Heliotropiaceae

Argusia Boehmer = Heliotropium

Euploca Nutt.

Last edited by Pieter B. Pelser, 29 May 2020

Photos
  1. Euploca bracteata (R.Br.) M.W.Frohl & M.W.Chase, Phytotaxa 434 (2020) 15; --Heliotropium bracteatum R.Br., Prodr. (1810) 493; --Merr., EPFP 3 (1923) 377. Java to tropical Australia. LUZON: Ilocos Norte, MINDANAO: Davao. Open sites at low elevation.
  2. Euploca strigosa (Willd.) Diane & Hilger, Bot. Jahrb. Syst. 125 (2003) 49; --Heliotropium strigosum Willd., Sp. Pl. 1 (1797) 743; --Merr., EPFP 3 (1923) 378. --Heliotropium brevifolium Wall. Fl. Ind., ed. 1820 2 (1824) 2; --Riedl, Fl. Males. ser. 1, 13 (1997) 102. Africa, Bangladesh, Bhutan, China, India, Laos, Middle East, Myanmar, Nepal, Pakistan, Sri Lanka, Thailand, Vietnam, Philippines. LUZON: Zambales. Open sandy places near the sea. Photos

Heliotropium L.

Last edited by Pieter B. Pelser, 14 July 2021

Photos
  1. Heliotropium arboreum (Blanco) Mabb., Mabberley's Pl.-Book, ed. 4 (2017) 1101; --Tournefortia arborea Blanco, Fl. Filip. (1837) 129. --Heliotropium foertherianum Diane & Hilger, Bot. Jahrb. Syst. 125 (2003) 46. --Tournefortia argentea L.f., Suppl. (1781) 133; --Merr., EPFP 3 (1923) 376; --Riedl, Fl. Males. ser. 1, 13 (1997) 117; --Argusia argentea (L.f.) Heine, Fl. New Caled. & Depend. 7 (1976) 108; --Dayang Awa in Tree Fl. Sabah & Sarawak 2 (1996) 95; --Messerschmiedia argentea (L.f.) I.M.Johnst., J. Arnold Arb. 16 (1935) 164; J. Arnold Arb. 32 (1951) 121; --Type: from Sri Lanka. Africa, Andaman Isls, Australia, Borneo, China, Indian Ocean, Java, Lesser Sunda Isls, Madagascar, Moluccas, New Caledonia, New Guinea, Nicobar Isls, Pacific, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Vietnam, Philippines. BATANES, LUZON, MINDANAO, PALAWAN, SULU ARCH. Along sandy seashores. Photos
  2. Heliotropium biblianum Craven, Blumea 50 (2005) 379; --Tournefortia tetrandra Blume, Bijdr. (1826) 845; I.M.Johnst., J. Arnold Arb. 16 (1935) 150; --Riedl, Fl. Males. ser. 1, 13 (1997) 123. Nicobars, Sri Lanka, Anambas & Natuna Isls, Sumatra, Peninsular Malaysia, Borneo, Java, New Guinea, Philippines. Montane forests and scrubs.
  3. Heliotropium curassavicum L., Sp. Pl. (1753) 130; --C.B.Rob., PJS 4 c (1909) 695; --Riedl, Fl. Males. ser. 1, 13 (1997) 103. Native of the Americas from Patagonia to the US and the West Indies, Java (near Bogor), Philippines. Occasionally occurring as weed. Photos
  4. Heliotropium indicum L., Sp. Pl. (1753) 130; --Merr., Fl. Manila (1912) 395; EPFP 3 (1923) 377; --I.M.Johnst., J. Arnold Arb. 32 (1951) 111; --Riedl, Fl. Males. ser. 1, 13 (1997) 105. Probably a native of tropical America, now pantropical. In Malesia: Sumatra, Peninsular Malaysia, Singapore, Borneo, Java, Philippines, Sulawesi, Lesser Sunda Isls (Lombok, Sumbawa, Sumba, Flores, Timor), Moluccas and New Guinea. A common weed in the settled areas throughout the Philippines at low elevation. Photos
  5. Heliotropium luzonicum (I.M. Johnst.) Craven, Blumea 50 (2005) 379; --Tournefortia luzonica I.M.Johnst., J. Arnold Arb. 16 (1935) 156; --Riedl, Fl. Males. ser. 1, 13 (1997) 118; --Type: Adduru 237 (A, K), Luzon.
  6. Heliotropium muelleri (I.M. Johnst.) Craven, Blumea 50 (2005) 380; --Tournefortia muelleri I.M.Johnst., J. Arnold Arb. 16 (1935) 157; --Riedl, Fl. Males. ser. 1, 13  (1997) 120; --Type: from Australia. N Australia, ?Java, Lesser Sunda Isls (?Bali, Sumba), New Guinea & Philippines. Along rivers and in tidal areas. 
  7. Heliotropium sarmentosum (Lam.) Craven, Blumea 50 (2005) 380; --Tournefortia sarmentosa Lam., Tabl. Encycl. 1 (1797) 416; --Merr., Fl. Manila (1912) 394; EPFP 3 (1923) 377; --I.M.Johnst., J. Arnold Arb. 16 (1935) 147; J. Arnold Arb. 32 (1951) 115; --Riedl, Fl. Males. ser. 1, 13 (1997) 121. --Tournefortia horsfieldii Miq., Fl. Ind. Bat. 2 (1858) 927; --Merr., EPFP 3 (1923) 376. Australia, Taiwan, Sumatra, Malay Peninsula, Borneo, Java, Philippines, Celebes, Lesser Sunda Islands (Lombok, Flores, Timor, Alor, Wetar), Moluccas, New Guinea. Photos

Messerschmidia Hebenstreit = Heliotropium

Tournefortia L. = Heliotropium


Literature

Craven, LA. 2005. Malesian and Australian Tournefortia transferred to Heliotropium and notes on delimitation of Boraginaceae. Blumea 50: 375-381

Frohlich, MW, M Thulin & MW Chase. 2020. Ninety-three new combinations in Euploca for species of Heliotropium section Orthostachys (Boraginaceae sensu APG). Phytotaxa 434: 13-21.

Hilger HH & N Diane. 2003. A systematic analysis of Heliotropiaceae (Boraginales) based on trnL and ITS1 sequence data. Botanische Jahrbücher 125: 19-51

Johnston, IM. 1951. Studies in the Boraginaceae. Journal of Arnold Arboretum 32: 1-26; 99-122.

Luebert, F, G Brokamp, J Wen, M Weigend & HH Hilger. 2011. Phylogenetic relationships and morphological diversity in Neotropical Heliotropium (Heliotropiaceae). Taxon 60: 663-680


Riedl, H. 1997. Boraginaceae. Flora Malesiana series 1, 13: 43-144

Pelser, P.B., J.F. Barcelona & D.L. Nickrent (eds.). 2011 onwards. Co's Digital Flora of the Philippines. www.philippineplants.org

Copyright © 2011, Co's Digital Flora of the Philippines

Last updated 14 July 2021