Myristicaceae

Endocomia W.J.de Wilde

Last edited by Pieter B. Pelser, 11 November 2017
  1. Endocomia macrocoma (Miq.) W.J.de Wilde, Blumea 30 (1984) 182; Blumea 41 (1996) 375; Fl. Males. ser. 1, 14 (2000) 32; ≡ Myristica macrocoma Miq., Ann. Mus. Bot. Lugduno-Batavi 1 (1864) 207. Distribution: BOHOL, CAMIGUIN, CEBU (photos), LEYTE, LUZON: Camarines Norte, Camarines Sur, Laguna, Quezon, MINDORO, NEGROS, PANAY, SAMAR, TABLAS. Notes: 0-1000m. Conservation status: Least Concern (IUCN 3.1). Native.
    • ssp. macrocoma. Distribution: Moluccas, Philippines, ?Sulawesi. LUZON. Notes: According to De Wilde (2000), the Palanan material strongly deviates in its very hairy appearance, and may represent a separate taxon. Forest with little undergrowth, on alluvial flats locally with stagnant water, or hill slopes; on porous volcanic soil, or loam soil with stones; also in disturbed forest. Fl. & fr. throughout the year. 0-400m. Native.
    • ssp. prainii (King) W.J.de Wilde, Blumea 30 (1984) 187; Fl. Males. Ser. 1, 14 (2000) 35; ≡ Myristica prainii King, Ann. Roy. Bot. Gard. (Calcutta) 3 (1891) 299. = Horsfieldia oblongata Merr., PJS 13c (1918) 286; Merrill, EPFP 2 (1923) 182. = Horsfieldia merrillii Warb., Fragm. Fl. Philipp. 1 (1904) 49. Distribution: Andaman Isls, Bangladesh, China, India, Indochina, Java, Myanmar, New Guinea, Philippines, Sumatra. BOHOL, CAMIGUIN, LEYTE, LUZON: Camarines Norte, Camarines Sur, Laguna, Quezon, MINDORO, NEGROS, PANAY, SAMAR, TABLAS. Notes: Lowland and hillside forest, riverine or swamp forest; by streams, on alluvial or clayey soil, in limestone country, or on copper-rich soil. Fl. & fr. throughout the year, but in Philippines mainly Mar-May. 0-500m. Native.

Gymnacranthera (A.DC.) Warb.

Last edited by Pieter B. Pelser, 16 January 2022
  1. Gymnacranthera farquhariana (Wall. ex Hook.f. & Thomson) Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 365; R.T.A.Schouten, Blumea 31 (1986) 476; De Wilde, Fl. Males. Ser. 1, 14 (2000) 44; ≡ Myristica farquhariana Wall. ex Hook.f. & Thomson, Fl. Ind. (Hooker f. & Thomson) 1 (1855) 161. Distribution: Bismarck Arch, Borneo, Malay Peninsula, Moluccas, New Guinea, Philippines, Sulawesi, Sumatra, Thailand. BASILAN, BUCAS GRANDE (photos), CALAYAN, LEYTE, LUZON: Albay, Aurora, Bataan, Benguet (photos), Cagayan, Camarines, Ilocos Norte, Isabela, Laguna, Nueva Vizcaya, Quezon, Rizal, Sorsogon, MINDANAO: Agusan del Norte, Surigao del Norte, Zamboanga, MINDORO, NEGROS, PALAWAN, PANAY, SAMAR, SIBUYAN, SULU ARCHIPELAGO, TABLAS, TAWI-TAWI, TICAO. Notes: 0-1400m. Conservation status: Least Concern (IUCN 3.1). Native.
    • var. paniculata (A.DC.) R.T.A.Schouten, Blumea 31 (1986) 481; De Wilde, Fl. Males. Ser. 1, 14 (2000) 47; ≡ Gymnacranthera paniculata (A.DC.) Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 370; Merrill, EPFP 2 (1923) 181; ≡ Myristica paniculata A.DC., Ann. Sci. Nat. (Paris), 4th Sér. 4 (1855) 31. = Gymnacranthera acuminata Merr., PJS 12c (1917) 265; Merrill, EPFP 2 (1923) 181. = Gymnacranthera macrobotrys Merr., PJS 13c (1918) 284; Merrill, EPFP 2 (1923) 181. Distribution: Endemic to the Philippines. BASILAN, CALAYAN, LEYTE, LUZON: Albay, Aurora, Bataan, Cagayan, Camarines, Ilocos Norte, Isabela, Laguna, Nueva Vizcaya, Rizal, Sorsogon, MINDANAO: Agusan del Norte, Surigao del Norte, Zamboanga, MINDORO, NEGROS, PANAY, SAMAR, SIBUYAN, TABLAS, TICAO. Notes: Forests, on ridges, as well as by rivers and lakes, common. 0-1400m. Native.
    • var. zippeliana (Miq.) R.T.A.Schouten, Blumea 31 (1986) 482; De Wilde, Fl. Males. Ser. 1, 14 (2000) 47; ≡ Myristica zippeliana Miq., Ann. Mus. Bot. Lugduno-Batavi 2 (1865) 50. = Gymnacranthera suluensis Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 373; Merrill, EPFP 2 (1923) 181; Type: Vidal 3546 (syn); Vidal 3561 (syn), Basilan. Distribution: Bismarck Arch, Malay Peninsula, New Guinea, Philippines. BASILAN, SULU ARCHIPELAGO. Notes: 0-1400m. Native.

