Lycopodiaceae

Dendrolycopodium A.Haines (Not native)

Possibly present

Dendrolycopodium juniperoideum (Sw.) A.Haines, Fam. Huperziac. Lycopodiac. New England (2003) 84; ≡ Lycopodium juniperoideum Sw., Syn. Fil. (1806) 178. Distribution: ?Philippines, Russia, Taiwan.

Diphasiastrum Holub

Last edited by Julie F. Barcelona & Pieter B. Pelser, 5 July 2022

Diphasiastrum multispicatum (J.H.Wilce) Holub, Preslia 47 (1975) 108; ≡ Lycopodium multispicatum J.H.Wilce, Nova Hedwigia 3 (1961) 103. Distribution: China, India, Philippines, ?Sulawesi, Taiwan, Thailand, Tibet, Vietnam. LUZON: Benguet, Ifugao, Ilocos Norte, Kalinga, Laguna, Mountain Province, Nueva Vizcaya, Quezon, MINDANAO: Cotabato, MINDORO. Native.
Diphasiastrum wightianum (Wall. ex Hook. & Grev.) Holub, Preslia 47 (1975) 108; ≡ Diphasium wightianum (Wall. ex Hook. & Grev.) Pic.Serm., Webbia 24 (1970) 720; ≡ Lycopodium wightianum Wall. ex Hook. & Grev., Bot. Misc. 2 (1831) 379. Distribution: Borneo, India, New Guinea, Philippines, Sri Lanka. LUZON: Benguet, Ifugao, Nueva Vizcaya. Native.

Possibly present

Diphasiastrum nikoense (Franch. & Sav.) Holub, Preslia 47 (1975) 108; ≡ Lycopodium nikoense Franch. & Sav., Enum. pl. Jap. 198 (1879) 613; ≡ Lycopodium sitchense var. nikoense (Franch. & Sav.) Takeda, Bot. Mag. (Tokyo) 23 (1909) 221; ≡ Diphasiastrum sitchense ssp. nikoense (Franch. & Sav.) Á.Löve & D.Löve, Taxon 26 (1977) 324. Distribution: Japan, Korea, ?Philippines.

Diphasium Rothm.

Last edited by Julie F. Barcelona, 13 March 2021

Diphasium scariosum (G.Forst.) Rothm., Repert. Spec. Nov. Reg. Veg. (1944) 54; ≡ Lycopodium scariosum G.Forst., Fl. Ins. Austr. (1786) 87. Distribution: Australia, Borneo, New Zealand, Pacific Ocean, Philippines, Sulawesi. LUZON: Benguet, Ifugao, Nueva Vizcaya, MINDANAO: Cotabato, Davao, Davao del Sur. Native.

Huperzia Bernh.

Last edited by Julie F. Barcelona, 15 April 2022

Huperzia javanica (Sw.) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 10; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22043; ≡ Lycopodium javanicum Sw., J. Bot. (Schrader) 1800 (1801) 114. = Huperzia serrata auct. non (Thunb.) Trevis. Distribution: America, Australia, Bhutan, Borneo, China, India, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, Nepal, New Caledonia, Pacific Ocean, Philippines, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam. MINDANAO: Bukidnon, South Cotabato. Native.
Huperzia miniata (Spring) Trevis., Atti Soc. Ital. Sci. Nat. 17 (1875) 248; ≡ Lycopodium selago var. miniatum (Spring) B.C.Tan & Tolentino, PJS 116 (1987) 437; ≡ Lycopodium miniatum Spring, Monogr. Lycop. 1 (1842) 28. Distribution: Australia, Eurasia, Indian Ocean, Japan, Java, Korea, Madagascar, N America, Philippines, Ryukyu Isls, Sulawesi. Native.
Huperzia minima (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 74; ≡ Urostachys minimus Herter, Bot. Arch. 3 (1923) 13. Distribution: Endemic to the Philippines. MINDANAO. Native.
Huperzia somae (Hayata) Ching, Acta Bot. Yunnan. 3 (1981) 301; ≡ Lycopodium somae Hayata, Ic. Pl. Formos. 5 (1915) 255. Distribution: Japan, Philippines, Taiwan. LUZON. Native.

Lateristachys Holub

Last edited by Julie F. Barcelona, 13 March 2021

Lateristachys halconensis (Copel.) Holub, Folia Geobot. Phytotax., 18 (1983) 441; ≡ Lycopodium halconense Copel., PJS 2 c (1907) 149. Distribution: Endemic to the Philippines. MINDORO. Native.

