Pteridaceae

Ceratopteridaceae, Sinopteridaceae, Antrophyaceae, Taenitidaceae, Parkeriaceae, Vittariaceae, Hemionitidaceae, Cheilanthaceae, Acrostichaceae, Adiantaceae

Acrostichum L.

Last edited by Julie F. Barcelona, 19 June 2019

Acrostichum aureum L., Sp. Pl. 2 (1753) 1069. = Parahemionitis arifolia (Burm.f.) Panigrahi, Amer. Fern J. 83 (1993) 90; ≡ Hemionitis arifolia (Burm.f.) T.Moore, Ind. Fil. (1859) 114. Distribution: Africa, America, Andaman Isls, Bangladesh, Bismarck Arch, Cambodia, China, Hong Kong, India, Indian Ocean, Lesser Sunda Isls, Malay Peninsula, Myanmar, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam. CATANDUANES, CORRIGIDOR, DINAGAT, LEYTE, LUZON: Abra, Albay, Bataan, Batangas, Benguet, Camarines Sur, Cavite, Ilocos Norte, Isabela, Kalinga-Apayao, Laguna, NCR, Pampanga, Quezon, Rizal, Tarlac, Zambales, MARINDUQUE, MINDANAO: Agusan del Norte, Davao del Sur, Maguindanao del Norte, Maguindanao del Sur, Misamis Oriental, Surigao del Sur, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY, POLILLO, SAMAR, SIASI, SULU ARCHIPELAGO. Native.
Acrostichum speciosum Willd., Sp. Pl., ed. 4 5 (1810) 117. Distribution: Andaman Isls, Australia, Bismarck Arch, China, Indian Ocean, Lesser Sunda Isls, Malay Peninsula, Moluccas, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, ?Sri Lanka, Sulawesi, Thailand, Vietnam. BALABAC, BILIRAN, LUZON: Bataan, NCR, Quezon, MINDANAO: Davao del Sur, Maguindanao del Norte, Zamboanga del Sur, MINDORO, PALAWAN, PANAY, SAMAR. Native.

Adiantum L.

Last edited by Pieter B. Pelser & Julie F. Barcelona, 8 November 2022

Adiantum capillus-veneris L., Sp. 2 (1753) 1096. Distribution: BATAN, CEBU, LUZON: Bataan, Benguet, Ifugao, Ilocos, Ilocos Norte, Mountain Province, Nueva Vizcaya, PALAWAN. Notes: Very widespread, including in the Philippines. Native.
Adiantum caudatum L., Mant. Pl. (1771) 308. Distribution: Bangladesh, Bhutan, Cambodia, China, India, Laos, Lesser Sunda Isls, Malay Peninsula, Nepal, New Caledonia, New Guinea, Pacific Ocean, Philippines, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam. BATANES, BOHOL, CATANDUANES, CEBU, DARAJUAY ISLS, GUIMARAS, LUZON: Bataan, Batangas, Benguet, Cavite, Ifugao, Ilocos, Ilocos Norte, Ilocos Sur, La Union, Laguna, Mountain Province, NCR, Nueva Ecija, Nueva Vizcaya, Pangasinan, Quezon, Rizal, Zambales, MINDANAO: Bukidnon, Davao, Zamboanga, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY, SULU ARCHIPELAGO. Native.
Adiantum ciliatum Blume, Enum. Pl. Javae 2 (1828) 215. = Adiantum malesianum Ghatak, Bull. Bot. Surv. India 5 (1963) 73. Distribution: Borneo, China, Malay Peninsula, Moluccas, Myanmar, Philippines, Sulawesi, Sumatra, Taiwan, Vietnam. BOHOL, LUZON: Benguet, Ifugao, Laguna, Rizal, MINDANAO: Bukidnon, MINDORO, PALAWAN. Native.
Adiantum cupreum Copel., LPB 4 (1911) 1152. Distribution: Endemic to the Philippines. LUZON: Isabela, PANAY, SIBUYAN. Conservation status: Vulnerable (DAO 2017-11). Native.
Adiantum diaphanum Blume, Enum. Pl. Javae 2 (1828) 215. = Adiantum setulosum J.Sm., Companion Bot. Mag. 72 (1846) 22. Distribution: Australia, Borneo, China, Japan, Lesser Sunda Isls, Moluccas, New Caledonia, New Guinea, Pacific Ocean, Philippines, Ryukyu Isls, Solomon Isls, Sulawesi, Taiwan, Vietnam. LUZON: Bataan, Batangas, Benguet, Cavite, Ifugao, Ilocos Norte, Kalinga, Laguna, Mountain Province, Pampanga, Quezon, Rizal, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao, Davao del Sur, Sarangani, South Cotabato, Zamboanga del Sur, SAMAR. Native.
Adiantum edgeworthii Hook., Sp. Fil. 2 (1851) 14. Distribution: Bhutan, China, India, Japan, Lesser Sunda Isls, Malay Peninsula, Myanmar, Nepal, Philippines, Taiwan, Thailand, Tibet, Vietnam. LUZON: Benguet. Native.
Adiantum flabellulatum L., Sp. 2 (1753) 1095. Distribution: Bangladesh, Cambodia, China, India, Japan, Laos, Lesser Sunda Isls, Malay Peninsula, Myanmar, Philippines, Ryukyu Isls, Singapore, Sri Lanka, Taiwan, Thailand, Vietnam. LUZON: Ilocos, Ilocos Norte. Native.
Adiantum hispidulum Sw., J. Bot. (Schrader) 1800 (1801) 82. Distribution: Africa, Australia, China, India, Indian Ocean, Java, Lesser Sunda Isls, Malay Peninsula, New Caledonia, Pacific Ocean, Philippines, Singapore, Sri Lanka, Sulawesi, Taiwan. LUZON: Abra, Bataan, Benguet, Ilocos Norte, Kalinga, Mountain Province, Rizal. Native.
Adiantum hosei Baker, J. Bot. 26 (1888) 324; Coritico et al., PJS 149 (2020) 782; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22048. Distribution: Borneo, Philippines, Sulawesi. MINDANAO: Agusan del Norte, Bukidnon, Davao Oriental, South Cotabato. Native.
Adiantum mindanaoense Copel., PJS 1 (Suppl. 2) (1906) 154. Distribution: Moluccas, New Guinea, Philippines, Sulawesi. MINDANAO: Zamboanga del Sur. Conservation status: Endangered (DAO 2017-11). Native.
Adiantum monosorum Baker, Syn. Fil. (Hooker & Baker), ed. 2 (1874) 472. = Adiantum scabripes Copel., PJS, C 7 (1912) 55. Distribution: Moluccas, Philippines, Solomon Isls, Sulawesi. MINDANAO: Agusan del Norte, Davao Oriental. Conservation status: Vulnerable (DAO 2017-11). Native.
Adiantum philippense L., Sp. Pl. 2 (1753) 1094. Distribution: BALABAC, GUIMARAS, LUBANG, LUZON: Abra, Bataan, Batangas, Benguet, Bulacan, Cavite, Ifugao, Kalinga, Laguna, Mountain Province, NCR, Pampanga, Pangasinan, Quezon, Rizal, Sorsogon, MINDORO, NEGROS, PALAWAN, PANAY. Native.
- ssp. philippense. Distribution: Africa, Andaman Isls, Australia, Bangladesh, Bhutan, Cambodia, China, India, Indian Ocean, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, Nepal, Pacific Ocean, Philippines, ?Solomon Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam. PALAWAN. Native.
- ssp. intermedium S.C.Verma & Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008). Distribution: Bhutan, Bismarck Arch, China, India, Myanmar, Nepal, New Guinea, Pacific Ocean, Philippines, Solomon Isls, Sulawesi, Thailand, Vietnam. Native.
Adiantum soboliferum Wall. ex Hook., Sp. Fil. 2 (1851) 13. Distribution: Africa, ?Cambodia, China, India, Laos, Lesser Sunda Isls, Madagascar, Malay Peninsula, Myanmar, Philippines, Taiwan, Thailand, Vietnam. CULION, LUZON: Bataan, Batangas, Ilocos Norte, Laguna, NCR, Rizal, MINDANAO: Cotabato, PALAWAN. Native.
Adiantum stenochlamys Baker, Ann. Bot. (Oxford) 5 (1891) 209. = Adiantum opacum Copel., PJS 1 (Suppl. 4) (1906) 255. Distribution: Andaman Isls, Borneo, Cambodia, Laos, Malay Peninsula, Philippines, Singapore, Sulawesi, Sumatra, Thailand, Vietnam. BICAWAYAN, LUZON: Isabela, PALAWAN, SIBUYAN. Conservation status: Vulnerable (DAO 2017-11). Native.
Adiantum tenerum Sw., Prodr. (1788) 135. Notes: Introduced from South America, not native to the Philippines. Naturalized.

Aleuritopteris Fée

Last edited by Julie F. Barcelona, 19 July 2019

Aleuritopteris albomarginata (C.B.Clarke) Ching, Hong Kong Naturalist 10 (1941) 199; ≡ Cheilanthes albomarginata C.B.Clarke, Trans. Linn. Soc. London, Bot. 1 (1880) 456. Distribution: Bhutan, China, India, Myanmar, Nepal, Pakistan, Philippines, Thailand, Tibet, Vietnam. Native.
Aleuritopteris dubia (C.Hope) Ching, Hongkong Nat. 10 (1941) 200; ≡ Cheilanthes dubia C.Hope, J. Bombay Nat. Hist. Soc. 12 (1899) 528. = Aleuritopteris rufa auct. non (D.Don) Ching; Copel. Distribution: LUZON: Benguet, Mountain Province. Native.
Aleuritopteris formosana (Hayata) Tagawa, Acta Phytotax. Geobot. 14 (1952) 191; ≡ Cheilanthes formosana Hayata, Enum. Pl. Formosa (1906) 61. Distribution: Bhutan, China, India, Myanmar, Nepal, Pakistan, Philippines, Taiwan, Thailand, Tibet. LUZON: Bataan, Batangas, Benguet, Camarines Norte, Ilocos Norte, Ilocos Sur, Mountain Province. Native.
Aleuritopteris krameri (Franch. & Sav.) Ching, Hongkong Nat. 10 (1941) 202; ≡ Cheilanthes krameri Franch. & Sav., Enum. Pl. Jap. 2 (1879) 212. = Aleuritopteris farinosa auct. non (Forssk.) Fée. Distribution: Japan, Philippines, Taiwan, Thailand. LUZON: Benguet, Ifugao, Mountain Province, Nueva Vizcaya. Native.

Antrophyum Kaulf.

