Putranjivaceae

Drypetes Vahl

1. Drypetes bawanii (Merr.) Airy Shaw, Kew Bull. 38 (1983) 68; Alphab. Enum. Euph. Philipp. (1983) 21; ≡ Cyclostemon bawanii Merr., PJS 20 (1922) 397; Merrill, EPFP 2 (1923) 405; Type: FB 25357 Bawan (A, US, iso), Luzon: Quezon, Atimonan, 12-Apr-1916. Distribution: Endemic to the Philippines. LUZON: Quezon. Notes: Lowland forests, along streams and slopes, 100-300m. Native.
2. Drypetes congestiflora Chun & T.Chen, Acta Phytotax. Sin. 8 (1963) 275. ≠ Drypetes confertiflora auct. non (Hook.f.) Pax & K.Hoffm.; Merr. & Chun, Sunyatsenia 2 (1935) 259. Distribution: China, Philippines. Native.
3. Drypetes convoluta Airy Shaw, Kew Bull. 19 (1965) 307; Airy Shaw, Alphab. Enum. Euph. Philipp. (1983) 21; Type: BS 13810 Ramos (K, lecto; A, L, NY, US, iso), Luzon: Cagayan, Jan-1912. Distribution: Endemic to the Philippines. LUZON: Cagayan. Conservation status: Critically Endangered (IUCN 3.1). Native.
4. Drypetes cumingii (Baill.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 238; Airy Shaw, Alphab. Enum. Euph. Philipp. (1983) 21; ≡ Cyclostemon cumingii Baill., Étude Euphorb. (1858) 562, as 'cummingii'; Müll.Arg. in DC., Prodr. 152 (1866) 485; S.Vidal, Phan. Cuming. Philip. (1885) 142; Rev. Pl. Vasc. Filip. (1886) 238; Merrill, EPFP 2 (1923) 406; Type: Cuming 1695 (G-DC, lecto; BM, G, K, L, M, P, isolecto), Samar. = Cyclostemon iwahigensis Elmer, LPB 4 (1911) 1278; Merrill, EPFP 2 (1923) 406; Levin, Gard. Bull. Singapore 73 (2021) 379; Type: Elmer 12695 (NY, lecto; E, G, HBG, K, L, U, isolecto), Palawan: Palawan, Brookes Point, Addison Peak, Feb-1911. Distribution: China, Philippines. LUZON: Bataan, Batangas, Quezon, Sorsogon, Zambales, MINDANAO: Davao Oriental, MINDORO, PALAWAN, SAMAR, TAMLANGON. Notes: Synonym of Drypetes littoralis (C.B.Rob.) Merr.?. The description of the flowers and No. 13175 of Cyclostemon iwahigensis Elmer in Elmer (1911) do not belong to this species. Native.
5. Drypetes ellipsoidea (Merr.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 250; Romeroso et al., J. Asia-Pacific Biodiv. 17 (2024) 12; ≡ Cyclostemon ellipsoideus Merr., PJS 9c (1914) 369; Merrill, EPFP 2 (1923) 406; Type: Wenzel 828 (A, lecto; BM, GH, US, isolecto), Leyte: Buenavista near Jaro, in forests, c. 500m, 3-Jun-1914. Distribution: Endemic to the Philippines. ?HOMONHON, LEYTE, SAMAR. Notes: Synonym of Drypetes celebica (Boerl. & Koord.) Pax & K.Hoffm.? Conservation status: Endangered (IUCN 3.1). Native.
6. Drypetes falcata (Merr.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 250; Airy Shaw, Alphab. Enum. Euph. Philipp. (1983) 22; ≡ Cyclostemon falcatus Merr., PJS 3c (1908) 415; Merrill, EPFP 2 (1923) 406; Type: BS 4033 E. Fenix (K, lecto; US, isolecto), Camiguin de Babuyanes: along the seashore, 22-Jun-1907. Distribution: Endemic to the Philippines. BABUYAN ISLS, CAMIGUIN DE BABUYANES, LUZON: Cagayan. Notes: Along the seashore. DAO category: Other Threatened Species. Native.
7. Drypetes gitingensis (Elmer) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 250; Airy Shaw, Alphab. Enum. Euph. Philipp. (1983) 22; ≡ Cyclostemon gitingensis Elmer, LPB 3 (1910) 912; Merrill, EPFP 2 (1923) 406; Type: Elmer 12317 (NY, lecto; A, E, GH, HBG, K, L, MO, U, US, isolecto), Sibuyan: Magallanes (=Magdiwang), Mt Giting-giting, Apr-1910. Distribution: Endemic to the Philippines. SIBUYAN. Notes: Lowland forests along streams, c. 300m. Conservation status: Critically Endangered (IUCN 3.1). Native.