Lycopodiastrum Holub ex R.D.Dixit

Last edited by Julie F. Barcelona, 13 March 2021

Lycopodiastrum casuarinoides (Spring) Holub ex R.D.Dixit, J. Bombay Nat. Hist. Soc. (1981) 77; ≡ Lycopodium casuarinoides Spring, Bull. Acad. Roy. Sci. Belg. 8 (1841) 521. Distribution: Bhutan, Borneo, China, India, Japan, Malay Peninsula, Myanmar, New Guinea, Philippines, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam. LUZON: Benguet, Ifugao, Ilocos Norte, Ilocos Sur, Mountain Province, Nueva Vizcaya, MINDANAO: Bukidnon, MINDORO. Native.

Lycopodium L.

Last edited by Julie F. Barcelona, 13 March 2021

Lycopodium clavatum L., Sp. Pl. (1753) 1101; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22043. Distribution: Africa, America, Borneo, Caribbean, Continental Asia, Europe, Japan, Korea, Madagascar, Malay Peninsula, Middle East, Moluccas, New Caledonia, New Guinea, Pacific Ocean, Philippines, Solomon Isls, Sulawesi, Thailand, Vietnam. LUZON: Benguet, Camarines Norte, Ifugao, Kalinga, Laguna, Mountain Province, Nueva Vizcaya, Quezon, MINDANAO: Davao del Sur, Lanao del Sur, South Cotabato, MINDORO. Native.
- ssp. clavatum. Notes: Widespread, incl. Philippines. Native.

Palhinhaea Franco & Vasc.

Last edited by Julie F. Barcelona, 2 May 2022

Palhinhaea cernua (L.) Vasc. & Franco, Bol. Soc. Brot. sér. 2, 41 (1967) 25; ≡ Lycopodiella cernua (L.) Pic.Serm., Webbia 23 (1968) 166; ≡ Lycopodium cernuum L., Sp. Pl. 2 (1753) 1103; Herter, PJS 22 (1923) 69. = Lycopodium cernuum forma ingens B.C.Tan & Tolentino, PJS 116 (1987) 438. Distribution: BASILAN, BILIRAN, CAMIGUIN, CATANDUANES, LEYTE, LUZON: Albay, Apayao, Aurora, Bataan, Batangas, Benguet, Cagayan, Camarines Norte, Camarines Sur, Ifugao, Kalinga, Laguna, Mountain Province, NCR, Nueva Ecija, Nueva Vizcaya, Pampanga, Quezon, Rizal, Sorsogon, Tarlac, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao, Davao Oriental, Davao del Norte, Davao del Sur, Lanao del Norte, Lanao del Sur, Misamis, South Cotabato, Surigao, Surigao del Norte, Zamboanga Sibugay, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY, POLILLO, SAMAR. Notes: Widespread, known from most tropical areas. Native.