Horsfieldia Willd.

Last edited by Pieter B. Pelser, 15 February 2024
  1. Horsfieldia ardisiifolia (A.DC.) Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 274; De Wilde, Gard. Bull. Sing. 28 (1975) 3; Gard. Bull. Sing. 38, 1 (1985) 72; Fl. Males. ser. 1, 14 (2000) 90; ≡ Myristica ardisiifolia A.DC., Ann. Sci. Nat. (Paris), 4th Sér. 4 (1855) 31. = Horsfieldia warburgiana Elmer, LPB 3 (1911) 1061; Merrill, EPFP 2 (1923) 183; Type: Elmer 12297. = Horsfieldia gigantifolia Elmer, LPB 9 (1925) 3120, nom. inval., nom. nud. Distribution: Endemic to the Philippines. BASILAN, CEBU, LEYTE, LUZON: Nueva Vizcaya, Quezon, Sorsogon, MINDANAO: Misamis Occidental, MINDORO, PALAWAN, PANAY (photos), POLILLO, SAMAR, SIBUYAN, SULU ARCHIPELAGO, TABLAS. Notes: Forest in moist valleys. Fl. & fr. throughout the year. 0-600m. Conservation status: Vulnerable (IUCN 3.1). Native.
  2. Horsfieldia costulata (Miq.) Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 350; De Wilde, Gard. Bull. Sing. 39 (1986) 38; Fl. Males. ser. 1, 14 (2000) 105; Buhay et al., Biodiversitas 26 (2025) 3086; ≡ Myristica costulata Miq., Ann. Mus. Bot. Lugduno-Batavi 2 (1865) 48. = Horsfieldia megacarpa Merr., PJS 13c (1918) 286; Merrill, EPFP 2 (1923) 182; Type: BS 16527 Ramos. = Horsfieldia confertiflora Merr., PJS 13c (1918) 285; Merrill, EPFP 2 (1923) 182; Type: FB 3183 Ahern’s Collector. Distribution: Philippines, Sulawesi. BASILAN, LEYTE, LUZON: Aurora, Bataan, Cagayan, Camarines, Ilocos Norte (photos), Isabela (photos), Laguna, Quezon, Rizal, Sorsogon, MINDANAO: Agusan del Norte, Davao Oriental, Misamis, Zamboanga Sibugay, MINDORO, NEGROS, PANAY, SAMAR. Notes: Mixed rain forest, primary dipterocarp forest; recorded from alluvial soil and volcanic soil, with Eucalyptus deglupta dominance. 50-1200m. Conservation status: Least Concern (IUCN 3.1). Native.
  3. Horsfieldia irya (Gaertn.) Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 317; J.Sinclair, Gard. Bull. Sing. 16 (1958) 382; Gard. Bull. Sing. 28 (1975) 61; De Wilde, Gard. Bull. Sing. 38, 1 (1985) 55; Fl. Males. ser. 1, 14 (2000) 132; ≡ Myristica irya Gaertn., Fruct. Sem. Pl. 1 (1788) 195. = Horsfieldia acuminata Merr., PJS 17 (1921) 253; Merrill, EPFP 2 (1923) 181; Type: FB 27507 de Mesa. Distribution: Andaman Isls, Borneo, Cambodia, Java, Malay Peninsula, Moluccas, Myanmar, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam. MINDANAO: Lanao, PALAWAN, SIBUYAN. Notes: 0-550m. Conservation status: Least Concern (IUCN 3.1). Native.
  4. Horsfieldia obscura W.J.de Wilde, Gard. Bull. Singapore 39 (1986) 44; Fl. Males. ser. 1, 14 (2000) 153; Type: from Borneo. Distribution: Borneo, Philippines. PALAWAN. Notes: 150-730m. Conservation status: Vulnerable (IUCN 3.1). Native.
  5. Horsfieldia obscurinervia Merr., PJS 12c (1917) 265; Merrill, EPFP 2 (1923) 182; De Wilde, Gard. Bull. Sing. 38, 1 (1985) 71; Fl. Males. Ser. 1, 14 (2000) 154; Type: FB 26503 de Mesa & Magistrado. = Horsfieldia ramosii Merr., PJS 17 (1921) 254; Merrill, EPFP 2 (1923) 182; Type: BS 35047 Ramos. Distribution: Endemic to the Philippines. BUCAS GRANDE, LUZON: Camarines, Camarines Norte, SAMAR. Notes: Low hills. Fl. Jul; fr. Nov-Dec. 20m. Conservation status: Endangered (IUCN 3.1). Native.
  6. Horsfieldia polyspherula (Hook.f.) J.Sinclair, Gard. Bull. Singapore 16 (1958) 422; Gard. Bull. Sing. 28 (1975) 101; De Wilde, Gard. Bull. Sing. 39 (1986) 17; Blumea 41(1996) 381; Fl. Males. ser. 1, 14 (2000) 167; ≡ Myristica polyspherula Hook.f., Fl. Brit. India 5 (1886) 108. Distribution: LUZON: Isabela, MINDANAO: Agusan, MINDORO, PALAWAN. Notes: 0-1100m. Conservation status: Least Concern (IUCN 3.1). Native.
    • var. polyspherula. Distribution: Borneo, Malay Peninsula, Philippines, Singapore, Sumatra. LUZON: Isabela, MINDANAO: Agusan, PALAWAN. Notes: Fl. & fr. throughout the year, but most flowering collections in Jun-Sep. 0-900m. Native.