Last edited by Julie F. Barcelona, 7 September 2020

Antrophyum callifolium Blume, Enum. Pl. Javae 2 (1828) 111. = Antrophyum nanum Fée, Mém. Foug. 4 (1851) 44. = Antrophyum alatum Brack., U.S. Expl. Exped., Filic. 16 (1854) 64. Distribution: Andaman Isls, Australia, Bismarck Arch, Borneo, Cambodia, China, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. CATANDUANES, CEBU, LUZON: Aurora, Bataan, Batangas, Benguet, Cagayan, Camarines, Ilocos Norte, Laguna, Nueva Ecija, Quezon, Rizal, Sorsogon, MANGSEE, MINDANAO: Agusan del Norte, Cotabato, Davao, Davao Oriental, Davao del Norte, Davao del Sur, Lanao del Sur, Zamboanga, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY, SAMAR, SIASI. Native.
Antrophyum latifolium Blume, Fl. Javae 2 (1829) 75. Distribution: Borneo, Java, Moluccas, New Guinea, Philippines, Sulawesi, Sumatra. MINDANAO: Bukidnon, Cotabato, Davao, Davao del Sur, South Cotabato, Zamboanga del Sur, PANAY. Native.
Antrophyum ledermannii Hieron., Bot. Jahrb. Syst. 56 (1920) 175. Distribution: Moluccas, New Guinea, Philippines. MINDANAO: Bukidnon, Davao del Sur, South Cotabato. Native.
Antrophyum parvulum Blume, Pl. Jav. (1828) 110; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22048. = Antrophyum immersum (Bory ex Willd.) Mett., Ann. Mus. Bot. Lugd.-Bat. 4 (1868) 171; ≡ Hemionitis immersa Bory ex Willd., Sp. 5 (1810) 127. Distribution: Africa, Borneo, ?Cambodia, China, India, Indian Ocean, Japan, Java, ?Laos, Lesser Sunda Isls, Malay Peninsula, Nicobar Isls, Philippines, Singapore, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. LUZON: Bataan, Laguna, Mountain Province, MINDANAO: Bukidnon, Davao Oriental, Davao del Sur, Lanao del Sur, South Cotabato, Zamboanga del Sur, NEGROS, PALAWAN. Native.
Antrophyum plantagineum (Cav.) Kaulf., Enum. Filic. (1824) 197; ≡ Hemionitis plantaginea Cav., Descr. Pl. (1802) 260. Distribution: Australia, Bangladesh, Bismarck Arch, India, Java, Lesser Sunda Isls, Malay Peninsula, Moluccas, New Caledonia, New Guinea, Pacific Ocean, Philippines, Solomon Isls, Sri Lanka, Sulawesi, Sumatra. LUZON: NCR, Pangasinan, Quezon, MINDANAO: Bukidnon, Cotabato, Davao, Davao del Sur, Zamboanga del Sur, NEGROS. Native.
Antrophyum reticulatum (G.Forst.) Kaulf., Enum. Filic. (1824) 198; ≡ Hemionitis reticulata G.Forst., Fl. Ins. Austr. (1786) 79. = Antrophyum clementis Christ, PJS 2 c (1907) 175. Distribution: Andaman Isls, Bhutan, Borneo, China, India, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, Nepal, Nicobar Isls, Pacific Ocean, Philippines, Sri Lanka, Sulawesi, Tibet, Vietnam. GENERAL ISLAND, LEYTE, LUZON: Albay, Aurora, Bataan, Benguet, Cagayan, Camarines, Camarines Norte, Cavite, Ilocos Norte, Isabela, Laguna, Mountain Province, Nueva Vizcaya, Pampanga, Pangasinan, Quezon, Rizal, Sorsogon, MINDANAO: Agusan del Norte, Davao del Sur, Lanao del Sur, Surigao, Surigao del Sur, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY, SAMAR. Native.
Antrophyum semicostatum Blume, Enum. Pl. Javae 2 (1828) 110. Distribution: Borneo, Lesser Sunda Isls, Malay Peninsula, Moluccas, Pacific Ocean, Philippines, Solomon Isls, Sulawesi, Sumatra. LUZON: Batangas, Laguna, MINDANAO: Agusan del Norte, Bukidnon, Davao del Sur, MINDORO, TAWI-TAWI. Native.
Antrophyum sessilifolium (Cav.) Spreng., Syst. Veg. 4 (1827) 67; ≡ Hemionitis sessilifolia Cav., Descr. Pl. (1802) 261. Distribution: Borneo, Philippines, Sulawesi, Taiwan. BATAN, BOHOL, CAMIGUIN, LEYTE, LUZON: Abra, Bataan, Benguet, Cagayan, Ifugao, Laguna, Nueva Vizcaya, Pampanga, Quezon, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Davao, Davao Oriental, Davao del Sur, Lanao del Sur, Sarangani, Surigao, NEGROS, PALAWAN, PANAY, POLILLO, SAMAR, TABLAS. Native.
Antrophyum williamsii Benedict, Amer. Fern J. 1 (1911) 72. Distribution: Endemic to the Philippines. LUZON: Benguet, MINDANAO: Lanao del Sur. Native.

Austrogramme E.Fourn.

Last edited by Julie F. Barcelona, 20 June 2019

Austrogramme luzonica (Alderw.) M.Kato, J. Fac. Sci. Univ. Tokyo, Bot. 14 (1988) 174; ≡ Taenitis luzonica (Alderw.) Holttum, Kew Bull. 30 (1975) 335; ≡ Syngramma luzonica Alderw., Bull. Jard. Bot. Buitenzorg Ser. 2I, 2 (1920) 174. Distribution: Endemic to the Philippines. LUZON: Apayao, Cagayan, Camarines Norte, Isabela, Laguna, Pangasinan, Quezon, SAMAR. Native.

Calciphilopteris Yesilyurt & H.Schneid.

Edited by Pieter B. Pelser, 15 August 2014

Calciphilopteris wallichii (J.Sm.) Yesilyurt & H.Schneid., Phytotaxa 7 (2010) 57; ≡ Doryopteris wallichii J.Sm., Journal of Botany 4 (1841) 404; Type: Cuming 238 (K, holo; BM, C, G, L, iso). Distribution: Endemic to the Philippines. LUZON: Benguet, Camarines, Pangasinan, Rizal. Notes: Note by J.F. Barcelona (15-Aug-14): Philippine specimens under Doryopteris ludens auct. non. (Wall. ex Hook) J.Sm. refer to this species. Native.

Ceratopteris Brongn.

Last edited by Julie F. Barcelona, 22 November 2021

Ceratopteris oblongiloba Masuyama & Watano, Acta Phytotax. Geobot. 61 (2010) 84; Type: from Indonesia. Distribution: Cambodia, Java, Philippines, Sumatra, Thailand. LUZON: Rizal. Native.
Ceratopteris thalictroides (L.) Brongn., Bull. Sci. Soc. Philom. Paris 8 (1821) 186; Batuyong et al., Biodiversitas 21 (2020) 3975; ≡ Acrostichum thalictroides L., Sp. Pl. 2(1753) 1070; Type: from Sri Lanka. = Naias obvoluta Blanco, Fl. Filip. Ed. 2 (1845) 460. = Ceratopteris siliquosa (L.) Copel., PJS 6 (1935) 107; ≡ Acrostichum siliquosum L., Sp. Pl. 2 (1753) 1070. Distribution: Asia, C America, Pacific Ocean. BASILAN, BOHOL, LUZON: Bataan, Batangas, Ifugao, Ilocos Norte, Laguna, NCR, Pampanga, Quezon, Rizal, Sorsogon, Tarlac, Zambales, MACTAN, MINDANAO: Agusan del Norte, Davao, Lanao del Sur, Maguindanao del Sur, Surigao, Surigao del Sur, Zamboanga, Zamboanga del Sur, MINDORO, SAMAR. Conservation status: Endangered (DAO 2017-11). Native.

Excluded

Ceratopteris gaudichaudii Brongn., Bull. Sci. Soc. Philom. Paris (1821) 187.
- var. vulgaris Masuyama & Watano, Acta Phytotax. Geobot. 61 (2010) 79. Notes: Mistakenly reported for the Philippines by Kinosian et al. (2020) (S.P. Kinosian, pers. comm. 8-Sep-20).

Coniogramme Fée

Last edited by Julie F. Barcelona, 3 June 2019

Coniogramme fraxinea (D.Don) Diels in Engl. & Prantl, Nat. Pflanzenfam. 1 (1899) 262; ≡ Diplazium fraxineum D.Don, Prodr. Fl. Nepal. (1825) 12. = Coniogramme macrophylla (Blume) Hieron., Hedwigia 57 (1916) 291; ≡ Coniogramme javanica var. macrophylla Blume, Enum. Pl. Javae 2 (1828) 113. Distribution: Bhutan, Borneo, ?Cambodia, China, India, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, Nepal, Pacific Ocean, Philippines, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam. LUZON: Aurora, Batangas, Benguet, Ifugao, Isabela, Kalinga, Laguna, Mountain Province, Nueva Vizcaya, Quezon, MINDANAO: Bukidnon, Cotabato, Davao, Davao Oriental, Davao del Sur, Lanao del Sur, South Cotabato, Zamboanga del Sur, NEGROS. Native.
Coniogramme serrulata (Blume) Fée, Mém. Foug., 5. Gen. Filic. 167 (1850) t. 14 B. f. 2; ≡ Gymnogramma serrulatum Blume, Enum. Pl. Jav. (1828) 113. = Coniogramme subcordata Copel., LPB 3 (1910) 823. Distribution: Bhutan, China, India, Japan, Java, Korea, Laos, Myanmar, Nepal, Pakistan, Philippines, Russia, Sri Lanka, Taiwan, Thailand, Tibet, Vietnam. LUZON: Benguet, Ifugao, Mountain Province, Nueva Vizcaya, MINDANAO: Bukidnon, Cotabato, Davao, Davao del Sur, South Cotabato. Native.

Doryopteris J.Sm.

Last edited by Julie F. Barcelona, 11 October 2020

Doryopteris kirkii (Hook.) Alston, Bol. Soc. Brot. sér. 2, 30 (1956) 14; Coritico & Amoroso, Nature Conservation Research 5 (2020) 82; ≡ Doryopteris concolor var. kirkii (Hook.) T.C.E.Fr., Wiss. Erg. d. Schwed. Rhodesia-Kongo Exp. 1911-12, Bot. 1 (1941) 4; ≡ Cheilanthes kirkii Hook., Sec. Cent. Ferns (1861) t. 81. = Doryopteris cuspidata Copel., PJS 38 (1929) 148. Distribution: Africa, Australia, Borneo, China, India, Indian Ocean, Lesser Sunda Isls, New Caledonia, New Guinea, Pacific Ocean, Philippines, Sri Lanka, Sulawesi, Taiwan, Vietnam. MINDANAO: Davao Oriental. Conservation status: Endangered (DAO 2017-11, as Doryopteris concolor). Native.