Phlegmariurus (Herter) Holub

Last edited by Julie F. Barcelona & Pieter B. Pelser, 5 July 2022

Phlegmariurus banayanicus (Herter) A.R.Field & Bostock, PhytoKeys 20 (2013) 36; ≡ Huperzia banayanica (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 71; ≡ Lycopodium banayanicum (Herter) Copel., Univ. Calif. Publ. Bot. 12 (1931) 384; ≡ Urostachys banayanicus Herter, Bot. Arch. 3 (1923) 17. Distribution: Endemic to the Philippines. CAMIGUIN, LUZON: Bataan, Ifugao, Laguna, Mountain Province, Quezon, Rizal, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao, Davao del Sur, MINDORO, NEGROS, PALAWAN, PANAY. Native.
Phlegmariurus carinatus (Desv. ex Poir.) Ching, Acta Bot. Yunnan. 20 (1982) 448; Coritico & Amoroso, Nature Conservation Research 5 (2020) 81; ≡ Huperzia carinata (Desv. ex Poir.) Trevis., Atti Soc. Ital. Sci. Nat. 17 (1874) 247; ≡ Lycopodium carinatum Desv. ex Poir., Encycl., Suppl. 3(2) (1814) 555. Distribution: Australia, Bangladesh, Bismarck Arch, Borneo, ?Cambodia, China, ?Laos, Malay Peninsula, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. LUZON: Bataan, Benguet, Mountain Province, NCR, MINDANAO: Bukidnon, Cotabato, Lanao del Sur, Zamboanga, MINDORO, NEGROS, POLILLO. Conservation status: Endangered (DAO 2017-11, as Huperzia carinata). Native.
Phlegmariurus delbrueckii (Herter) A.R.Field & Bostock, PhytoKeys 20 (2013) 39; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22043; ≡ Huperzia delbrueckii (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 72; ≡ Urostachys delbrueckii Herter, Bot. Arch. 3 (1923) 19. Distribution: Endemic to the Philippines. MINDANAO: Bukidnon, Cotabato, Davao, Davao del Sur, Misamis, South Cotabato. Native.
Phlegmariurus elmeri (Herter) A.R.Field & Bostock, PhytoKeys 20 (2013) 39; ≡ Huperzia elmeri (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 72; ≡ Urostachys elmeri Herter, Bot. Arch. 3 (1923) 16. Distribution: Endemic to the Philippines. CULION, LUZON: Bataan, Benguet, Cagayan, NCR, Quezon, Rizal, Tayabas, MINDANAO: Cotabato, Davao, Davao del Sur, MINDORO, PALAWAN. Conservation status: Endangered (DAO 2017-11, as Huperzia elmeri). Native.
Phlegmariurus filiformis (Sw.) W.H.Wagner, Contr. Univ. Michigan Herb. 20 (1995) 242; ≡ Huperzia filiformis (Sw.) Holub, Folia Geobot. Phytotax. 20 (1985) 72. Distribution: Africa, America, Australia, Borneo, Indian Ocean, Malay Peninsula, Pacific Ocean, Philippines. MINDANAO: Bukidnon. Native.
Phlegmariurus fordii (Baker) Ching, Acta Phytotax. Sin. 20 (1982) 447; ≡ Huperzia fordii (Baker) Holub, Folia Geobot. Phytotax. 20 (1985) 72; ≡ Lycopodium fordii Baker, Fern Allies (1887) 17; Hatusima, Mem. Fac. Agr. Kagoshima Univ. 5 (1966) 18. Distribution: China, Japan, Philippines, Ryukyu Isls, Taiwan, Vietnam. Native.
Phlegmariurus giganteus (Herter) A.R.Field & Bostock, PhytoKeys 20 (2013) 40; ≡ Huperzia gigantea (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 73; ≡ Urostachys verticillatus var. giganteus Herter, Bot. Arch. 3 (1923) 13. Distribution: Endemic to the Philippines. LUZON, MINDANAO: Cotabato, Davao, Misamis Occidental. Native.
Phlegmariurus magnusianus (Herter) A.R.Field & Testo, Molec. Phylogen. Evol. 94 (2015) 645; Coritico & Amoroso, Nature Conservation Research 5 (2020) 80; ≡ Huperzia magnusiana (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 74; ≡ Lycopodium magnusianum Herter, Hedwigia 49 (1909) 91; Type: MS Clemens s.n., Apr-1907. = Huperzia whitfordii (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 78; ≡ Urostachys whitfordii Herter, Bot. Arch. 3 (1923) 14. Distribution: Endemic to the Philippines. BASILAN, LEYTE, LUZON: Quezon, MINDANAO: Bukidnon, Cotabato, Lanao del Sur, Surigao, Zamboanga del Sur, MINDORO, PANAY, SULU ARCHIPELAGO. Conservation status: Critically Endangered (DAO 2017-11, as Huperzia magnusiana). Native.
Phlegmariurus merrillii (Herter) A.R.Field & Bostock, PhytoKeys 20 (2013) 43; ≡ Huperzia merrillii (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 74; ≡ Lycopodium merrillii (Herter) Copel., Univ. Calif. Publ. Bot. 12 (1931) 384; ≡ Urostachys merrillii Herter, Bot. Arch. 3 (1923) 15; Type: Merrill 965. Distribution: Endemic to the Philippines. LUZON: Benguet, Ifugao, Ilocos Sur, Kalinga, Mountain Province, Nueva Vizcaya, MINDANAO: Bukidnon. Native.
Phlegmariurus nummulariifolius (Blume) Ching, Acta Bot. Yunnanica 3 (1981) 298; ≡ Huperzia nummulariifolia (Blume) Jermy in T.C.Chambers, Jermy & Crabbe, Brit. Fern Gaz. 10 (1971) 176; ≡ Lycopodium nummulariifolium Blume, Enum. Pl. Javae 2 (1828) 263. Distribution: Bismarck Arch, Borneo, Java, Malay Peninsula, Moluccas, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sulawesi, Sumatra, Thailand. MINDANAO: Bukidnon. Native.