  7. Horsfieldia romblonensis W.J.de Wilde, Blumea 47 (2002) 355; Type: PP1 (Stone, Reynoso & Fernando) 6782 (L, holo; PNH, iso), Romblon. Distribution: Endemic to the Philippines. ROMBLON, SIBUYAN. Notes: Lowland forest on red-brown soil over ultra-basic. Native.
  8. Horsfieldia samarensis W.J.de Wilde, Gard. Bull. Singapore 38 (1985) 76; Fl. Males. ser. 1, 14 (2000) 182; Type: PNH 147374 Gutierrez et al. (L, holo), Samar: Eastern Samar, Mt Apoy, 600-800ft, May-1969. Distribution: Endemic to the Philippines. SAMAR. Notes: L.L.Co: Known only from the type collection. 200-270m. Conservation status: Critically Endangered (IUCN 3.1). DAO category: Vulnerable. Native.

Knema Lour.

Last edited by Pieter B. Pelser, 31 January 2025
  1. Knema alvarezii Merr., PJS 13c (1918) 288; Merrill, EPFP 2 (1923) 183; De Wilde, Blumea 25 (1979) 437; Fl.Males. Ser. 1, 14 (2000) 243; Type: FB 22395 Alvarez (L, iso); FB 22397 (US, isosyn), Luzon: Nueva Ecija, Mt Macasandal. 10-Feb-1911. Distribution: Endemic to the Philippines. LUZON: Cavite, Isabela, Nueva Ecija, SAMAR, SIARGAO. Notes: Lower and montane forests, possibly on ultramafics. Fl. Feb.; fr. May. L.L.Co: Known only from two collections. 0-850m. Conservation status: Vulnerable (IUCN 3.1). Native.
  2. Knema cinerea (Poir.) Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 611; J.Sinclair, Gard. Bull. Sing. 18 (1961) 169; De Wilde, Blumea 25 (1979) 457; Blumea 32 (1987) 129; Fl. Males. Ser. 1, 14 (2000) 249; ≡ Myristica cinerea Poir., Encycl. (Lamarck) Suppl. 4 (1816) 35; Type: from Pulau Buton, SE off Sulawesi. Distribution: Lesser Sunda Isls, Moluccas, Philippines, Sulawesi. MINDANAO: Zamboanga. Notes: L.L.Co: Known in the Philippines only from a single collection: FB 23328. 0-1000m. Conservation status: Least Concern (IUCN 3.1). Native.
  3. Knema glomerata (Blanco) Merr., J. Straits Branch Roy. Asiat. Soc. 76 (1917) 81; Merrill, EPFP 2 (1923) 183; J.Sinclair, Gard. Bull. Sing. 18 (1961) 219, pro parte excl. syn. Knema stenocarpa Warb; De Wilde, Blumea 25 (1979) 437; Blumea 32 (1987) 125; Fl. Males. Ser. 1, 14 (2000) 267; ≡ Sterculia glomerata Blanco, Fl. Filip., 1st Ed. (1837) 764; Type: Merrill Sp. Blancoanae 504 (neo). = Knema acuminata Merr., PJS 17 (1920) 256; Merrill, EPFP 2 (1923) 183; Type: FB 11266 Klemme. = Knema gitingensis Elmer, LPB 3 (1911) 1065. = Knema vidalii Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 615. ≠ Knema glauca auct. non (Blume) Warb.; Merr., EPFP 2 (1923) 183, pro parte. Distribution: Borneo, Moluccas, Philippines. ALABAT, BASILAN, BATAN, BILIRAN, CALAYAN, CAMIGUIN DE BABUYANES, CATANDUANES, CEBU, GUIMARAS, LEYTE, LUZON: Albay, Apayao, Aurora, Bataan, Batangas, Benguet, Bulacan, Cagayan, Camarines Norte, Camarines Sur, Cavite, Ilocos Norte, Isabela, Laguna, Nueva Ecija, Nueva Vizcaya, Quezon, Rizal, Sorsogon, Tarlac, Zambales, MARINDUQUE, MASBATE, MINDANAO: Agusan del Norte, Bukidnon (photos), Cotabato, Davao Oriental, Davao del Sur, Lanao, Lanao del Sur, Surigao, Surigao del Norte, Zamboanga Sibugay, Zamboanga del Norte, Zamboanga del Sur, MINDORO, NEGROS, PALAUI, PALAWAN, PANAY, POLILLO, SAMAR, SIBUYAN, TABLAS, TAWI-TAWI, TICAO. Notes: Common in primary and degraded lowland and medium elevation forests. Fl. throughout the year; fr. Jun-Dec. 0-600m. Conservation status: Least Concern (IUCN 3.1). Native.