Haplopteris C.Presl

Last edited by Pieter B. Pelser & Julie F. Barcelona, 5 November 2022

Haplopteris alternans (Copel.) S.Linds. & C.W.Chen, Gard. Bull. Singapore 66 (2014) 170; ≡ Vittaria alternans Copel., PJS 1 (Suppl. 2) (1906) 157. = Vittaria hecistophylla Copel., PJS 81 (1952) 45. Distribution: Philippines, Sulawesi. LEYTE, LUZON: Benguet, Mountain Province, MINDANAO: Agusan, Agusan del Norte, Bukidnon, Cotabato, Davao Oriental, Davao del Sur, Lanao del Sur, Zamboanga del Sur, NEGROS. Native.
Haplopteris anguste-elongata (Hayata) E.H.Crane, Syst. Bot. 22 (1998) 514; ≡ Vittaria anguste-elongata Hayata, Icon. Pl. Formosan. 6 (1916) 161. Distribution: China, Philippines, Ryukyu Isls, Taiwan. LUZON: Aurora. Native.
Haplopteris capillaris (Copel.) C.W.Chen, S.Linds. & K.T.Yong, Syst. Bot. 44 (2019) 491; ≡ Vaginularia capillaris (Copel.) C.Chr., Index Filic., Suppl. Tert. (1934) 193; ≡ Monogramma capillaris Copel., PJS 6c (1911) Bot. 147; Type: Merrill 6961 (PNH, holo, lost; MICH, lecto; NY, P, isolecto), Negros: Mt Kanlaon, Apr-1910. Distribution: Borneo, Philippines, Taiwan. NEGROS. Native.
Haplopteris dareicarpa (Hook.) S.Linds. & C.W.Chen, Gard. Bull. Singapore 66 (2014) 171; ≡ Monogramma dareicarpa Hook., Sp. Fil. 5 (1864) 121. Distribution: ?Australia, Borneo, Malay Peninsula, New Guinea, Philippines, Singapore, Solomon Isls. LUZON: Camarines Norte, Laguna, Quezon, MINDANAO: Agusan del Sur, MINDORO, NEGROS, PANAY (photos). Native.
Haplopteris elongata (Sw.) E.H.Crane, Syst. Bot. 22 (1998) 514; ≡ Vittaria elongata Sw., Syn. Fil. 109 (1806) 302. = Vittaria intermedia Blume, Enum. Pl. Javae 2 (1828) 199. = Vittaria philippinensis Christ, Bull. Herb. Boissier II. 6 (1906) 1007. = ?Vittaria crispomarginata Christ, Bull. Herb. Boissier II. 6 (1907) 1007; Type: Loher s.n. (BM, P), Luzon: Makiling. Distribution: Africa, Andaman Isls, Australia, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, New Caledonia, New Guinea, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Tibet, Vietnam. BASILAN, BOHOL, CAMIGUIN DE BABUYANES, CATANDUANES, LEYTE, LUZON: Abra, Albay, Aurora, Bataan, Batangas, Benguet, Camarines Sur, Ifugao, Isabela, Kalinga, Laguna, Mountain Province, Nueva Vizcaya, Pampanga, Quezon, Rizal, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao, Davao del Sur, Lanao del Sur, Sarangani, South Cotabato, Sultan Kudarat, Surigao, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY, POLILLO, SAMAR, SIBUTU. Notes: Note by J.F. Barcelona (8-Aug-2020): It is presently unclear if Vittaria crispomarginata is taxonomically distinct from Haplopteris elongata. Native.
Haplopteris ensiformis (Sw.) E.H.Crane, Syst. Bot. 22 (1998) 514; ≡ Vittaria ensiformis Sw., Ges. Nat. Fr. Berl. Neu. Schr. 2 (1799) 134. = Vittaria incurvata Cav., Descr. (1802) 270. = Vittaria lineata auct. non Sm. = Vittaria pachystemma Christ, PJS 2 c (1907) 174. Distribution: Africa, Australia, ?Bangladesh, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, New Guinea, Pacific Ocean, Philippines, Ryukyu Isls, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. BALABAC, BASILAN, BATAN DE ALBAY, CAMIGUIN, LUZON: Albay, Apayao, Aurora, Bataan, Benguet, Cagayan, Camarines Norte, Ifugao, Ilocos Norte, Laguna, Mountain Province, Nueva Vizcaya, Quezon, Rizal, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao, Davao del Sur, MINDORO, NEGROS, PALAWAN, POLILLO, SAMAR, SIBUTU. Native.
Haplopteris mediosora (Hayata) X.C.Zhang, Ann. Bot. Fenn. 40 (2003) 460; Delos Angeles et al., 151 PJS (2022) 1933; ≡ Vittaria mediosora Hayata, Icon. Pl. Formosan. 5 (1915) 346. Distribution: China, Japan, Myanmar, Nepal, Philippines, Taiwan, Tibet. LUZON: Benguet, Mountain Province, SAMAR. Native.
Haplopteris mindanaoensis S.Linds. & C.W.Chen, Gard. Bull. Singapore 70 (2018) 184; Type: L.-Y. Kuo 3659 (PNH, holo; CMUH, SING, TAIF, iso), Mindanao: Mt Kitanglad, 18-Dec-2012. Distribution: Endemic to the Philippines. MINDANAO: Bukidnon, Davao del Sur. Native.
Haplopteris scolopendrina (Bory) C.Presl, Tent. Pterid. (1836) 141; ≡ Vittaria scolopendrina (Bory) Thwaites, Enum. pl. Zeyl. (1864) 381; ≡ Pteris scolopendrina Bory, Voy. 2 (1804) 323. = Vittaria subcoriacea Christ, PJS 2 c (1907) 175. Distribution: Borneo, Indian Ocean, Malay Peninsula, Moluccas, New Guinea, Pacific Ocean, Philippines, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand. BASILAN, JOLO, LEYTE, LUZON: Cagayan, Laguna, MINDANAO: Bukidnon, Davao Oriental, Davao del Sur, Surigao, MINDORO, NEGROS, PALAWAN, POLILLO, SAMAR. Native.
Haplopteris stenophylla (Copel.) C.W.Chen & S.Linds., Kew Bull. 76 (2021); ≡ Vittaria stenophylla Copel., PJS 40 (1929) 312; Type: Copeland 122 (MICH, lecto; BO, K, M, MU, NY, SING, UC, US, isolecto), Luzon: Benguet, Mt Santo Tomas, May-1909, 2300m. Distribution: Endemic to the Philippines. LUZON: Abra, Benguet, Kalinga, SAMAR. Native.
Haplopteris taeniophylla (Copel.) E.H.Crane, Syst. Bot. 22 (1998) 514; ≡ Vittaria taeniophylla Copel., PJS 1 (Suppl. 2) (1906) 157. Distribution: Bhutan, Cambodia, China, India, Japan, Laos, Myanmar, Nepal, Philippines, Sulawesi, Thailand, Tibet, Vietnam. LUZON: Benguet, Ifugao, Mountain Province, Nueva Vizcaya, Sorsogon, MINDANAO: Agusan del Norte. Native.
Haplopteris zosterifolia (Willd.) E.H.Crane, Syst. Bot. 22 (1998) 514; ≡ Vittaria zosterifolia Willd., Sp. Pl. 5 (1810) 406. = Vittaria copelandii Alderw., Bull. Jard. Bot. Buitenzorg Ser. 2, VII (1912) 28. = Vittaria merrillii Christ, PJS 2 (1907) 174. Distribution: Andaman Isls, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, Java, Korea, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Ryukyu Isls, Solomon Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam. BALABAC, BASILAN, CAMIGUIN DE BABUYANES, LUZON: Apayao, Bataan, Laguna, Quezon, Sorsogon, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao, Davao del Sur, South Cotabato, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN. Native.

Mickelopteris Fraser-Jenk.

Last edited by Julie F. Barcelona, 19 June 2019

Mickelopteris cordata (Roxb. ex Hook. & Grev.) Fraser-Jenk., Annot. Checkl. Ind. Pterid. 1 (2016) 247; ≡ Parahemionitis cordata (Roxb. ex Hook. & Grev.) Fraser-Jenk., New Sp. Syndr. Indian Pteridol. (1997) 187; ≡ Hemionitis cordata Roxb. ex Hook. & Grev., Ic. Fil. (1828) t. 64. Distribution: Bangladesh, Cambodia, China, India, Laos, Lesser Sunda Isls, Malay Peninsula, Myanmar, Philippines, Sri Lanka, Taiwan, Thailand, Vietnam. LUZON: Bataan, Benguet, Laguna, NCR, Rizal. Native.

Oeosporangium Vis.

Last edited by Julie F. Barcelona & Pieter B. Pelser, 6 August 2022

Oeosporangium belangeri (Bory) Fraser-Jenk., Annot. Checkl. Ind. Pterid. 1 (2016) 255; ≡ Cheilanthes belangeri (Bory) C.Chr., Index Filic. (1905) 172; ≡ Pteris belangeri Bory, Voy. Indes Or. 2 (1833) 44. Distribution: Bangladesh, Borneo, Cambodia, China, India, Laos, Malay Peninsula, Myanmar, Nepal, Philippines, Thailand, Vietnam. LUZON: NCR. Native.
Oeosporangium chusanum (Hook.) Fraser-Jenk., Annot. Checkl. Ind. Pterid. 1 (2016) 256; ≡ Cheilanthes mysurensis var. chusana (Hook.) Christ, Ind. (1901) 293; ≡ Cheilanthes chusana Hook., Sp. Fil. 2 (1852) 95. Distribution: China, Japan, Korea, Laos, Philippines, Sulawesi, Taiwan, Vietnam. LUZON: Batangas, Ilocos Norte, Laguna, MINDANAO: Davao del Sur. Native.
Oeosporangium tenuifolium (Burm.f.) Fraser-Jenk. & Pariyar, Annot. Checkl. Ind. Pterid. 1 (2016) 261; ≡ Cheilanthes tenuifolia (Burm.f.) Sw., Syn. Fil. (1806) 129. Distribution: Andaman Isls, Australia, Bangladesh, Borneo, Cambodia, China, India, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Nepal, New Caledonia, Pacific Ocean, Pakistan, Philippines, Singapore, Solomon Isls, Sri Lanka, Taiwan, Thailand, Vietnam. LUZON: NCR, PALAWAN. Native.

Incertae sedis

Cheilanthes nudiuscula (R.Br.) T.Moore, Index Fil. (1861) 249; ≡ Hemionitis nudiuscula (R.Br.) Christenh., Global Fl. 4 (2018) 18; ≡ Pteris nudiuscula R.Br., Prodr. Fl. Nov. Holland. (1810) 155. = Cheilanthes javensis (Desv.) T.Moore, Index Fil. (1861) 244; ≡ Cincinalis javensis Desv., Berl. Mag. 5 (1811) 329. = Cheilanthes densa (J.Sm.) Fée, Mém. Foug. 5 (1852) 156; ≡ Notholaena densa J.Sm., J. Bot. (Hooker) 3 (1841) 395; Type: Cuming 282 (K, MICH, UC, US), Luzon. = Cheilanthes dilimanensis P.M.Zamora, Kalikasan 3 (1974) 193; Type: Zamora 71-886 (PUH, holo; MICH, iso), Luzon: Quezon City, Diliman, 1971. Distribution: Australia, China, Java, Lesser Sunda Isls, Moluccas, New Caledonia, New Guinea, Pacific Ocean, Philippines, Sulawesi, Taiwan. LUZON: Quezon. Notes: Pelser (6-Aug-22): It is presently unclear to which species of Oeosporangium (or a different genus?) Philippine specimens of this taxon belong. Native.

