelser, 5 August 2020

Photos
  1. Stranvaesia nussia (Buch.-Ham. ex D.Don) Decne., Nouv. Arch. Mus. Hist. Nat. Ser. I, 10 (1874) 178; --Li et al., J. Syst. Evol. (2019); --Photinia nussia (Buch.-Ham. ex D.Don) Kalkm., Blumea 21 (1973) 429; Fl. Males. ser. 1, 11 (1993) 314; --Pyrus nussia Buch.-Ham. ex D.Don, Prodr. Fl. Nepal (1825) 237. --Stranvaesia ambigua (Merr.) Nakai, J. Arn. Arb. 5 (1924) 72; --Merr., EPFP 4 (1925) 242; --Eriobotrya ambigua Merr., Gov’t. Lab. Publ. (Philipp.) 17 (1904) 18; EPFP 2 (1923) 226; --Type: FB 2796 Meyer, Luzon. India, Myanmar, Thailand, Laos, China, Philippines. In the Philippines up to 2500m. Photos

Cultivated, not naturalized:

  1. Fragaria vesca L., Sp. Pl. (1753) 494; --Kalkm., Fl. Males. ser. 1, 11 (1993) 285. Temperate Eurasia. LUZON: Benguet, Mountain Province. Not native to the Philippines; commercially grown in N Luzon.
  2. Malus baccata (L.) Borkh., Theor. Prakt. Handb. Forstbot. 2 (1803) 1280; --Hyland, Plant Inventory 169 (1967) 260; --Pyrus baccata L., Mant. Pl. (1767) 75. Continental Asia. Not native to the Philippines.
  3. Malus domestica Borkh., Handb. Forstbot. 2 (1803) 1272. W Asia. LUZON: Mountain Province, Benguet. Apple. Not native to the Philippines. Photos
  4. Pyrus pyrifolia (Burm.f.) Nakai, Bot. Mag. (Tokyo) 40 (1926) 564; --Hyland, Plant Inventory 169 (1967) 260; --Ficus pyrifolia Burm.f., Fl. Ind. (1768) 226. --Pyrus serotina Rehder, Proc. Amer. Acad. Arts l (1915) 231. Oriental pear.

Literature

DENR Administrative Order. 2017. Updated national list of threatened Philippine plants and their categories. DAO 2017-11

Feng, H-Z, S-J Wei, L-Y Wang, S-F Chen, Q Fan & W-B Liao. 2021. A taxonomic revision of the Pyracantha crenulata complex (Rosaceae, Maleae). Phytotaxa 478: 239-252.

Kalkman, C. 1965. The Old World species of Prunus subg. Laurocerasus including those formerly referred to Pygeum. Blumea 13: 1-115

Kalkman, C. 1968. Potentilla, Duchesnea, and Fragaria in Malesia (Rosaceae). Blumea 16: 325-354


Kalkman, C. 1973. The Malesian species of the subfamily Maloideae (Rosaceae). Blumea 21: 413-442

Kalkman, C. 1973. The genus Rosa in Malesia. Blumea 21: 281-291

Kalkman, C. 1984. The genus Rubus (Rosaceae) in Malesia. 2. The subgenus Malachobatus. Blumea 29: 319-386

Kalkman, C. 1987. The genus Rubus (Rosaceae) in Malesia. 3. The subgenus Micranthobatus. Blumea 32: 323-341


Kalkman, C. 1992. Rosaceae. Flora Malesiana ser. 1, 11: 227-351

Liu, BB, DY Hong, SL Zhou, C Xu, WP Dong, G Johnson & J Wen. 2019. Phylogenomic analyses of the Photinia complex support the recognition of a new genus Phippsiomeles and the resurrection of a redefined Stranvaesia in Maleae (Rosaceae). Journal of Systematics and Evolution 57: 678-694

Liu, BB, GN Liu, DY Hong & J Wen. 2020. Eriobotrya belongs to Rhaphiolepis (Maleae, Rosaceae): evidence from chloroplast genome and nuclear ribosomal DNA data. Frontiers in Plant Science 10: 1731

Liu, BB, YB Wang, DY Hong & J Wen. 2020. A synopsis of the expanded Rhaphiolepis (Maleae, Rosaceae). PhytoKeys 154: 19-55

Zandee, M & C Kalkman. 1981. The genus Rubus (Rosaceae) in Malesia. 1. Subgenera Chamaebatus and Idaeobatus. Blumea 27: 75-113

Zhao L, D Potter, Y Xu, PL Liu, G Johnson, ZY Chang & J Wen. 2018. Phylogeny and spatio-temporal diversification of Prunus subgenus Laurocerasus section Mesopygeum (Rosaceae) in the Malesian region. Journal of Systematics and Evolution 56: 637-651


Pelser, P.B., J.F. Barcelona & D.L. Nickrent (eds.). 2011 onwards. Co's Digital Flora of the Philippines. www.philippineplants.org

Copyright © 2011, Co's Digital Flora of the Philippines

Last updated 15 January 2022