  4. Knema korthalsii Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 557; J.Sinclair, Gard. Bull. Sing. 18 (1961) 232; De Wilde, Blumea 25 (1979) 428; Blumea 43 (1998) 242; Fl. Males. Ser. 1, 14 (2000) 273; Type: from Borneo (Kalimantan). = Knema insularis Merr., PJS 30 (1926) 394; Type: BS 44288 Ramos & Edaño (NY, US, iso), Tawi-tawi: damp forest at low elevation, 24-Jul-1924. = Knema cenabrei Merr. & Quisumb., PJS 37 (1928) 144; Type: FB 29973 Cenabre (NY, iso), Dumaran Is, along Linuatan Creek, damp forest at low elevation, 28-Jul-1925. ≠ Knema mindanaensis auct. non (Warb.) Merr.; Merr., EPFP 2 (1923) 184, pro parte excl. type. Distribution: Borneo, Philippines. BALABAC, BASILAN, DUMARAN, MINDANAO: Lanao del Sur, Misamis, Zamboanga, PALAWAN, SULU ARCHIPELAGO, TAWI-TAWI, TURTLE IS. Notes: Lowland forests. Fl. May-Jun; fr. May-Oct. 0-900m. Conservation status: Least Concern (IUCN 3.1). Native.
  5. Knema kunstleri (King) Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 568; De Wilde, Fl. Males. Ser. 1, 14 (2000) 277; ≡ Myristica kunstleri King, Ann. Roy. Bot. Gard. (Calcutta) 3 (1891) 314; Type: from Peninsular Malaysia. Distribution: Borneo, Malay Peninsula, Philippines, Sumatra. LUZON: Albay, Camarines, Camarines Norte, Isabela, MINDORO, POLILLO, SAMAR, SIBUYAN. Notes: 0-2000m. Conservation status: Least Concern (IUCN 3.1). Native.
    • ssp. parvifolia (Merr.) W.J.de Wilde, Blumea 25 (1979) 470; Blumea 43 (1998) 244; Fl. Males. Ser. 1, 14 (2000) 277; ≡ Knema parvifolia Merr., PJS 13c (1918) 287; Merrill, EPFP 2 (1923) 184; Type: FB 10573 Curran, Luzon: Albay, Manito. Forests, c. 30m. Distribution: Endemic to the Philippines. LUZON: Albay, Camarines, Camarines Norte, Isabela, POLILLO, SAMAR. Notes: Primary dipterocarp forest. Fl. Jun, Nov-Dec.; fr. Mar-May. 0-300m. Native.
  6. Knema latericia Elmer, LPB 5 (1913) 1815; Merrill, EPFP 2 (1923) 183; J.Sinclair, Gard. Bull. Sing. 16 (1958) 278; Gard. Bull. Sing. 18 (1961) 238; De Wilde, Blumea 25 (1979) 395; Blumea 43 (1998) 245; Fl. Males. Ser. 1, 14 (2000) 284; Type: Elmer 12757, Palawan. Distribution: MINDORO, PALAWAN. Notes: 0-1500m. Conservation status: Least Concern (IUCN 3.1). Native.
    • ssp. latericia. Distribution: Endemic to the Philippines. PALAWAN. Native.
      • var. latericia. Distribution: Endemic to the Philippines. PALAWAN. DAO category: Vulnerable. Native.
      • var. subtilis W.J.de Wilde, Blumea 32 (1987) 118; Blumea 41 (1996) 388; Fl. Males. Ser. 1, 14 (2000) 287; Type: Ridsdale SMHI 1642, Palawan. Distribution: ?Borneo, Philippines. PALAWAN. Notes: Primary and logged ridges in dipterocarp forests, once recorded from limestone. Fl. & fr. Apr. 0-100m. DAO category: Vulnerable. Native.
  7. Knema ridsdaleana W.J.de Wilde, Blumea 41 (1996) 392; Fl. Males. Ser. 1, 14 (2000) 335; Type: Ridsdale ISU 451 (PNH, holo; L, iso), Luzon: Isabela, Palanan, Digallorin (Digollorin), Divinisa Camp site, 16 50’N, 122 26’E, forest on ultrabasic, streamside forest, 50m, 9-Apr-1992. Distribution: Endemic to the Philippines. LUZON: Isabela. Notes: L.L.Co: Known only from the type collection. Streamside forest over ultramafics. 50m. Conservation status: Critically Endangered (IUCN 3.1). DAO category: Critically Endangered. Native.
  8. Knema stellata Merr., PJS 11c (1916) 182; Merrill, EPFP 2 (1923) 184; De Wilde, Blumea 25 (1979) 471; Blumea 27 (1981) 234; Fl. Males. Ser. 1, 14 (2000) 343; Type: BS 24276 Ramos, Samar. Distribution: Philippines, Sulawesi. CATANDUANES, LUZON: Cagayan, Camarines Norte, MINDANAO: Agusan del Norte, Surigao, Surigao del Norte, MINDORO, PANAY, SAMAR, SIBUYAN, TABLAS. Notes: 250-1000m. Native.
    • var. stellata. Distribution: Endemic to the Philippines. SAMAR. Notes: Forests at low elevation. Fr. Feb-May. Native.
    • ssp. cryptocaryoides (Elmer) W.J.de Wilde, Blumea 25 (1979) 471; Fl. Males. Ser. 1, 14 (2000) 344; ≡ Gymnacranthera cryptocaryoides Elmer, LPB 3 (1911) 1060; J.Sinclair, Gard. Bull. Sing. 16 (1958) 115; Type: Elmer 12262, Sibuyan. = Knema kunstleri (King) Warb. var. sheurigaoensis J.Sinclair, Gard. Bull. Singapore 18 (1961) 238; ≡ Myristica kunstleri King, Ann. Roy. Bot. Gard. (Calcutta) 3 (1891) 314; Type: Wenzel 2680, Mindanao. Distribution: Endemic to the Philippines. CATANDUANES, LUZON: Cagayan, Camarines Norte, MINDANAO: Agusan del Norte, Surigao, Surigao del Norte, PANAY, SIBUYAN, TABLAS. Notes: Lowland forests on fertile soil. Fl. Sep-Oct; fr. Feb-Apr. 250m. Native.