Onychium Kaulf.

Last edited by Julie F. Barcelona, 20 June 2019

Onychium japonicum (Thunb.) Kunze, Bot. Zeitung (Berlin) 6 (1848) 507. Distribution: China, Japan, Java, Korea, Lesser Sunda Isls, Pacific Ocean, Philippines, Ryukyu Isls, Taiwan, Thailand. LUZON: Benguet, Ifugao, Kalinga, Laguna, Mountain Province, Nueva Vizcaya, Quezon. Native.
Onychium siliculosum (Desv.) C.Chr., Index Filic. fasc. 8 (1906) 469; ≡ Pteris siliculosa Desv., Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5 (1811) 324. Distribution: Andaman Isls, ?Andaman Isls, Australia, Bangladesh, Bhutan, Bismarck Arch, Cambodia, China, India, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Myanmar, Nepal, New Guinea, Philippines, Sulawesi, Taiwan, Thailand, Tibet, Vietnam. LEYTE, LUZON: Apayao, Batangas, Benguet, Bulacan, Cavite, Ifugao, Ilocos Norte, Kalinga, Laguna, Mountain Province, NCR, Nueva Ecija, Pampanga, Pangasinan, Quezon, Rizal, Sorsogon, Tarlac, Zambales, MINDANAO: Bukidnon, Lanao del Sur, Zamboanga del Sur, PALAWAN, PANAY. Native.

Paragymnopteris K.H.Shing

Last edited by Julie F. Barcelona, 3 June 2019

Paragymnopteris vestita (Wall. ex C.Presl) K.H.Shing, Indian Fern J. 10 (1994) 230. 1994; ≡ Gymnopteris vestita (Wall. ex C.Presl) Underw., Bull. Torrey Bot. Club 29 (1902) 627; ≡ Gymnogramma vestita Wall. ex C.Presl, Tent. Pterid. (1836) 218. Distribution: Bhutan, China, India, Myanmar, Nepal, Pakistan, Philippines, Taiwan, Thailand, Tibet. LUZON: Benguet, Mountain Province. Native.

Pityrogramma Link (Not native)

Last edited by Julie F. Barcelona, 7 September 2020

Pityrogramma calomelanos (L.) Link, Handb. Gewachs. 3 (1833) 20; ≡ Acrostichum calomelanos L., Sp. Pl. 2 (1753) 1072. Distribution: America. MINDANAO: Bukidnon. Naturalized.

Pteris L.