Phlegmariurus phlegmaria (L.) Holub, Preslia 36 (1964) 21; Batuyong et al., Biodiversitas 21 (2020) 3974; Coritico & Amoroso, Nature Conservation Research 5 (2020) 81; ≡ Huperzia phlegmaria (L.) Rothm., Fedde Repert. 54 (1944) 62; ≡ Urostachys phlegmaria (L.) Herter ex Nessel, Bärlappgewächse (1939) 215; Herter, Bot. Arch. 3 (1923) 17, nom. inval.; PJS 22 (1923) 66; ≡ Lycopodium phlegmaria L., Sp. Pl. 2 (1753) 1101. Distribution: Africa, Andaman Isls, Australia, Bangladesh, Bhutan, Bismarck Arch, Borneo, Cambodia, China, India, Japan, Laos, Malay Peninsula, Moluccas, Nepal, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. GENERAL ISLAND, LEYTE, LUZON: Aurora, Bataan, Benguet, Cagayan, Ifugao, Ilocos Norte, Kalinga, Laguna, Mountain Province, NCR, Pampanga, Quezon, Sorsogon, Tarlac, Zambales, MINDANAO: Agusan del Norte, Agusan del Sur, Bukidnon, Cotabato, Davao, Davao Oriental, Davao del Sur, Lanao del Sur, Misamis Occidental, Zamboanga del Norte, MINDORO, NEGROS, PALAWAN, POLILLO, SULU ARCHIPELAGO. Conservation status: Endangered (DAO 2017-11, as Huperzia phlegmaria). Native.
Phlegmariurus phyllanthus (Hook. & Arn.) R.D.Dixit, J. Bombay Nat. Hist. Soc. 77 (1981) 541; ≡ Huperzia phyllantha (Hook. & Arn.) Holub, Folia Geobot. Phytotax. 20 (1985) 75; ≡ Lycopodium phyllanthum Hook. & Arn., Bot. Beechey Voy. (1832) 102. Distribution: Andaman Isls, Borneo, India, Malay Peninsula, New Caledonia, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Sri Lanka. LUZON: Batangas, Laguna, NCR, MINDANAO: Agusan del Norte, Zamboanga del Sur. Native.
Phlegmariurus pinifolius (Trevis.) Kiew, PhytoKeys 96 (2018) 108; ≡ Urostachys pinifolius (Trevis.) Herter, Bot. Arch. 3 (1923) 65; ≡ Huperzia pinifolia Trevis., Atti Soc. Ital. Sci. Nat. 17 (1874) 247; ≡ Lycopodium pinifolium Blume, Enum. Pl. Javae 2 (1828) 264, nom. illeg., non Kaulf. = Huperzia piscium (Herter) Holub, Folia Geobot. Phytotax. 26 (1991) 93; ≡ Lycopodium piscium (Herter) Tagawa & K.Iwats., Acta Phytotax. Geobot. 22 (1967) 103; ≡ Urostachys piscium Herter, Ind. Lycopod. (1949) 75. Distribution: Borneo, Java, Malay Peninsula, Moluccas, Pacific Ocean, Philippines, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Thailand. LEYTE, LUZON: Abra, Bataan, Batangas, Benguet, Ilocos Norte, Laguna, Nueva Vizcaya, Quezon, Zambales, MINDANAO: Agusan del Norte, Bukidnon, NEGROS, PALAWAN. Native.
Phlegmariurus salvinioides (Herter) Ching, Acta Bot. Yunnan. 4 (1982) 122; Coritico & Amoroso, Nature Conservation Research 5 (2020) 81; ≡ Huperzia salvinioides (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 76; ≡ Lycopodium salvinioides (Herter) Tagawa, Acta Phytotax. Geobot. 8 (1939) 20; ≡ Urostachys salvinioides Herter, Bot. Arch. 3 (1923) 18; Type: Curran s.n. (B20011394, lecto), Luzon: Cagayan, Musure. = Huperzia edanoi (Copel.) Holub, Folia Geobot. Phytotax. 20 (1985) 72; ≡ Lycopodium edanoi Copel., PJS 46 (1931) 209; Type: BS 77930 Edaño. Distribution: Japan, Philippines, Taiwan, ?Vietnam. CATANDUANES, LEYTE, LUZON: Abra, Apayao, Bataan, Cagayan, Camarines Sur, Ifugao, Kalinga, Laguna, Nueva Vizcaya, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao, Davao Oriental, Davao del Sur, Lanao del Norte, North Cotabato, Zamboanga, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY (photos), POLILLO, SAMAR. Conservation status: Endangered (DAO 2017-11). Native.
Phlegmariurus squarrosus (G.Forst.) Á.Löve & D.Löve, Taxon 26 (1977) 324; Coritico & Amoroso, Nature Conservation Research 5 (2020) 81; ≡ Huperzia squarrosa (G.Forst.) Trevis., Atti Soc. Ital. Sci. Nat. 17 (1874) 247; ≡ Lycopodium squarrosum G.Forst., Fl. Ins. Aust. (1786) 479. Distribution: Andaman Isls, Australia, Bangladesh, Bhutan, Borneo, China, India, Indian Ocean, Japan, Java, Laos, Lesser Sunda Isls, Madagascar, Malay Peninsula, Moluccas, Myanmar, Nepal, New Caledonia, ?Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam. CAMIGUIN DE BABUYANES, LEYTE, LUZON: Albay, Aurora, Bataan, Benguet, Camarines Sur, Kalinga, Laguna, Mountain Province, Pampanga, Quezon, Rizal, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao Oriental, Davao del Sur, Misamis Occidental, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN. Conservation status: Endangered (DAO 2017-11). Native.
Phlegmariurus tetrastichus (Kunze) A.R.Field & Bostock, PhytoKeys 20 (2013) 48; ≡ Lycopodium tetrastichum Kunze, Bot. Zeitung (Berlin) 6 (1848) 99; Type: from Java. Distribution: Borneo, Java, Malay Peninsula, Moluccas, Philippines, Sulawesi, Sumatra, Thailand, Vietnam. LUZON: Bataan, Benguet, Quezon. Native.
Phlegmariurus toppingii (Herter) A.R.Field & Bostock, PhytoKeys 20 (2013) 48; ≡ Huperzia toppingii (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 77; ≡ Urostachys toppingii Herter, Bot. Arch. 3 (1923) 15. Distribution: Endemic to the Philippines. LUZON: Benguet. Native.
Phlegmariurus verticillatus (L.f.) A.R.Field & Testo, Molec. Phylogen. Evol. 94 (2015) 645; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22043; ≡ Huperzia verticillata (L.f.) Trevis., Atti Soc. Ital. Sci. Nat. 17 (1874) 248; ≡ Lycopodium verticillatum L.f., Suppl. Pl. (1782) 448. = Lycopodium verticillatum var. maximum Herter in Engl., Bot. Jahrb. 54 (1917) 227. Distribution: Indian Ocean, Philippines. LUZON: Mountain Province (photos), MINDANAO: Bukidnon, Cotabato, Davao, Davao del Sur, South Cotabato, MINDORO. Notes: Like the other Malesian populations, those in the Philippines probably belong to Phlegmariurus ignambiensis (Compton) A.R.Field & Testo (Hassler & Schmitt 2019). Native.