  9. Knema stenocarpa Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 577; Merrill, EPFP 2 (1923) 184; De Wilde, Blumea 25 (1979) 436; Fl. Males. ser. 1, 14 (2000) 345; Type: Warburg 13303 (syn); Warburg 13303a (syn), Mindanao. Distribution: Endemic to the Philippines. BASILAN, MINDANAO: Davao, Davao del Sur, Lanao, Misamis Occidental, Zamboanga Sibugay, Zamboanga del Norte, Zamboanga del Sur, TAWI-TAWI. Notes: Only the fruiting collection, not the male specimens, mentioned by Warburg are Knema stenocarpa Warb. Primary and degraded forests. Fl. & fr. throughout the year. 300-500m. Conservation status: Near Threatened (IUCN 3.1). DAO category: Other Threatened Species. Native.
  10. Knema tomentella (Miq.) Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 588; J.Sinclair, Gard. Bull. Sing. 18 (1961) 277; De Wilde, Blumea 25 (1979) 438; Blumea 41 (1996) 394; Fl. Males. Ser. 1, 14 (2000) 351; ≡ Myristica corticosa (Lour.) Hook.f. & Thomson var. tomentella Miq., Ann. Mus. Bot. Lugduno-Batavi 1 (1864) 207; Type: from Moluccas (Ambon). Distribution: Moluccas, New Guinea, Philippines, Sulawesi. LUZON: Nueva Ecija, SAMAR. Notes: 0-1500m. Native.

Myristica Gronov.

Last edited by Pieter B. Pelser, 5 October 2022
  1. Myristica agusanensis Elmer, LPB 8 (1915) 2775; Merrill, EPFP 2 (1923) 178; J.Sinclair, Gard. Bull. Sing. 23 (1968) 218; De Wilde, Blumea 42 (1997) 146; Fl. Males. ser. 1, 14 (2000) 398; Type: Elmer 13284 (L, iso), Mindanao: Agusan del Norte, Mt Urdaneta, Jul-1912. = Myristica laevis ssp. badia W.J.de Wilde, Blumea 42 (1997) 172; Fl. Males. ser. 1, 14 (2000) 509; Type: Rojo 159 (L, holo), Mindanao: Surigao del Sur, Lanuza. Hillside forest, 400m, Oct/Nov-1972. Distribution: Endemic to the Philippines. BASILAN, LUZON: Bataan, Cagayan, Camarines Norte, Camarines Sur, Quezon, Sorsogon, MINDANAO: Agusan del Norte, Agusan del Sur, Bukidnon, Davao del Sur, Surigao del Sur, Zamboanga, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, POLILLO, SAMAR, SIBUYAN, SULU ARCHIPELAGO. Notes: If Myristica agusanensis Elmer and Gymnacranthera lanceolata Merr. are considered conspecific, the epithet of the latter name is the oldest available for this species (and is not blocked by Myristica lanceolata Wall. 1828, a nomen nudum with no nomenclatural priority). The combination Myristica lanceolata (Merr.) ined. therefore needs to be made to replace Myristica agusanensis Elmer as the correct name, because the epithet of Gymnacranthera lanceolata Merr. (1906) has priority over the epithet of Myristica agusanensis Elmer (1915) (John Boggan, pers. comm. 3-Nov-18). 50-1380m. Conservation status: Near Threatened (IUCN 3.1). Native.
    • ssp. agusanensis. = Myristica sorsogonensis Elmer ex Merr., EPFP 2 (1923) 178, nom. inval., in obs., pro syn.; Elmer, LPB 10 (1939) 3809, nom. inval., no Latin descr.; Voucher: Elmer 16010 (L, NY), Luzon: Sorsogon, Mt Bulusan, May-1916. = Myristica laxiflora Merr., PJS 17 (1920) 254; Merrill, EPFP 2 (1923) 179; Type: BS 15498 Reillo (L, NY, US, iso), Basilan. Aug-1912. Throughout the Philippines, except Palawan. Inland forests at low elevation (50 m). Fl. Jul-Aug.; fr. throughout the year. Distribution: Endemic to the Philippines. BASILAN, LUZON: Sorsogon, MINDANAO: Agusan del Norte, Agusan del Sur, Davao del Sur, Zamboanga del Sur, NEGROS, PALAWAN, POLILLO, SAMAR, SIBUYAN, SULU ARCHIPELAGO. Notes: 50m. Native.
    • ssp. squamulosa W.J.de Wilde, Blumea 42 (1997) 146; ≡ Gymnacranthera lanceolata Merr., PJS 1(Suppl.) (1906) 55; Elmer, LPB 3 (1911) 1058; ≡ Myristica lancifolia Merr., EPFP 2 (1923) 178, nom. illeg., non Poir.; Type: FB 3236 Meyer, Luzon. Distribution: Endemic to the Philippines. LUZON: Cagayan, Camarines Sur, SIBUYAN. Native.
  2. Myristica basilanica W.J.de Wilde, Blumea 42 (1997) 148; Fl. Males. Ser. 1, 14 (2000) 411; Type: FB 18929 Miranda (L, US, iso), Basilan, Aug/Sep-1912. Distribution: Endemic to the Philippines. BASILAN. Notes: L.L.Co: Known only from the type collection. Flowers and mature fruits in Aug-Sep. Conservation status: Critically Endangered (IUCN 3.1). DAO category: Other Threatened Species. Native.