Last edited by Julie F. Barcelona & Pieter B. Pelser, 5 February 2023

Pteris amoena Blume, Enum. Pl. Jav. (1828) 210. = Pteris purpureorhachis Copel., PJS 12 (1917) 48. Distribution: Borneo, China, India, Japan, Java, Laos, Myanmar, Philippines, Ryukyu Isls, Solomon Isls, Taiwan, Thailand, Tibet, Vietnam. MINDANAO: Bukidnon. Native.
Pteris arisanensis Tagawa, Acta Phytotax. Geobot. 6 (1936) 102. Distribution: Bhutan, China, Hong Kong, India, Myanmar, Nepal, Philippines, Sri Lanka, Taiwan, Thailand, Tibet, Vietnam. Native.
Pteris armata C.Presl, Reliq. Haenk. 1 (1825) 56. Distribution: Endemic to the Philippines. BASILAN, BOHOL, LEYTE, LUZON: Bataan, Batangas, Benguet, Cagayan, Isabela, Laguna, Nueva Ecija, Pangasinan, Quezon, Rizal, Sorsogon, MINDANAO: Lanao, Lanao del Norte, Surigao del Norte, Zamboanga, Zamboanga del Sur, PALAWAN, PANAY, SAMAR, SULU ARCHIPELAGO. Native.
Pteris biaurita L., Sp. Pl. 2 (1753) 1076. Distribution: BOHOL, LUZON: Abra, Bataan, Benguet, Cavite, Ifugao, Ilocos Norte, Laguna, Mountain Province, NCR, Nueva Vizcaya, Pampanga, Pangasinan, Quezon, Rizal, Zambales, MINDANAO: Bukidnon, Zamboanga, MINDORO, PALAWAN. Native.
- ssp. fornicata Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 116. = Pteris pacifica Hieron., Hedwigia 66 (1914) 355; Rufila et al., Ruhuna J. Science 13 (2022) 159. = Pteris cumingii Hieron., Hedwigia 66 (1914) 358. Distribution: Andaman Isls, Australia, Bangladesh, Bhutan, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, Nepal, Pacific Ocean, Philippines, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam. LUZON, MINDANAO: Bukidnon, Davao del Sur, Zamboanga, PALAWAN. Native.
- ssp. walkeriana Fraser-Jenk. & Rajkumar, Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 115. Distribution: Andaman Isls, Bangladesh, Bhutan, China, India, Java, Laos, Myanmar, Nepal, Philippines, Thailand, Tibet. Native.
Pteris blumeana J.Agardh, Rec. Pterid. (1839) 22. Distribution: Bangladesh, Bismarck Arch, Cambodia, ?China, India, Java, Laos, Moluccas, Myanmar, New Guinea, Philippines, Solomon Isls, Sulawesi, Thailand, Vietnam. BALABAC, BASILAN, JOLO, LEYTE, LUZON: Apayao, Batangas, Benguet, Bulacan, Cagayan, Laguna, Nueva Ecija, Pampanga, Quezon, MINDANAO: Agusan del Norte, Davao, Davao del Sur, Lanao del Sur, Zamboanga, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY. Native.
Pteris brevis Copel., PJS 81 (1952) 7. Distribution: Endemic to the Philippines. BASILAN. Conservation status: Vulnerable (DAO 2017-11). Native.
Pteris calocarpa (Copel.) M.G.Price, Kew Bull. 29 (1975) 727; ≡ Copelandiopteris calocarpa (Copel.) Stone, Webbia 28 (1973) 493; ≡ Schizostege calocarpa Copel., PJS 1 (1906) 155. Distribution: Endemic to the Philippines. MINDANAO: Zamboanga, Zamboanga del Sur. Conservation status: Critically Endangered (DAO 2017-11). Native.
Pteris cretica L., Mant. Pl. 1 (1767) 130. = Pteris stenophylla auct. non Hook. & Grev.; Copel., FFP 1 (1958) 129. Distribution: CORRIGIDOR, MINDORO, PALAWAN. Native.
- ssp. cretica. Distribution: Africa, Bangladesh, Bhutan, China, Europe, India, Indian Ocean, Japan, Korea, ?Malay Peninsula, Middle East, Myanmar, Nepal, Pacific Ocean, Pakistan, Philippines, Ryukyu Isls, Sri Lanka, Taiwan, Thailand, Tibet, Vietnam, W Asia. CORRIGIDOR, PALAWAN. Native.
- ssp. laeta (Wall. ex Ettingsh.) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 100; ≡ Pteris laeta Wall. ex Ettingsh., Farnkr. Jetztw. (1865) 96. Distribution: Afghanistan, Bhutan, Cambodia, China, India, Japan, Laos, Lesser Sunda Isls, Middle East, Myanmar, Nepal, Pakistan, Philippines, Sri Lanka, Sulawesi, Taiwan, Tibet, Vietnam, W Asia. Native.
Pteris dataensis Copel., PJS 81 (1952) 7. Distribution: Endemic to the Philippines. LUZON: Benguet, Ifugao, Mountain Province, Nueva Vizcaya, MINDANAO: Bukidnon. Conservation status: Vulnerable (DAO 2017-11). Native.
Pteris distans J.Sm., J. Bot. 3 (1841) 405. Distribution: Endemic to the Philippines. LUZON: Bulacan, Rizal. Notes: Possibly a hybrid or a bizarre freak. Native.
Pteris edanoi Copel., PJS 81 (1952) 8, as 'edanyoi'. Distribution: Endemic to the Philippines. LUZON. Native.
Pteris elmeri Christ ex Copel., PJS 81 (1952) 7. Distribution: Endemic to the Philippines. LUZON: Benguet, Bulacan, Mountain Province, NEGROS. Native.
Pteris endoneura M.G.Price, Kew Bull. 29 (1975) 727; ≡ Copelandiopteris endoneura (M.G.Price) Stone, Kalikasan 5 (1976) 330. Distribution: Endemic to the Philippines. LUZON: Camarines Norte, Quezon, SAMAR. Conservation status: Endangered (DAO 2017-11). Native.
Pteris ensiformis Burm.f., Fl. Ind. (1768) 230. Distribution: ALABAT, BATAN, BATANES, BOHOL, GENERAL ISLAND, GUIMARAS, JOLO, LEYTE, LUZON: Batangas, Bulacan, Cagayan, Camarines Sur, Cavite, Ifugao, Ilocos Norte, Isabela, Laguna, NCR, Nueva Ecija, Quezon, Rizal, Sorsogon, MINDANAO: Agusan del Norte, Bukidnon, Davao, Davao del Sur, Surigao del Norte, Zamboanga del Sur, MINDORO, NEGROS, PANAY, SAMAR, SIBUYAN, TAWI-TAWI. Native.
- var. ensiformis. = Pteris mutilata auct. non L. Distribution: Bangladesh, Bhutan, Bismarck Arch, Borneo, Cambodia, China, India, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, Nepal, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam. Native.
- var. victoriae Baker, Gard. Chr. III, 7 (1890) 576. Distribution: LUZON: NCR. Notes: Probably best treated as a garden cultivar. Cultivated, not naturalized.
Pteris faba Y.S.Chao, New Zealand J. Bot. 60 (2022) 8; Type: Y.S. Chao 1405 (TAIF, holo; CMUH, TAIF, iso), Mindanao: Bukidnon, Mt Kitanglad. 17-Nov-2007. = Pteris amoena ssp. firma K.U.Kramer, ined. Distribution: New Guinea, Philippines. MINDANAO: Bukidnon, Davao del Sur. Conservation status: Data Deficient (Chao et al. 2022). Native.
Pteris glaucovirens Goldm. ex Hieron., Hedwigia 66 (1914) 339. Distribution: Borneo, Philippines. CAMIGUIN DE BABUYANES, LUZON: Bataan, Batangas, Benguet, Camarines Norte, Ilocos Norte, Kalinga, La Union, Laguna, Mountain Province, NCR, Pampanga, Quezon, Rizal, Zambales, MINDANAO: Bukidnon, Davao del Sur, Lanao del Sur, Zamboanga Sibugay, MINDORO, NEGROS (photos), PALAWAN. Native.
Pteris heteromorpha Fée, Mém. Foug. 5 (1852) 127. Distribution: Borneo, China, India, Laos, Myanmar, Philippines, Sulawesi, Thailand, Vietnam. LUZON: Bataan, Ilocos Norte, Laguna, NCR, Pampanga, Pangasinan, Rizal. Native.
Pteris khasiana (C.B.Clarke) Hieron., Hedwigia 55 (1914) 364; ≡ Pteris quadriaurita var. khasiana C.B.Clarke, Trans. Linn. Soc. London, Bot. 1 (1880) 466. Distribution: BATAN, LUZON: Aurora, Benguet, Ifugao, Laguna, Nueva Vizcaya. Native.
- ssp. fauriei (Hieron.) Fraser-Jenk., Annot. Checkl. Ind. Pterid. 1 (2016) 292; ≡ Pteris fauriei Hieron., Hedwigia 55 (1914) 345; Hatusima, Mem. Fac. Kagoshima Univ. 5 (1966) 20. = Pteris rigidula Copel., FFP 1 (1958) 137. Distribution: China, Japan, Korea, Pacific Ocean, Philippines, Ryukyu Isls, Taiwan, Vietnam. BATAN, LUZON: Aurora, Benguet, Ifugao, Laguna, Nueva Vizcaya. Native.
Pteris loheri Copel., PJS 81 (1952) 8. Distribution: Endemic to the Philippines. LUZON: Benguet, Kalinga. Native.
Pteris longipinnula Wall. ex J.Agardh, Recens. Spec. Pter. (1839) 19; Rufila et al., Ruhuna J. Science 13 (2022) 159. Distribution: Bangladesh, Borneo, China, India, Japan, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, Nepal, Philippines, Sulawesi, Thailand. LUZON: Albay, Ilocos Norte, Isabela, Laguna, Pampanga, Quezon, Rizal, Sorsogon, MINDANAO: Agusan del Norte, Davao del Sur, PALAWAN. Native.
Pteris luzonensis Hieron., Hedwigia 66 (1914) 370. Distribution: Endemic to the Philippines. LUZON: Cagayan, MINDORO. Native.
Pteris macgregorii Copel., PJS 81 (1952) 9. Distribution: Endemic to the Philippines. LUZON: Nueva Vizcaya. Conservation status: Vulnerable (DAO 2017-11). Native.
Pteris melanocaulon Fée, Gen. (1850-52) 127. Distribution: Philippines, Sulawesi. BASILAN, CORRIGIDOR, JOLO, LEYTE, LUZON: Albay, Apayao, Bulacan, Camarines Norte, Cavite, Laguna, Quezon, Rizal, Sorsogon, MINDANAO: Agusan del Norte, Davao del Norte, Surigao, Surigao del Sur, Zamboanga, Zamboanga del Sur, MINDORO, PALAWAN, PANAY, SAMAR. Native.
Pteris melanorhachis Copel., PJS 81 (1952) 9. Distribution: Endemic to the Philippines. LUZON: Camarines Norte, NEGROS. Native.
Pteris mertensioides Willd., Sp. Pl. (1810) 394. = Pteris gardneri (Fée) Hook., Sp. Fil. 2 (1858) 206; ≡ Litobrochia gardneri Fée, Gen. (1850-52) 139. = Pteris patens Hook., Sp. Fil. 2 (1858) 177, nom. illeg., non Kunze. Distribution: Andaman Isls, Bhutan, Borneo, India, Java, Malay Peninsula, Moluccas, Pacific Ocean, Philippines, Singapore, Sri Lanka, Sulawesi, Thailand. BOHOL, CAMIGUIN, CATANDUANES, LUZON: Batangas, Kalinga, Laguna, Mountain Province, Quezon, Rizal, Sorsogon, Zambales, MINDANAO: Bukidnon, Davao del Sur, Lanao del Sur, Surigao, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY. Native.
Pteris micracantha Copel., PJS 81 (1952) 10. Distribution: Endemic to the Philippines. LUZON: Abra, Benguet. Conservation status: Vulnerable (DAO 2017-11). Native.
Pteris mucronulata Copel., PJS 81 (1952) 10. Distribution: Endemic to the Philippines. MINDANAO: Bukidnon, NEGROS, PALAWAN. Conservation status: Vulnerable (DAO 2017-11). Native.
Pteris opaca (C.Presl) J.Sm. ex Fée, Mém. Foug., 5. Gen. Filic. 242 (1850) 358; ≡ Pycnodoria opaca C.Presl, Abh. Kön. Böhm. Ges. Wiss. 5(6) (Epim. Bot.) (1851) 101. = Pteris taenitis Copel., PJS 7 (1912) 63. Distribution: Borneo, Moluccas, New Guinea, Philippines, Sulawesi. BILIRAN, BOHOL, CEBU, LUZON: Camarines Norte, Kalinga, Quezon, Tayabas, MINDANAO: Agusan del Norte, Davao Oriental, Zamboanga, Zamboanga del Sur, PALAWAN, SAMAR. Conservation status: Vulnerable (DAO 2017-11 as Pteris taenitis). Native.
Pteris oppositipinnata Fée, Mém. Foug. 10 (1865) 17; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22048. = Pteris asperula J.Sm. ex Hieron., Hedwigia 66 (1914) 302. = Pteris caesia Copel., PJS 1 (Suppl. 2) (1906) 156. Distribution: Borneo, Philippines, Sulawesi. LUZON: Abra, Bataan, Benguet, Ilocos Sur, Laguna, Nueva Ecija, Pampanga, Pangasinan, Quezon, Rizal, Zambales, MINDANAO: Davao del Sur, South Cotabato, MINDORO, PALAWAN. Native.
Pteris pachysora (Copel.) M.G.Price, Kew Bull. 29 (1975) 727; ≡ Copelandiopteris pachysora (Copel.) Stone, Webbia 28 (1973) 493; ≡ Schizostege pachysora Copel., PJS 1 (Suppl. 2) (1906) 155. Distribution: Endemic to the Philippines. MINDANAO: Zamboanga, Zamboanga del Sur. Conservation status: Critically Endangered (DAO 2017-11). Native.
Pteris pedicellata Copel., PJS 81 (1952) 10. Distribution: Endemic to the Philippines. LUZON: Batangas, Benguet, Cagayan, Kalinga, Pampanga, MINDANAO: Lanao del Sur. Native.
Pteris pellucens J.Agardh, Rec. Sp. Gen. Pterid. (1839) 32. = Pteris philippinensis Fée, Mém. Foug. 8 (1857) 114; ≡ Pteris longipes D.Don var. philippinensis (Fée) C.Chr., Gard. Bull., S. S. 7 (1934) 286. = Pteris longipes D.Don, Prod. Fl. Nepal. (1825) 15; Copel., FFP 1 (1958) 133. Distribution: Bhutan, Borneo, China, India, Laos, Lesser Sunda Isls, Myanmar, Nepal, Philippines, Sulawesi, Taiwan, Thailand, Vietnam. BOHOL, LUZON: Laguna, PALAWAN. Native.
Pteris pellucida C.Presl, Reliq. Haenk. 1 (1825) 55. Distribution: Borneo, Java, Lesser Sunda Isls, New Guinea, Philippines. BOHOL, LUZON: Bataan, Batangas, Benguet, Laguna, Quezon, Sorsogon, MINDANAO: Davao Oriental, NEGROS. Native.
Pteris ramosii Copel., PJS 81 (1952) 11. Distribution: Endemic to the Philippines. LUZON: Abra. Notes: Possibly a variant of Pteris philippinensis. Conservation status: Vulnerable (DAO 2017-11). Native.
Pteris rangiferina C.Presl ex Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1868) 95. Distribution: Borneo, Java, Philippines. Native.
Pteris ryukyuensis Tagawa, Acta Phytotax. Geobot. 4 (1935) 204; Hatusima, Mem. Fac. Kagoshima Univ. 5 (1966) 20. = ?Pteris cretica auct. non L.; Copel., FFP 1 (1958) 129, quoad pl. ex Batanes Islands. Distribution: Japan, Philippines, Ryukyu Isls, Taiwan. Native.
Pteris schlechteri Brause, Bot. Jahrb. Syst. 49 (1912) 33. Distribution: Moluccas, New Guinea, Philippines, Sulawesi. LUZON: Kalinga, Laguna, Quezon, MINDANAO: Bukidnon, Cotabato, Davao, Davao del Sur, Sarangani, South Cotabato, NEGROS. Native.
Pteris semipinnata L., Sp. Pl. 2 (1753) 1076. Distribution: Bangladesh, Bhutan, Borneo, China, India, Korea, Laos, Moluccas, Myanmar, Nepal, Philippines, Ryukyu Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. BALABAC, LUZON: Bataan, Bulacan, Ilocos Norte, Nueva Vizcaya. Native.
Pteris setulosocostulata Hayata, Ic. Pl. Formos. 4 (1914) 241. Distribution: China, Japan, Philippines, Taiwan. LUZON: Benguet, Ilocos Norte, Laguna, SABTANG. Native.
Pteris squamipes Copel., PJS 81 (1952) 11. Distribution: Endemic to the Philippines. MINDANAO: Agusan del Norte, Bukidnon. Conservation status: Vulnerable (DAO 2017-11). Native.
Pteris tripartita Sw., J. Bot. (Schrader) 1800 (1801) 67. = Pteris microdictyon (Fée) Hook., Sp. Fil. 2 (1858) 228; ≡ Litobrochia microdictyon Fée, Gen. (1850-52) 138. Distribution: Africa, Andaman Isls, Australia, Bismarck Arch, Borneo, China, India, Indian Ocean, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, New Caledonia, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, ?Taiwan, Thailand, Vietnam. ALABAT, BATAN, BATANES, CATANDUANES, DINAGAT, JOLO, LEYTE, LUZON: Albay, Aurora, Bataan, Batangas, Benguet, Cagayan, Camarines Norte, Camarines Sur, Ilocos Norte, Laguna, Mountain Province, NCR, Nueva Ecija, Nueva Vizcaya, Pampanga, Quezon, Rizal, Sorsogon, MINDANAO: Agusan del Norte, Bukidnon, Davao del Sur, Lanao del Norte, Lanao del Sur, Sultan Kudarat, Zamboanga, Zamboanga del Norte, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, POLILLO, SAMAR, SIARGAO. Native.
Pteris vittata L. ex Hieron., Hedwigia 64 (1914) 290; Batuyong et al., Biodiversitas 21 (2020) 3975; Rufila et al., Ruhuna J. Science 13 (2022) 159. Distribution: Africa, Andaman Isls, Australia, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, Europe, India, Indian Ocean, Japan, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Middle East, Moluccas, Myanmar, Nepal, New Guinea, Nicobar Isls, Pakistan, Philippines, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, W Asia. BABUYAN, BATAN, BOHOL, CEBU, GUIMARAS, LEYTE, LUZON: Abra, Batangas, Benguet, Cavite, Ifugao, Ilocos, Ilocos Norte, La Union, Laguna, Mountain Province, NCR, Nueva Vizcaya, Pampanga, Pangasinan, Quezon, Rizal, MINDANAO: Bukidnon, Cotabato, Davao, Davao del Sur, Lanao del Norte, Zamboanga, Zamboanga del Sur, MINDORO, PALAWAN, PANAY, POLILLO. Native.
Pteris wallichiana J.Agardh, Rec. Sp. Gen. Pterid. (1839) 69. Distribution: Bhutan, Borneo, China, India, Japan, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Nepal, New Guinea, Philippines, Ryukyu Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Tibet, Vietnam. LUZON. Native.
Pteris whitfordii Copel., PJS 1 (Suppl. 4) (1906) 255. Distribution: Endemic to the Philippines. BOHOL, MINDANAO: Bukidnon, MINDORO, NEGROS. Native.