Possibly present

Phlegmariurus cryptomerinus (Maxim.) Satou, Hikobia 12 (1997) 268; ≡ Lycopodium cryptomerinum Maxim., Bull. Acad. Imp. Sci. Saint-Petersbourg 15 (1870) 231. Distribution: China, Japan, Korea, ?Philippines, Ryukyu Isls, Taiwan.
Phlegmariurus hippuris (Desv. ex Poir.) A.R.Field & Testo, Molec. Phylogen. Evol. 94 (2015) 645; ≡ Lycopodium hippuris Desv. ex Poir., Encycl. Suppl. 3 (1814) 559. Distribution: Java, Moluccas, ?Pacific Ocean, ?Philippines.

Pseudodiphasium Holub

Last edited by Julie F. Barcelona, 13 March 2021

Pseudodiphasium volubile (G.Forst.) Holub, Folia Geobot. Phytotax. 18 (1983) 442; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22043; ≡ Lycopodium volubile G.Forst., Fl. Ins. Austr. (1786) 86. Distribution: Australia, Borneo, Java, Lesser Sunda Isls, Malay Peninsula, Moluccas, New Caledonia, New Guinea, New Zealand, Pacific Ocean, Philippines, Solomon Isls, Sulawesi. LUZON: Abra, Benguet, Camarines Norte, Ifugao, Ilocos Norte, Kalinga, Laguna, Mountain Province, Nueva Vizcaya, MINDANAO: Bukidnon, Cotabato, Davao, Davao del Sur, Misamis Occidental, South Cotabato, MINDORO, PANAY. Native.