  3. Myristica cagayanensis Merr., PJS 17 (1921) 255; Merrill, EPFP 2 (1923) 178; J.Sinclair, Gard. Bull. Sing. 18 (1961) 226; De Wilde, Blumea 42 (1997) 154; Fl. Males. Ser. 1, 14 (2000) 426; ≡ Myristica ceylanica A.DC. var. cagayanensis (Merr.) J.Sinclair, Gard. Bull. Singapore 23 (1968) 442; Type: Bernardo 24277 (PNH, lost). Distribution: Philippines, Taiwan. BATAN, CAMIGUIN DE BABUYANES, LUZON: Cagayan. Notes: Forest of mountain slopes in coastal areas. Fl. & fr. throughout the year. Native.
  4. Myristica cinnamomea King, Ann. Roy. Bot. Gard. (Calcutta) 3 (1891) 292; De Wilde, Blumea 42 (1997) 156; Fl. Males. Ser. 1, 14 (2000) 431; Type: from Peninsular Malaysia. Distribution: Borneo, Malay Peninsula, Philippines, Singapore, Sumatra. MINDANAO: Surigao, TINAGO. Notes: Rare in the Philippines (de Wilde 2000; no material seen by him). 0-1000(-1400)m. Conservation status: Least Concern (IUCN 3.1). Native.
  5. Myristica colinridsdalei W.J.de Wilde, Blumea 42 (1997) 156; Fl. Males. Ser. 1, 14 (2000) 434; Type: Ridsdale at al. ISU 499 (L, holo), Luzon: Isabela, Palanan, Dimapnat, forest on ultrabasic, high canopy forest on low coastal hills, 50m, 15-Apr-1992. Distribution: Endemic to the Philippines. LUZON: Isabela. Notes: Forest on ultramafics, low stature forest with many large-girth trees, streamside forests, or high canopy forest on low coastal hills; locally common on coastal flat, riverine areas. Fl & fr. Apr. 0-50m. Conservation status: Critically Endangered (IUCN 3.1). DAO category: Critically Endangered. Native.
  6. Myristica cumingii Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 442; Merrill, EPFP 2 (1923) 178; De Wilde, Blumea 42 (1997) 154; Fl. Males. ser. 1, 14 (2000) 446; Type: Cuming 1570 (G-DC, lecto; G-Boissier, B, FI, K, L, LE, M, P, W, isolecto). = Myristica mindorensis Merr., PJS 13c (1918) 281; Merrill, EPFP 2 (1923) 179; Type: Merrill 3698. = Myristica nitida Merr., PJS 13c (1918) 282; Merrill, EPFP 2 (1923) 179; Type: FB 22199 Alvarez. = Gymnacranthera negrosensis Elmer, LPB 2 (1909) 576. ≠ Myristica ceylanica var. ceylanica auct. non A.DC.; J.Sinclair, Gard. Bull. Sing. 23 (1968) 437, pro parte. Distribution: Endemic to the Philippines. ALABAT, BILIRAN, CATANDUANES, LEYTE, LUZON: Albay, Camarines, Camarines Sur, Laguna, Nueva Ecija, Quezon, Sorsogon, MASBATE, MINDANAO: Agusan del Norte, Surigao, Surigao del Norte, Zamboanga, MINDORO, NEGROS, PALAWAN, PANAY, POLILLO, SAMAR, SIBUYAN, TABLAS, TICAO. Notes: Forests, common. Fl. mainly May-Aug; fr. throughout the year. 0-1000m. Conservation status: Least Concern (IUCN 3.1). Native.
  7. Myristica fatua Houtt., Nat. Hist. (Houttuyn) 2(3) (1774) 337; J.Sinclair, Gard. Bull. Sing. 23 (1968) 268, pro parte; De Wilde, Blumea 35 (1990) 242; Blumea 40 (1995) 279; Blumea 42 (1997) 166; Fl. Males. ser. 1, 14 (2000) 460; Fernando et al., Forest Flora of Dinagat (2017) 66; Type: Clusius, Tab. 14 (lecto). = Myristica plumeriifolia Elmer, LPB 3 (1911) 1063; Merrill, EPFP 2 (1923) 180; Type: Elmer 11063, Mindanao. Distribution: DINAGAT, LUZON: Aurora (photos), Pangasinan (photos), Quezon, MINDANAO: Agusan del Norte, Davao de Oro, Davao del Sur, Misamis Occidental, Surigao, Surigao del Sur (photos), MINDORO, PALAWAN (photos), SAMAR. Notes: 0-1235m. Native.
    • ssp. fatua. Distribution: Borneo, Moluccas, New Guinea, Philippines. LUZON: Quezon, MINDANAO: Agusan del Norte, Davao del Sur. Notes: Forest along Baracatan River. 0-600m. Native.
  8. Myristica frugifera W.J.de Wilde, Blumea 42 (1997) 166; Fl. Males. Ser. 1, 14 (2000) 472; Type: Ridsdale 1219 (L, iso), Mindoro. Distribution: Endemic to the Philippines. CALAYAN, LEYTE, LUZON: Isabela, Quezon, Sorsogon, MINDORO, PALAUI, POLILLO. Notes: Understorey tree of primary and disturbed lowland forest. Fr. Feb, Jun. 0-500m. Conservation status: Vulnerable (IUCN 3.1). Native.