Possibly present

Pteris cadieri Christ, J. Bot. (Morot) 19 (1905) 72.
- ssp. cadieri. Distribution: Bangladesh, Cambodia, China, ?Philippines, Ryukyu Isls, Taiwan, Thailand, Vietnam.
Pteris grevilleana Wall. ex J.Agardh, Recens. Spec. Pter. (1839) 23.
- ssp. digitata (Baker) Fraser-Jenk., Annot. Checkl. Ind. Pterid. 1 (2016) 289; ≡ Pteris quadriaurita var. digitata Baker, J. Bot. (1879) 40. Distribution: Borneo, China, Myanmar, ?Philippines, Taiwan, Thailand, Vietnam.
Pteris keysseri Rosenst., Repert. Spec. Nov. Regni Veg. 12 (1913) 167. Distribution: New Guinea, ?Philippines, Solomon Isls. Notes: Reported by Chao et al. (2021) from the Philippines (Y.-S. Chao 1403, TAIF).
Pteris terminalis Wall. ex J.Agardh, Recens. Spec. Gen. Pterid. (1839) 20. = Pteris excelsa Gaudich., Voy. Uranie (1829) 388. Distribution: Bhutan, Bismarck Arch, Borneo, China, India, Japan, Korea, Lesser Sunda Isls, Malay Peninsula, Moluccas, Nepal, New Guinea, Pacific Ocean, Pakistan, ?Philippines, Solomon Isls, Sulawesi, Taiwan, Tibet, Vietnam.

Syngramma J.Sm.

Last edited by Julie F. Barcelona, 7 September 2020

Syngramma alismifolia (C.Presl) J.Sm., Lond. J. Bot. 4 (1845) 168; ≡ Diplazium alismifolium C.Presl, Rel. Haenk. 1 (1825) 49. Distribution: Borneo, Cambodia, Java, Malay Peninsula, ?New Guinea, Nicobar Isls, Philippines, Singapore, Sulawesi, Sumatra, Thailand, Vietnam. CAMIGUIN, CEBU, LEYTE, LUZON: Quezon, Sorsogon, MINDANAO: Agusan del Norte, Bukidnon, Davao Oriental, Davao del Sur, Lanao del Sur, Surigao del Sur, Zamboanga del Sur, MINDORO, NEGROS, SIBUYAN. Native.
Syngramma vittiformis (J.Sm.) J.Sm., Lond. J. Bot. 4 (1845) 168; ≡ Callipteris vittiformis J.Sm., J. Bot. (Hooker) 3 (1841) 409. Distribution: Borneo, Philippines, Sulawesi. LUZON: Batangas, Camarines, Camarines Norte, Camarines Sur, Isabela, Laguna, Quezon, MINDANAO: Surigao, Surigao del Sur, MINDORO, NEGROS, PANAY, SAMAR, SIBUYAN. Native.
Syngramma wallichii (Hook.) Bedd., Handb. Ferns Brit. India (1866) t.153; Rufila et al., Ruhuna J. Science 13 (2022) 159; ≡ Gymnogramma wallichii Hook., Sp. 5 (1864) 155. Distribution: Borneo, Malay Peninsula, Moluccas, Philippines, Sulawesi, Sumatra. LUZON: Aurora, Laguna, Quezon, MINDANAO: Davao del Sur, Surigao, SAMAR, SIBUYAN. Native.

Taenitis Willd.

Last edited by Julie F. Barcelona, 7 September 2020

Taenitis blechnoides (Willd.) Sw., Syn. Fil. 24 (1806) 220; ≡ Pteris blechnoides Willd., Phyt. (1794) 13. Distribution: Australia, Bismarck Arch, Borneo, Cambodia, China, ?India, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam. BALABAC, BASILAN, BILIRAN, DINAGAT, LEYTE, LUZON: Aurora, Bataan, Benguet, Cagayan, Camarines Norte, Camarines Sur, Cavite, Ilocos, Ilocos Norte, Isabela, La Union, Laguna, NCR, Quezon, Rizal, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Davao Oriental, Lanao del Norte, Surigao, Zamboanga, Zamboanga del Sur, NEGROS, PALAWAN, PANAY, POLILLO, SAMAR, SIBUYAN, TAWI-TAWI. Native.
Taenitis cordata (Gaudich.) Holttum, Amer. Fern J. 50 (1960) 112; ≡ Schizoloma cordatum Gaudich., Ann. Sci. Nat. 3 (1824) 507. Distribution: Philippines, Sulawesi. BOHOL, PONEAS, SAMAR. Conservation status: Vulnerable (DAO 2017-11). Native.

Vaginularia Fée

Last edited by Julie F. Barcelona, 7 September 2020

Vaginularia junghuhnii Mett., Fil. Hort. Bot. Lips. (1856) 25; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22048. = Vaginularia paradoxa auct. non (Fée) Mett; Rufila et al., Ruhuna J. Science 13 (2022) 159. = Monogramma paradoxa auct. non (Fée) Mett. Distribution: Borneo, Lesser Sunda Isls, Moluccas, New Guinea, Philippines, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand. LUZON: Camarines Sur (photos), Quezon (photos), MINDANAO: Bukidnon, Cotabato (photos), Davao del Sur, South Cotabato, NEGROS (photos). Native.
Vaginularia trichoidea Fée, Mém. Foug. 3 ((1851)) 54; ≡ Monogramma trichoidea (Fée) J.Sm., J. Bot. 3 (1841) 394. Distribution: Borneo, China, Java, Lesser Sunda Isls, Malay Peninsula, Moluccas, New Guinea, Philippines, Singapore, Sulawesi, Sumatra, Taiwan, Thailand. BILIRAN, CATANDUANES, LEYTE, LUZON: Albay, Apayao, Aurora, Batangas, Cagayan, Camarines Norte, Camarines Sur, Kalinga, Laguna, Nueva Ecija, Quezon, Sorsogon, MINDANAO: Agusan del Norte, Davao del Sur, Lanao del Sur, Surigao, Surigao del Norte, Zamboanga, Zamboanga Sibugay, Zamboanga del Sur, MINDORO, NEGROS, SAMAR. Native.