References

📋 Barcelona, JF, NE Dolotina, GS Madroñero, WG Granert & DD Sopot. 2006. The ferns and fern allies of the karst forests of Bohol Island, Philippines. American Fern Journal 96: 1-20
Barcelona, JF. 2002. Philippine pteridophyte collections as a resource for conservation planning. Proceedings of the “International Symposium on the Fern Flora Worldwide: Threats and Responses” sponsored by the British Pteridological Society and IUCN- Species Survival Commission.” Fern Gazette 16(6, 7 & 8): 307-312.
📋 Barcelona, JF. 2003. Preliminary report on the ferns and fern allies (pteridophytes) of Mt. Bali-it, Balbalasang-Balbalan National Park, Kalinga, Northern Luzon, Philippines. Sylvatrop: The Technical Journal of Philippine Ecosystems and Natural Resources 13: 81-92
Barcelona, JF. 2003. The taxonomy and ecology of the pteridophytes of Mt. Iraya & vicinity, Batan Island, Batanes Province, Northern Philippines, in Pteridology in the New Millenium (Chandra and Srivastava, eds). Kluwer Academy Publishers. The Netherlands, 299-325.
Barcelona, JF. 2004. Collection and conservation status of pteridophytes in Panay Island, Philippines. Philipp. Scient. 41:57-73.
📋 Barcelona, JF. 2005. Noteworthy fern discoveries in the Philippines at the turn of the 21st Century. Fern Gazette 17: 139-146
📋 CABI Invasive Species Compendium
Christenhusz, MJM, X-C Zhang & H Schneider. 2011. A linear sequence of extant families and genera of lycophytes and ferns. Phytotaxa 19: 7-54.
Copeland, E.B. 1961. Fern Flora of the Philippines 3: 547-553.
📋 Coritico FP & VB Amoroso. 2020. Threatened lycophytes and ferns in four protected areas of Mindanao, Philippines. Nature Conservation Research 5: 78-88
📋 Coritico, FP, VB Amoroso, FM Acma, YLL Cariño & PW Fritsch. 2020. Ferns and lycophytes of Mt. Tago Range, Bukidnon, southern Philippines: species richness, distribution, and conservation status. Philippine Journal of Science 149: 773-790
📋 DENR Administrative Order. 2017. Updated national list of threatened Philippine plants and their categories. DAO 2017-11
📋 Delos Angeles, MD, IE Buot Jr, L Hongmei & H Schneider. 2022. Pteridophyte diversity in the Samar Island Natural Park (SINP), Samar Island, Philippines. Philippine Journal of Science 151: 1929-1942
📋 Ebihara A, CR Fraser-Jenkins, BS Parris, XC Zhang, YH Yang, WL Chiou, HM Chang, S Lindsay, D Middleton, M Kato, TN Praptosuwiryo, VB Amoroso, JF Barcelona, RHG Ranil, CH Park, N Murakami & A Hoya. 2012. Rare and threatened pteridophytes of Asia 1. An enumeration of narrowly distributed taxa. Bulletin of the National Museum of Nature and Science, Series B. 38: 93-119
📋 Field, AR, W Testo, PD Bostock, JAM Holtum & M Waycott. 2016. Molecular phylogenetics and the morphology of the Lycopodiaceae subfamily Huperzioideae supports three genera: Huperzia, Phlegmariurus and Phylloglossum. Molecular Phylogenetics and Evolution 94: 635-657
📋 Hassler, M & B Schmitt. 2019. Checklist of ferns and lycophytes of the world
📋 Hassler, M & B Schmitt. World ferns: checklist of ferns and lycophytes of the world
Hassler, M. 2019. World ferns: checklist of ferns and lycophytes of the world. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
📋 Smith AR, Pryer KM, Schuettpelz E, Korall P, Schneider H & Wolf PG. 2006. A classification for extant ferns. Taxon 55: 705–731
📋 TPG. 2019. Updating Taiwanese pteridophyte checklist: a new phylogenetic classification. Taiwania 64: 367-395