8. Drypetes globosa (Merr.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 247; Airy Shaw, Alphab. Enum. Euph. Philipp. (1983) 22; ≡ Cyclostemon globosus Merr., PJS 9c (1914) 478; Merrill, EPFP 2 (1923) 406; Type: BS 21362 Escritor (NY, lecto; F, K, L, SING, US, isolecto), Cuyo, 20-Jul-1913. Distribution: Endemic to the Philippines. CUYO, CUYO ARCHIPELAGO, LUZON: Sorsogon, MASBATE, MINDANAO: Zamboanga, MINDORO, NEGROS, PALAWAN. Notes: Lowland thickets and forests. Conservation status: Vulnerable (IUCN 3.1). Native.
9. Drypetes grandifolia (C.B.Rob.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 245; Airy Shaw, Alphab. Enum. Euph. Philipp. (1983) 22; ≡ Cyclostemon grandifolius C.B.Rob., PJS 3c (1908) 197; Merrill, EPFP 2 (1923) 406; Type: FB 6554 Hutchinson (US, lecto), Mindanao: Zamboanga, Lumbiag, 40m, 22-Feb-1907. = Drypetes megacarpa (Merr.) Pax & K.Hoffm., Bull. Jard. Bot. Buitenzorg, 3rd Sér. 6 (1924) 248; ≡ Cyclostemon megacarpus Merr., PJS 7c (1912) 387; Merrill, EPFP 2 (1923) 407; Type: BS 14953 (A, lecto), Luzon: Laguna, San Antonio, in forests, Jun-1912. = Cyclostemon ramiflorus Merr., PJS 7c (1912) 387; Type: FB 13640 Curran & Foxworthy (US, lecto), Negros: Negros Occidental, Mt Marapara, on forested ridges, c. 700m, Sep-1909. Distribution: Borneo, Philippines. BASILAN, CATANDUANES, LEYTE, LUZON: Aurora, Camarines, Isabela, Laguna, Nueva Ecija, Pangasinan, Quezon, Sorsogon, MINDANAO: Agusan del Norte, Cotabato, Zamboanga, Zamboanga del Sur, MINDORO, NEGROS, PANAY, SAMAR. Notes: In primary forests at low and medium elevation. Native.
10. Drypetes heptandra Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 254; Airy Shaw, Alphab. Enum. Euph. Philipp. (1983) 22; Type: Whitford & Hutchinson 9336 (US, lecto; L, NY, US, isolecto), Mindanao: Zamboanga, Port Banga, Jan/Feb-1908. Distribution: Endemic to the Philippines. LUZON: Camarines, Camarines Sur, MINDANAO: Zamboanga, Zamboanga del Sur. Conservation status: Endangered (IUCN 3.1). Native.
11. Drypetes littoralis (C.B.Rob.) Merr., PJS 29 (1926) 380; Airy Shaw, Euph. Born. (1975) 103; Alphab. Enum. Euph. Philipp. (1983) 22; ≡ Cyclostemon littoralis C.B.Rob., PJS 3c (1908) 198; Merrill, EPFP 2 (1923) 407; Type: R.S. Williams 377 (NY, lecto; GH, K, US, isolecto), Luzon: Bataan, along the coast at Lamao River, Mt Mariveles, staminate, 30-Dec-1903. = Drypetes mindorensis (Merr.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 274; ≡ Cyclostemon mindorensis Merr., PJS 9c (1914) 479; Merrill, EPFP 2 (1923) 408; Type: BS 21311 Escritor (NY, lecto; A, F, GH, K, L, P, SING, US, VT, isolecto), Mindoro: Mindoro Oriental, Naujan, Jul-1913. Distribution: Borneo, Lesser Sunda Isls, Philippines, Sulawesi. HUNDRED ISLS (photos), LUZON: Bataan, Batangas, Cagayan, Camarines, Ilocos Sur, La Union, Laguna, Nueva Ecija, Pangasinan, Quezon, Sorsogon, MINDANAO: Misamis, MINDORO, PALAWAN, POLILLO, ?SAMAR. Notes: Lowland primary forests, also along the seashore. Native.