References

📋 Alston, AHG. 1959. Isoetaceae. Flora Malesiana series 2, 1: 63-64
📋 Amoroso, VB, FP Coritico & PW Fritsch. 2020. Actinostachys minuta, a new species of grass fern from Mindanao, Philippines. PhytoKeys 151: 59-66
📋 Amoroso, VB, YLL Cariño, JC Nobleza & FP Coritico. 2020. Ophioderma subsessile (Ophioglossaceae), a new snake tongue fern species from Mindanao, Philippines. Philippine Journal of Science 150 (S1): 215-221
Amoroso, VB, YLL Cariño, JC Nobleza & FP Coritico. 2021. A new species of Actinostachys (Schizaeaceae) from Mindanao, Philippines. Phytotaxa 508: 101-106.
Barcelona, JF & MG Price. 1999. A new Philippine Goniophlebium. Fern Gazette 15: 261-264.
Barcelona, JF & PB Pelser. 2014. Phanerosorus (Matoniaceae), a new fern genus record for the Philippines. Phytotaxa 170: 133-135.
📋 Barcelona, JF, B. Hernaez, & MG Price. 1996. Philippine Schizaea. Asia Life Sciences 5: 27-34
📋 Barcelona, JF, DL Nickrent, JV LaFrankie, JRC Callado & PB Pelser. 2013. Co’s Digital Flora of the Philippines: plant identification and conservation through cybertaxonomy. Philippine Journal of Science 142: 57-67
📋 Barcelona, JF, NE Dolotina, GS Madroñero, WG Granert & DD Sopot. 2006. The ferns and fern allies of the karst forests of Bohol Island, Philippines. American Fern Journal 96: 1-20
Barcelona, JF. 2002. Philippine pteridophyte collections as a resource for conservation planning. Proceedings of the “International Symposium on the Fern Flora Worldwide: Threats and Responses” sponsored by the British Pteridological Society and IUCN- Species Survival Commission.” Fern Gazette 16(6, 7 & 8): 307-312.
📋 Barcelona, JF. 2003. Preliminary report on the ferns and fern allies (pteridophytes) of Mt. Bali-it, Balbalasang-Balbalan National Park, Kalinga, Northern Luzon, Philippines. Sylvatrop: The Technical Journal of Philippine Ecosystems and Natural Resources 13: 81-92
Barcelona, JF. 2003. The taxonomy and ecology of the pteridophytes of Mt. Iraya & vicinity, Batan Island, Batanes Province, Northern Philippines, in Pteridology in the New Millenium (Chandra and Srivastava, eds). Kluwer Academy Publishers. The Netherlands, 299-325.
Barcelona, JF. 2004. Collection and conservation status of pteridophytes in Panay Island, Philippines. Philipp. Scient. 41:57-73.
📋 Barcelona, JF. 2005. Noteworthy fern discoveries in the Philippines at the turn of the 21st Century. Fern Gazette 17: 139-146
📋 Bosman, MTM. 1991. A monograph of the fern genus Microsorum (Polypodiaceae) including an attempt towards a reconstruction of the phylogenetic history of the microsoroids. Leiden Botanical Series 14: 1-161
📋 Brownsey PJ. 1987. A review of the fern genus Hypolepis (Dennstaedtiaceae) in the Malesian and Pacific regions. Blumea 32: 227-276
📋 CABI Invasive Species Compendium
📋 Chambers TC & PA Farrant. 2001. Revision of Blechnum (Blechnaceae) in Malesia. Blumea 46: 283-350
📋 Chang, Y-H, L-Y Kuo, W-L Chiou, C-W Chen, H-M Chang & VB Amoroso. 2013. Stenolepia Alderw. (Dryopteridaceae), a fern genus new to the Philippines. Philippine Journal of Science 142: 83-88
Chao, Y-S, C-W Chen, W-L Chiou, FP Coritico, Y-H Chang, & T-Y A Yang. 2022. Pteris faba and Pteris rubella, two new species in section Hypsopodium (Pteridaceae). New Zealand Journal of Botany: 1-13
📋 Chao, Y-S, W-L Chiou, A Ebihara, T-C Hsu, Y-H Chang & C-Y Lin. 2021. Taxonomic and nomenclatural novelties in the Pteris fauriei group (Pteridaceae). Taiwania 66: 307-316
📋 Chen, C-W, CJ Rothfels, AMA Mustapeng, M Gubilil, DK Karger, M Kessler & Y-M Huang. 2018. End of an enigma: Aenigmopteris belongs in Tectaria (Tectariaceae: Polypodiopsida). Journal of Plant Research 131: 67-76
📋 Chen, C-W, LT Ngan, A Hidayat, L Evangelista, HP Nooteboom & W-L Chiou. 2014. First insights into the evolutionary history of the Davallia repens complex. Blumea 59: 49-58
📋 Chen, C-W, S Lindsay, CM Mynssen, L-Y Kuo, VB Amoroso & Y-M Huang. 2018. A new Haplopteris species from the Philippines and clarification of the status of H. amboinensis. Garden's Bulletin Singapore 70: 177-190
📋 Chen, C-W, S Lindsay, KT Yong, AMA Mustapeng, VB Amoroso, VD Dang & Y-M Huang. 2019. Clarification of two poorly known vittarioid ferns (Pteridaceae): Haplopteris angustissima and H. capillaris. Systematic Botany 44: 483-493
Chen, C-W, SK Tang, R Knapp, MAM Andi, P-F Lu, M Kessler & Y-M Huang. 2021. Leucostegia amplissima, the third species of the genus (Hypodematiaceae). Systematic Botany 46: 656-665.
Christenhusz, MJM, X-C Zhang & H Schneider. 2011. A linear sequence of extant families and genera of lycophytes and ferns. Phytotaxa 19: 7-54.
Copeland, E.B. 1961. Fern Flora of the Philippines 3: 547-553.
📋 Coritico FP & VB Amoroso. 2020. Threatened lycophytes and ferns in four protected areas of Mindanao, Philippines. Nature Conservation Research 5: 78-88
📋 Coritico FP, VB Amoroso & YC Liu. 2019. Athyrium nakanoi Makino (Athyriaceae), a new record from the Philippines and an identification key to the Malesian Athyrium sect. Polystichoides. Philippine Journal of Systematic Biology 13: 46-50
📋 Coritico, FP, VB Amoroso & M Lehnert. 2017. New records, names and combinations of scaly tree ferns (Cyatheaceae) in eastern Malesia. Blumea 62: 92-96
📋 Coritico, FP, VB Amoroso, FM Acma, YLL Cariño & PW Fritsch. 2020. Ferns and lycophytes of Mt. Tago Range, Bukidnon, southern Philippines: species richness, distribution, and conservation status. Philippine Journal of Science 149: 773-790
📋 Coritico, FP, YLL Cariño, MMM Guiang & VB Amoroso. 2022. New record of Ophioderma redactophylla (Ophioglossaceae) in the Philippines and new insights to its morphology. Philippine Journal of Science 151: 227-233
📋 Crane, E.H. 1997. A revised circumscription of the genera of the fern family Vittariaceae. Systematic Botany 22: 509-517
📋 DENR Administrative Order. 2017. Updated national list of threatened Philippine plants and their categories. DAO 2017-11
📋 Delos Angeles, MD, IE Buot Jr, L Hongmei & H Schneider. 2022. Pteridophyte diversity in the Samar Island Natural Park (SINP), Samar Island, Philippines. Philippine Journal of Science 151: 1929-1942
Ding, H-H, Y-S Chao & S-Y Dong. 2013. Taxonomic novelties in the fern genus Tectaria (Tectariaceae). Phytotaxa 122: 61-64.
Dong, S-H. 2014. Nomenclatural novelties in the fern genus Tectaria (Tectariaceae). Phytotaxa 178: 225-228.
📋 Ebihara A, CR Fraser-Jenkins, BS Parris, XC Zhang, YH Yang, WL Chiou, HM Chang, S Lindsay, D Middleton, M Kato, TN Praptosuwiryo, VB Amoroso, JF Barcelona, RHG Ranil, CH Park, N Murakami & A Hoya. 2012. Rare and threatened pteridophytes of Asia 1. An enumeration of narrowly distributed taxa. Bulletin of the National Museum of Nature and Science, Series B. 38: 93-119
📋 Ebihara, A, J-Y Dubuisson, K Iwatsuki, S Hennequin, & M Ito. 2006. A taxonomic revision of the Hymenophyllaceae. Blumea 51: 221-280
📋 Field, AR, W Testo, PD Bostock, JAM Holtum & M Waycott. 2016. Molecular phylogenetics and the morphology of the Lycopodiaceae subfamily Huperzioideae supports three genera: Huperzia, Phlegmariurus and Phylloglossum. Molecular Phylogenetics and Evolution 94: 635-657
Gruezo, WS. 2012. Tmesipteris zamorae: a new species of Tmesipteris Bernh. (Psilotaceae) from the Philippines. Asia Life Sciences 21: 653-670.
📋 Hassler, M & B Schmitt. 2019. Checklist of ferns and lycophytes of the world
📋 Hassler, M & B Schmitt. World ferns: checklist of ferns and lycophytes of the world
Hassler, M. 2019. World ferns: checklist of ferns and lycophytes of the world. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
Hauke, RI. 1963. A taxonomic monograph of the genus Equisetum subgenus Hippochaete. Beih. Nova Hedwigia 8: 1-123.
📋 Hennipman, E & MC Roos. 1983. Phylogenetic Systematics of the Polypodiaceae (Filicales). Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversiteit te Utrecht 525: 321-342
📋 Hennipman, E & WLA Hetterscheid. 1984. The emendation of the fern genus Christiopteris, including the transference of two taxa to the microsorioid Polypodiaceae. Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversiteit te Utrecht 539: 1-10
📋 Hennipman, E. 1970. Some novelties in Bolbitis from Asia and the Pacific (Filices). Blumea 18: 147-149
📋 Hennipman, E. 1974. The lectotypification of Acrostichum acuminatum Willd. and the reinstatement of the name Photinopteris speciosa (Bl.) Presl. Blumea 22: 51-52
📋 Hennipman, E. 1977. A monograph of the fern genus Bolbitis (Lomariopsidaceae). Leiden Botanical Series 2: 1-329
📋 Hennipman, E. 1987. A new species of Lecanopteris (Polypodiaceae) from the Philippines. Blumea 32: 311-312
Holltum, RE. 1975. Studies in the family Thelypteridaceae. IX. The genus Sphaerostephanos in the Philippines. Kalikasan 4: 47-68.
📋 Holttum, RE & E Hennipman. 1978. Lomariopsis group. Flora Malesiana series 2, 1: 255-330
📋 Holttum, RE, U Sen & D Mittra. 1970. Studies in the family Thelypteridaceae II. A comparative study of the type-species of Thelypteris Schmidel, Cyclosorus Link, and Ampelopteris Kunze. Blumea 18: 195-215
📋 Holttum, RE. 1959. Cyatheaceae. Flora Malesiana series 2, 1: 65-176
📋 Holttum, RE. 1959. Gleicheniaceae. Flora Malesiana series 2, 1: 1-36
📋 Holttum, RE. 1959. Keys to the genera of Pteropsida. Flora Malesiana series 2, 1: 15-19
📋 Holttum, RE. 1959. Schizaeaceae. Flora Malesiana series 2, 1: 37-61
📋 Holttum, RE. 1966. The genera Lomariopsis, Teratophyllum, and Lomagramma in the islands of the Pacific and Australia. Blumea 14: 215-223
📋 Holttum, RE. 1966. The genus Arthropteris J.Sm. in Malesia. Blumea 14: 225-229
📋 Holttum, RE. 1968. A re-definition of the fern-genus Taenitis Willd. Blumea 16: 87-95
📋 Holttum, RE. 1969. Studies in the family Thelypteridaceae. The genera Phegopteris, Pseudophegopteris, and Macrothelypteris. Blumea 17: 5-32
📋 Holttum, RE. 1971. Studies in the Family Thelypteridaceae. III. A new system of genera in the Old World. Blumea 19: 17-52
📋 Holttum, RE. 1972. Studies in the family Thelypteridaceae IV. The genus Pronephrium Presl. Blumea 20: 105-126
📋 Holttum, RE. 1973. Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai. Blumea 21: 293-325
Holttum, RE. 1975. A comparative account of the fern-genera Syngramma J.Sm. and Taenitis Willd., with discussion of their relationships to each other and to other genera. Kew Bulletin 30: 327-343.
📋 Holttum, RE. 1975. Studies in the family Thelypteridaceae VIII. The genera Mesophlebion and Plesioneuron. Blumea 22: 223-250
📋 Holttum, RE. 1976. Studies in the family Thelypteridaceae X. The genus Coryphopteris. Blumea 23: 18-47
📋 Holttum, RE. 1977. Studies in the family Thelypteridaceae. XII. The genus Amphineuron Holttum. Blumea 23: 205-218
📋 Holttum, RE. 1981. Thelypteridaceae. Flora Malesiana Ser. 2, 1: 331-599