  9. Myristica guatteriifolia A.DC., Ann. Sci. Nat. (Paris), 4th Sér. 4 (1855) 20, as 'guatteriaefolia'; J.Sinclair, Gard. Bull. Sing. 16 (1958) 350; Gard. Bull. Sing. 23 (1968) 213; De Wilde, Blumea 42 (1997) 168; Fl. Males. ser. 1, 14 (2000) 483; Type: Cuming 1582, Luzon: Batangas. Distribution: Borneo, Java, Lesser Sunda Isls, Malay Peninsula, Myanmar, Philippines, ?Thailand, Vietnam. BALABAC, BASILAN, CAMIGUIN DE BABUYANES, CULION, GUIMARAS, LAGEN, LUZON: Batangas, Camarines Norte, Camarines Sur, Laguna, Quezon, MINDANAO: Lanao del Sur, Zamboanga, Zamboanga del Sur, NEGROS, PALAWAN, PANAY, SAMAR, SIBUYAN. Notes: Throughout the Philippines. 0-900m. Conservation status: Least Concern (IUCN 3.1). Native.
  10. Myristica iners Blume, Bijdr. fl. Ned. Ind. (1826) 575; De Wilde, Blumea 42 (1997) 168; Fl. Males. ser. 1, 14 (2000) 493. Distribution: Borneo, Cambodia, Java, Malay Peninsula, Philippines, Sumatra, Thailand. Notes: Philippine record, fide De Wilde (2000), is based on Wenzel 3537 (K) which agrees with specimens from NE Borneo. Its locality however is remarkable, far from main distributional area of the species. 0-1200(-1800)m. Conservation status: Least Concern (IUCN 3.1). Native.
  11. Myristica laevis W.J.de Wilde, Blumea 42 (1997) 170; Fl. Males. ser. 1, 14 (2000) 508; Type: PNH 118091 Gutierrez (L, holo), Samar: Mt Sohoton, Pesikohon site, Apr-1970. Distribution: Endemic to the Philippines. SAMAR. Notes: Fl. & fr. Apr. 400m. Native.
  12. Myristica longepetiolata W.J.de Wilde, Blumea 42 (1997) 173; Fl. Males. ser. 1, 14 (2000) 515; Type: PNH 21548 Sulit (L, holo), Biliran: Mt Suiro, N slope, 830m. Distribution: Endemic to the Philippines. BILIRAN, LUZON: Camarines, Camarines Norte, Sorsogon. Notes: Sloping forests. Fr. Apr-May, Jul-Aug, Dec. 830m. Conservation status: Endangered (IUCN 3.1). DAO category: Other Threatened Species. Native.
  13. Myristica mindanaensis Warb., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 68 (1897) 497; De Wilde, Blumea 35 (1990) 249; Blumea 42 (1997) 178; Fl. Males. ser. 1, 14 (2000) 529; Odulio et al., Biodivers. Data J. 14 (2026) 26; Type: Warburg 13300 (L, iso), Mindanao. Distribution: Moluccas, Philippines. MINDANAO: Agusan del Norte, Davao del Sur, SAMAR. Notes: 0-250(-700)m. Native.
  14. Myristica nivea Merr., PJS 1(Suppl.) (1906) 191; Merrill, EPFP 2 (1923) 179; De Wilde, Blumea 42 (1997) 178; Fl. Males. ser. 1, 14 (2000) 532; Buhay et al., Biodiversitas 26 (2025) 3086; Type: FB 3454 Hutchinson (US, iso), Basilan: 14-Dec-1905. Distribution: Moluccas, Philippines. BASILAN, CEBU (photos), MINDANAO: Zamboanga, Zamboanga Sibugay, SAMAR. Notes: Lowland forests. Fl. Dec; fr. Oct-Nov, Mar. Native.
  15. Myristica philippensis Lam., Hist. Acad. Roy. Sci. (Paris) 788 (1761) 161; Merrill, EPFP 2 (1923) 179; J.Sinclair, Gard. Bull. Sing. 23 (1968) 138; De Wilde, Blumea 42 (1997) 179; Fl. Males. ser. 1, 14 (2000) 541; Coritico et al., PJS 151 (2022) 2439; Romeroso et al., J. Asia-Pacific biodiv. 17 (2024) 12; Type: Sonnerat s.n., Philippines. Distribution: Endemic to the Philippines. BASILAN, CALAYAN, HOMONHON, LEYTE, LUZON: Abra, Aurora, Bataan, Batangas, Benguet, Camarines, Camarines Sur, Ilocos Sur, Isabela, Laguna, Pampanga, Pangasinan, Quezon, Rizal, Sorsogon, Zambales, MINDANAO: Bukidnon, Zamboanga, Zamboanga del Sur, MINDORO, PALAWAN, POLILLO, SAMAR, SULU ARCHIPELAGO. Notes: Primary forests. Fl. & fr. possibly throughout the year. 0-400m. Conservation status: Least Concern (IUCN 3.1). Native.
  16. Myristica pilosigemma W.J.de Wilde, Blumea 42 (1997) 179; Fl. Males. ser. 1, 14 (2000) 543; Type: PNH 118078 Gutierrez (L, holo), Samar: Western Samar, Mt Sohoton. Along trail, 1800ft., Apr-1970. Distribution: Endemic to the Philippines. MINDANAO: Bukidnon, SAMAR. Notes: Fl. Apr; fr. Jun-Jul. 600m. Conservation status: Critically Endangered (IUCN 3.1). DAO category: Other Threatened Species. Native.