12. Drypetes longifolia (Blume) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 245; Airy Shaw, Euph. Born. (1975) 103; Alphab. Enum. Euph. Philipp. (1983) 22; Whitmore, Tree Fl. Malaya 2 (1973) 88; ≡ Cyclostemon longifolius Blume, Bijdr. fl. Ned. Ind. (1826) 598. = Drypetes bordenii (Merr.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 244; ≡ Cyclostemon bordenii Merr., Publ. Bur. Sci. Gov. Lab. 17 (1904) 26; Merrill, EPFP 2 (1923) 405; Type: FB 333 P.T. Barnes (US, lecto; F, NY, isolecto), Luzon: Bataan, Lamao River, Feb-1904. = Cyclostemon incarnatus Elmer, LPB 7 (1915) 2639, as 'incarnata'; Levin, Gard. Bull. Singapore 73 (2021) 379; Type: Elmer 13265 (NY, lecto; A, G, GH, HBG, K, L, P, U, US, isolecto), Mindanao: Agusan del Norte, Cabadbaran, Mt Urdaneta, Jul-1912. = Cyclostemon merrittii Elmer, LPB 3 (1910) 913; Type: Elmer 12247 (NY, lecto; BISH, G, HBG, L, MO, US, isolecto), Sibuyan: Magallanes, Mt Giting-Giting, Apr-1910. Distribution: Andaman Isls, Borneo, Java, Lesser Sunda Isls, Malay Peninsula, Moluccas, New Guinea, Philippines, Sulawesi, Sumatra, Thailand. BABUYAN ISLS, LEYTE, LUZON: Apayao, Bataan, Cagayan, Camarines, Isabela (photos), Nueva Ecija, Nueva Vizcaya, Pampanga, Quezon, Sorsogon, Zambales (photos), MINDANAO: Agusan del Norte, Surigao, MINDORO, PALAWAN, PANAY, SAMAR, SIBUYAN. Notes: In primary forests at low elevation. Common. Conservation status: Least Concern (IUCN 3.1). Native.
13. Drypetes maquilingensis (Merr.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 240; Airy Shaw, Euph. Born. (1975) 104; Alphab. Enum. Euph. Philipp. (1983) 23; ≡ Cyclostemon maquilingensis Merr., PJS 9c (1914) 477; Merrill, EPFP 2 (1923) 407; Type: FB 20877 Villamil (US, lecto), Luzon: Laguna, Mt Makiling, Feb-1914. Distribution: Borneo, Lesser Sunda Isls, Moluccas, Philippines, Sulawesi. BALABAC, LUZON: Laguna, Sorsogon, MINDANAO: Agusan del Norte, Davao, Zamboanga, PALAWAN. Notes: Lowland forests, ascending to 700m. Native.
14. Drypetes microphylla (Merr.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 237; Airy Shaw, Euph. Born. (1975) 105; Alphab. Enum. Euph. Philipp. (1983) 23; Whitmore, Tree Fl. Malaya 2 (1973) 88; Romeroso et al., J. Asia-Pacific Biodiv. 17 (2024) 12; ≡ Cyclostemon microphyllus Merr., Publ. Bur. Sci. Gov. Lab. 17 (1904) 27; Merrill, EPFP 2 (1923) 407; Type: FB 175 P.T. Barnes (US, neo; BM, CAL, F, NY, P, isoneo), Luzon: Bataan, Lamao River, Jan-1904. = Cyclostemon oligophlebium Merr., PJS 20 (1922) 398; Merrill, EPFP 2 (1923) 408; Type: FB 12210 R. Rosenbluth (NY, lecto), Mindoro: Mindoro Oriental, Bongabong River, 21-May-1908. Distribution: Borneo, Malay Peninsula, Philippines. ALABAT, HOMONHON, LUZON: Aurora, Bataan, Cagayan, Camarines, Isabela, Nueva Ecija, Quezon, Rizal, Sorsogon, MASBATE, MINDANAO: Agusan del Norte, MINDORO, PALAWAN, PANAY, SAMAR. Notes: Throughout Luzon. Low and medium elevation primary forests. Native.
15. Drypetes mindanaensis (Merr.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 248; Airy Shaw, Alphab. Enum. Euph. Philipp. (1983) 23; ≡ Cyclostemon mindanaensis Merr., PJS 9c (1914) 479; Merrill, EPFP 2 (1923) 408; Type: FB 12458 V. Tarrosa (US, lecto; K, NY, P, isolecto), Mindanao: Zamboanga, Lumbiag, 21-Jul-1908. Distribution: Endemic to the Philippines. MINDANAO: Davao, Zamboanga. Notes: Lowland primary forests. Synonym of Drypetes sibuyanensis (Elmer) Pax & K.Hoffm.? Native.
16. Drypetes monosperma (Merr.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 255; Airy Shaw, Alphab. Enum. Euph. Philipp. (1983) 23; ≡ Cyclostemon monospermus Merr., PJS 1(Suppl.) (1906) 76; Merrill, EPFP 2 (1923) 408; Type: FB 2802 Meyer (NY, lecto; F, K, P, US, isolecto), Luzon: Bataan, Mt Mariveles, Lamao River, 3-Mar-1905. Distribution: Endemic to the Philippines. BABUYAN ISLS, CALAYAN, LUZON: Bataan, Bulacan, Cagayan, Camarines, Ilocos Norte, Laguna, Nueva Ecija, Rizal, MINDORO. Notes: In primary forests at low and medium elevation ascending to 1300m. Conservation status: Near Threatened (IUCN 3.1). Native.