📋 Holttum, RE. 1984. Studies in the fern-genera allied to Tectaria. III. Aenigmopteris and Ataxipteris, two new genera allied to Tectaria Cav., with comments on Psomiocarpa Presl. Blumea 30: 1-11
📋 Holttum, RE. 1985. Studies in the fern genera allied to Tectaria Cav. IV. The genus Ctenitis in Asia, Malesia and the Western Pacific. Blumea 31: 1-38
📋 Holttum, RE. 1991. New taxa in the Tectaria Group (Polypodiaceae) from Malesia. Blumea 35: 547-557
📋 Holttum, RE. 1991. Tectaria group. Flora Malesiana series 2, 2: 1-132
📋 Holttum. RE. 1966. Florae Malesianae Precursores XLIV. The fern-genus Elaphoglossum in Malesia, with descriptions of new species. Blumea 14: 317-326
📋 Hovenkamp, PH & F Miyamoto. 2005. A conspectus of the native and naturalized species of Nephrolepis (Nephrolepidaceae) in the world. Blumea 50: 279-322
📋 Hovenkamp, PH & F Miyamoto. 2012. Nephrolepidaceae. Flora Malesiana series 2, 4: 97-122
📋 Hovenkamp, PH & H Boon-Chuan. 2012. A revision of the fern genus Oleandra (Oleandraceae) in Asia. PhytoKeys 11: 1-37
📋 Hovenkamp, PH & NAP Franken. 1993. An account of the fern genus Belvisia Mirbel (Polypodiaceae). Blumea 37: 511-527
📋 Hovenkamp, PH, MTM Bosman, E Hennipman, HP Nooteboom, G. Rodl-Linder & MC Roos. 1998. Polypodiaceae. Flora Malesiana series 2, 3: 1-234
📋 Hovenkamp, PH. 1986. A monograph of the fern genus Pyrrosia (Polypodiaceae). Leiden Botanical Series 9: 1-280
📋 Hovenkamp, PH. 1998. An account of the Malay-Pacific species of Selliguea (Polypodiaceae). Blumea 43: 1-108
📋 Hovenkamp, PH. 1998. Lepisorus in Malesia. Blumea 43: 109-115
📋 Hovenkamp, PH. 2019. Flora of Singapore precursors, 13. New names and lectotypifications in Athyriaceae and Polypodiaceae. Gard. Bull. Singapore 71: 61-67
Hsu, T-C, Y-F Huang & Y-S Chao. 2019. Systematics of Hymenophyllum subgenus Mecodium (Hymenophyllaceae) in Taiwan. Systematic Botany 44: 753-767.
Johns, RJ. 1995. Lecanopteris lomarioides: Polypodiaceae. Curtis's Botanical Magazine 12: 89-95.
📋 Karger, DN, S Lehtonen, VB Amoroso, M Kessler. 2012. A new species of Lindsaea (Lindsaeaceae, Polypodiopsida) from Mt. Hamiguitan, Mindanao, Philippines. Phytotaxa 56: 15-20
📋 Kato, M, Y Yatabe, N Sahashi & N Murakami. 2001. Taxonomic studies of Cheiropleuria (Dipteridaceae). Blumea 46: 513-525
📋 Kato, M. 1993. A taxonomic study of the genus Matonia (Matoniaceae). Blumea 38: 167-172
📋 Kato, M. 1998. Matoniaceae. Flora Malesiana series 2, 3: 289-294
Kinosian, SP, WD Pearse & PG Wolf. 2020. Cryptic diversity in the model fern genus Ceratopteris (Pteridaceae). Molecular Phylogenetics and Evolution 152: 106938.
📋 Kramer, KU. 1967. The Lindsaeoid ferns of the Old World II. A revision of Tapeinidium. Blumea 15: 545-556
📋 Kramer, KU. 1967. The Lindsaeoid ferns of the Old World III. Notes on Lindsaea and Sphenomeris in the Flora Malesiana area. Blumea 15: 557-574
📋 Kramer, KU. 1971. Lindsaea Group. Flora Malesiana series 2, 1: 177-254
📋 Laferrière, JE. 1998. Cheiropleuriaceae. Flora Malesiana series 2, 3: 285-286
📋 Laferrière, JE. 1998. Equisetaceae. Flora Malesiana series 2, 3: 287-288
📋 Lehnert, M & FP Coritico. 2018. The genus Dicksonia (Dicksoniaceae - Cyatheales) in western Malesia. Blumea 63: 268-278
📋 Lin, YX & R Viane. 2013. Aspleniaceae. In: Pp. 267–316 in Wu, ZY, PH Raven & DY Hong (eds.), Flora of China, Vol. 2-3 (Pteridophytes). Beijing: Science Press; St. Louis: Missouri Botanical Garden Press
📋 Lindsay, S, PH Hovenkamp & DJ Middleton. 2017. New combinations and typifications in Aglaomorpha (Polypodiaceae). Gardens' Bulletin Singapore 69: 149-155
📋 Liu, H-M, R-H Jiang, J Guo, P Hovenkamp, LR Perrie, L Shepherd, S Hennequin & H Schneider. 2013. Towards a phylogenetic classification of the climbing fern genus Arthropteris. Taxon 62: 688-700
📋 Liu, Y-C, CR Fraser-Jenkins, VB Amoroso & W-L Chiou. 2008. Athyrium erythropodum (Woodsiaceae, Pteridophyta), a new Philippine record. Blumea 53: 447-452
📋 Luong, TT, PH Hovenkamp & MSM Sosef. 2015. Revision of the fern genus Orthiopteris (Saccolomataceae) in Malesia and adjacent regions. PhytoKeys 53: 39–71
📋 Masuyama, S & Y Watano. 2010. Cryptic species in the fern Ceratopteris thalictroides (L.) Brongn. (Parkeriaceae). IV. Taxonomic revision. Acta Phytotaxonomica et Geobotanica 61: 75-86
📋 Moody, K. 1989. Weeds reported in rice in South and Southeast Asia. IRRI, Philippines
📋 Murdock, AG. 2008. A taxonomic revision of the eusporangiate fern family Marattiaceae, with description of a new genus Ptisana. Taxon 57: 737-755
📋 Nooteboom, HP. 1992. Notes on Davalliaceae I. The genera Araiostegia, Davallodes, Leucostegia, and Gymnogrammitis. Blumea 37: 165-187
📋 Nooteboom, HP. 1994. Notes on Davalliaceae II. A revision of the genus Davallia. Blumea 39: 151-214
📋 Nooteboom, HP. 1997. The microsoroid ferns (Polypodiaceae). Blumea 42: 261-395
📋 Nooteboom, HP. 1998. Davalliaceae. Flora Malesiana series 2, 3: 235-276
📋 Paclibar, GCB & ER Tadiosa. 2020 Plant species diversity and assessment in Quezon Protected Landscape, Southern Luzon, Philippines. Philippine Journal of Systematic Biology 4: 1-19
Panigrahi, G.1993. Parahemionitis, a new genus of Pteridaceae. American Fern Journal 83: 90-92.
📋 Parris, BS. 2007. Five new genera and three new species of Grammitidaceae (Filicales) and the re-establishment of Oreogrammitis. Garden's Bulletin Singapore 59: 233-274
📋 Parris, BS. 2009. New genera of Malesian Grammitidaceae (Monilophyta). Blumea 54: 217-219
Parris, BS. 2021. Calymmodon pectinatus, a new species of grammitid fern (Polypodiaceae) from Malesia. Fern Gazette 21: 254-259.
📋 Perrie, LR, AR Field, DJ Ohlsen & PJ Brownsey. 2021. Expansion of the fern genus Lecanopteris to encompass some species previously included in Microsorum and Colysis (Polypodiaceae). Blumea 66: 242-248
📋 Petchsri, S, T Boonkerd, BR Baum, D Karladee, S Suriyong & A Lungkaphin. 2012. Phenetic study of the Microsorum punctatum complex (Polypodiaceae). ScienceAsia 38: 1-12
📋 Picasa albums of Taiwanese pterophytes and gynosperms
Price, MG. 1990. Philippine fern notes. Contr. Univ. Mich. Herb. 17: 267-278.
📋 Ravensberg, WJ & E Hennipman. 1986. The Pyrrosia species formerly referred to Drymoglossum and Saxiglossum (Filicales, Polypodiaceae). Leiden Botanical Series 9: 281-310
Reid, JD, GM Plunkett, GA Peters. 2006. Phylogenetic relationships in the heterosporous fern genus Azolla (Azollaceae) based on DNA sequence data from three noncoding regions. International Journal of Plant Sciences 167(3): 529-538.
📋 Rolleri, C. 1993. Revision of the genus Christensenia. American Fern Journal 83: 3-19
📋 Roos, MC. 1985. New combinations in Aglaomorpha Schott (Polypodiaceae). Blumea 31: 153
📋 Ruhfel, B, S Lindsay & CC Davis. 2008. Phylogenetic placement of Rheopteris and Monogramma (Pteridaceae, s.l.): evidence from Rbcl sequence data. Systematic Botany 33: 37-43
📋 Rödl-Linder, G. 1994. A monograph of the fern genus Thylacopteris (Polypodiaceae). Blumea 39: 351-364
📋 Rödl-Linder. G. 1990. A monograph of the fern genus Goniophlebium (Polypodiaceae). Blumea 34: 277-423
📋 Salgado, AE. 2017. Does Asplenium falcatum exist in the Philippines? Edinburgh Journal of Botany 74: 1-4
Salgado, AE. 2021. Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524: 71-84.
📋 Salgado, AE. 2022. Taxonomic and nomenclatural Notes on Philippine ferns. V. On the identity of Asplenium lepturus, A. militare, and A. nigrescens (Polypodiales, Aspleniaceae. Philippine Journal of Science 151: 291-301
📋 Saunders, RMK. 1998. Azollaceae. Flora Malesiana series 2, 3: 277-284
📋 Schuettpelz, E & KM Pryer. 2007. Fern phylogeny inferred from 400 leptosporangiate species and three plastid genes. Taxon 56: 1037-1050
📋 Schuettpelz, E, H Schneider, L Huiet, MD Windham & KM Pryer. 2007. A molecular phylogeny of the fern family Pteridaceae: assessing overall relationships and the affinities of previously unsampled genera. Molecular Phylogenetics and Evolution 44:1172-1185
📋 Senterre B, G Rouhan, C Morel & JY Dubuisson. 2017. New species and records in the fern genus Didymoglossum for the flora of Seychelles, with notes on the Southeast Asian D. motleyi complex (Hymenophyllaceae). Phytotaxa 292: 201-217
Shang, H, Z-L Liang & L-B Zhang. 2021. Taxonomy of the fern genus Didymochlaena (Didymochlaenaceae) from Asia and Pacific islands based on morphological and molecular evidence with the description of four new species and one new status. Phytotaxa 479: 71-82.
📋 Smith AR, Pryer KM, Schuettpelz E, Korall P, Schneider H & Wolf PG. 2006. A classification for extant ferns. Taxon 55: 705–731
📋 TPG. 2019. Updating Taiwanese pteridophyte checklist: a new phylogenetic classification. Taiwania 64: 367-395
📋 Tan, BC. 2013. The Selaginella flora, a good indicator of the Philippine pleistocene island groups. Philippine Journal of Science 142: 211-213
📋 Testo, WL, AR Field, AB Sessa & M Sundue. 2019. Phylogenetic and morphological analyses support the resurrection of Dendroconche and the recognition of two new genera in Polypodiaceae subfamily Microsoroideae. Systematic Botany 44: 1-16
Tryon. 1941. A revision of the genus Pteridium. Rhodora 43: 1-70.
📋 Wang, LI, Z-Q Wu, QP Xiang, J Heinrichs, H Schneider & XC Zhang. 2010. A molecular phylogeny and a revised classification of tribe Lepisoreae (Polypodiaceae) based on an analysis of four plastid DNA regions. Botanical Journal of the Linnean Society 162: 28-38
📋 Wardani, W & B Adjie. 2018. The genus Athyrium Roth in Malesia: a checklist with working names and distributions. Sibbaldia 16: 39-48
📋 Wieffering, JH. 1964. A preliminary revision of the Indo-Pacific species of Ophioglossum (Ophioglossaceae). Blumea 12: 321-337
Yesilyurt, JC & H Schneider. 2010. The new fern genus Calciphilopteris (Pteridaceae). Phytotaxa 7: 52-59.
📋 Zhang XC & HP Nooteboom. 1997. Some new combinations in Plagiogyria (Plagiogyriaceae, Pteridophyta). Blumea 42: 483-484
📋 Zhang XC & HP Nooteboom. 1998. A taxonomic revision of Plagiogyriaceae (Pteridophyta). Blumea 43: 401-469
📋 Zhang XC & HP Nooteboom. 1998. Plagiogyriaceae. Flora Malesiana series 2, 3: 295-316
📋 Zhang, L-B & L Zhang. 2012. The inclusion of Acrophorus, Diacalpe, Nothoperanema, and Peranema in Dryopteris: The molecular phylogeny, systematics, and nomenclature of Dryopteris subg. Nothoperanema (Dryopteridaceae). Taxon, 61: 1199-1216
📋 Zhang, L-B. 2012. Reducing the fern genus Dryopsis to Dryopteris and the systematics and nomenclature of Dryopteris subgenus Erythrovariae section Dryopsis (Dryopteridaceae). Phytotaxa 71: 17-27.
📋 Zhang, XC. 2003. New combinations in Haplopteris (Pteridophyta: Vittariaceae). Annales Botanici Fennici 40: 459-461
📋 Zhou, S, W Dong, X Chen, X Zhang, J Wen & H Schneider. 2014. How many species of bracken (Pteridium) are there? Assessing the Chinese brackens using molecular evidence. Taxon 63: 509-521
Zhou, X-M & L-B Zhang. 2019. A monograph of the fern genus Pteridrys (Pteridryaceae). Systematic Botany 44 (2019) 243-273.