  17. Myristica rubrinervis W.J.de Wilde, Blumea 42 (1997) 180; Fl. Males. Ser. 1, 14 (2000) 554; Type: Ridsdale SMHI 258 (L, holo), Palawan: Palawan, Puerto Princesa, Mt Beaufort, W spur, 815m, 29-Mar-1984. Distribution: Endemic to the Philippines. LEYTE, MINDANAO: Misamis Occidental, MINDORO, NEGROS, PALAWAN. Notes: 40-900m. Conservation status: Vulnerable (IUCN 3.1). Native.
    • var. rubrinervis. Distribution: Endemic to the Philippines. PALAWAN. Notes: Known from Puerto Princesa: Mt Beaufort. Stunted montane forest with many epiphytes but little moss; closed broad-leaved rainforest on ultramafics. Fr. Mar-May. 400-900m. Native.
    • var. duplex W.J.de Wilde, Blumea 42 (1997) 182; Fl. Males. ser. 1, 14 (2000) 555; Type: Ridsdale SMHI 1528 (L, holo), Palawan: Pagdanan Range, Ibangley, Brookside Hill. Closed broad-leaved rainforest with Dipterocarpus spp., 40m, 23-Apr-1984. Distribution: Endemic to the Philippines. LEYTE, MINDORO, PALAWAN. Notes: Lowland dipterocarp to montane ridge forests (with Agathis). Fl. (immature) & fr. Feb-Apr. 40-900m. Native.
  18. Myristica simiarum A.DC., Ann. Sci. Nat. (Paris), 4th Sér. 4 (1855) 29; Merrill, EPFP 2 (1923) 180; De Wilde, Blumea 35 (1990) 254; Fl. Males. ser. 1, 14 (2000) 565; ≡ Myristica elliptica Wall. ex Hook.f. & Thomson var. simiarum (A.DC.) J.Sinclair, Gard. Bull. Singapore 16 (1958) 356; Gard. Bull. Sing. 23 (1968) 190; Type: Callery 34, Luzon: “Manilla”. Distribution: BATAN, DINAGAT (photos), LEYTE, LUZON: Aurora, Bataan, Cagayan, Camarines Norte, Camarines Sur, Ifugao, Ilocos Norte, Isabela, La Union, Laguna, Quezon, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Surigao del Sur, Zamboanga, Zamboanga del Norte, MINDORO, PALAWAN, SAMAR. Notes: 0-1400m. Native.
    • ssp. simiarum. Distribution: Philippines, Taiwan. BATAN, LEYTE, LUZON: Aurora, Bataan, Cagayan, Ifugao, Ilocos Norte, Isabela, La Union, Laguna, Quezon, MINDANAO: Agusan del Norte, Zamboanga del Norte, SAMAR. Notes: Forests. Fl. & fr. throughout the year. 0-600m. Native.
    • ssp. celebica (Miq.) W.J.de Wilde, Blumea 35 (1990) 255; Blumea 42 (1997) 185; Fl. Males. ser. 1, 14 (2000) 569; ≡ Myristica elliptica Wall. ex Hook.f. & Thomson var. celebica (Miq.) J.Sinclair, Gard. Bull. Singapore 16 (1958) 356; Gard. Bull. Sing. 23 (1968) 194; Type: from Sulawesi, Bacan, and Buru (syn). Distribution: Moluccas, Philippines, Sulawesi. LUZON: Camarines Sur, Zambales, MINDANAO: Zamboanga. Notes: 10-900m. Native.
  19. Myristica umbellata Elmer, LPB 5 (1913) 1816; Merrill, EPFP 2 (1923) 180; J.Sinclair, Gard. Bull. Sing. 23 (1968) 174; De Wilde, Blumea 42 (1997) 188; Fl. Males. ser. 1, 14 (2000) 600; Type: Elmer 12820 (L, MO, NY, SING, US, syn), Palawan: Palaawan, Puerto Princesa, Mt Pulgar (Thumb Peak), forested ridge, 1000ft, Mar-1911; Elmer 13166 (L, NY, US, syn), Palawan: Palawan, Puerto Princesa, Mt Pulgar, on trail to Napsan, 750ft, May-1911. Distribution: Endemic to the Philippines. LAGEN, PALAWAN. Notes: Low canopy lowland and montane forest (with Agathis celebica), also on limestone and ultramafic alluvial deposits. Fl. Mar; fr. Apr-May. 50-800m. Native.
  20. Myristica wenzelii Merr., PJS 10c (1915) 270; Merrill, EPFP 2 (1923) 180; De Wilde, Blumea 42 (1997) 188; Fl. Males. ser. 1, 14 (2000) 615; ≡ Myristica fatua Houtt. var. wenzelii (Merr.) J.Sinclair, Gard. Bull. Singapore 23 (1968) 309; Type: Wenzel 1152 (MO, iso), Leyte: Masaganap near Jaro, in forests, 700m, Sep-1914. Distribution: Endemic to the Philippines. LEYTE, MINDANAO: Surigao del Norte, SAMAR. Notes: Fl. Oct; fr. Oct-Mar. Low elevation forests. 350m (photos). Conservation status: Critically Endangered (IUCN 3.1). Native.

References