17. Drypetes ovalis (J.J.Sm.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 269; Airy Shaw, Alphab. Enum. Euph. Philipp. (1983) 23; ≡ Hemicyclia ovalis J.J.Sm., Meded. Dept. Landb. Ned.-Indië 10 (1910) 221; Merrill, EPFP 2 (1923) 409. = Drypetes calcicola (Merr.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 266; ≡ Cyclostemon calcicola Merr., PJS 9c (1914) 478; Type: Merrill 9430 (US, lecto; BRI, CM, K, isolecto), Palawan: Taytay Bay, on a small limestone island known as Apulit, in dry thickets and crevices and ravines, steep slopes, c. 10m, 31-May-1913. Distribution: Java, Lesser Sunda Isls, Philippines, Sulawesi. APULIT, MALAMAWI, MINDORO, NEGROS, PALAWAN. Notes: Lowland thickets and forests on limestone. Native.
18. Drypetes rhakodiskos (Hassk.) Bakh.f., Blumea 12 (1963) 62; Kew Bull. 27 (1972) 76; Euph. Born. (1975) 106; Whitmore, Tree Fl. Malaya 2 (1973) 88; Odulio et al., Biodivers. Data J. 14 (2026) 33; ≡ Hemicyclia rhakodiskos Hassk., Verslagen Meded. Afd. Natuurk. Kon. Akad. Wetensch., 1st Ser. 4 (1855) 140. = Drypetes palawanensis (Merr.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 249; ≡ Cyclostemon palawanensis Merr., PJS 9c (1914) 480; Merrill, EPFP 2 (1923) 408; Type: Merrill 9451(BM, lecto; K, NY, US, isolecto), Palawan: Lake Manguao, on dry forested ridges, c. 80m, 27-Apr-1913. Distribution: Borneo, Java, Malay Peninsula, Philippines, Sumatra. BALABAC, PALAWAN, SAMAR. Notes: Lowland dry forests. DAO category: Other Threatened Species. Native.
19. Drypetes sibuyanensis (Elmer) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 247; Airy Shaw, Kew Bull. 23 (1969) 59; Euph. Born. (1975) 108; Alphab. Enum. Euph. Philipp. (1983) 24; ≡ Cyclostemon sibuyanensis Elmer, LPB 3 (1910) 914; Merrill, EPFP 2 (1923) 408; Type: Elmer 12275 (NY, lecto; A, BISH, BM, E, F, G, GH, HBG, K, L, MO, P, U, US), Sibuyan: Magallanes (Magdiwang), Mt Giting-giting, Apr-1910. Distribution: Borneo, Philippines, Sulawesi, Sumatra. SIBUYAN. Notes: Lowland forests, c. 225m. Conservation status: Least Concern (IUCN 3.1). Native.
20. Drypetes subcrenata (Merr.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 274; Airy Shaw, Alphab. Enum. Euph. Philipp. (1983) 24; ≡ Hemicyclia subcrenata (Merr.) Merr., EPFP 2 (1923) 409; ≡ Cyclostemon subcrenatus Merr., PJS 7c (1912) 388; Type: FB 21567 Tamesis (A, lecto; US, isolecto), Luzon: Batangas, near Sto Tomas, in forests at c. 200m, 22-May-1910. Distribution: Endemic to the Philippines. BANCALAN, LUZON: Albay, Batangas, Benguet, Camarines. Notes: Lowland forests. Conservation status: Endangered (IUCN 3.1). Native.

Possibly present

1. Drypetes celebica (Boerl. & Koord.) Pax & K.Hoffm., Pflanzenr. (Engler) IV (147 XV) (1922) 272; Airy Shaw, Alphab. Enum. Euph. Philipp. (1983) 21; ≡ Cyclostemon celebicus Boerl. & Koord., Meded. Lands Plantentuin 19 (1898) 591. Distribution: New Guinea, ?Philippines, Sulawesi. ?BANCALAN, ?LEYTE, ?LUZON: ?Quezon, ?SAMAR. Notes: Note: these locality records might pertain to Drypetes ellipsoidea (Merr.) Pax & K.Hoffm. Lowland forests.

Putranjiva Wall.

1. Putranjiva roxburghii Wall., Tent. Fl. Napal. 2 (1826) 61; ≡ Drypetes roxburghii (Wall.) Hurus., J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 6 (1954) 335. Distribution: Bangladesh, Borneo, Cambodia, India, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, Nepal, New Guinea, Pakistan, Philippines, Sri Lanka, Sulawesi, Thailand, Vietnam. POLILLO, SAMAR. Notes: Known from the Philippines from Barbon, Garcia & Sagcal PPI 6064 (L) and Fox 9213 (L). Conservation status: Least Concern (IUCN 3.1). Native.