Heliotropiaceae

Euploca Nutt.

Last edited by Pieter B. Pelser, 29 May 2020

Euploca bracteata (R.Br.) M.W.Frohl. & M.W.Chase, Phytotaxa 434 (2020) 15; ≡ Heliotropium bracteatum R.Br., Prodr. (1810) 493; Merr., EPFP 3 (1923) 377. Distribution: Australia, Java, Pacific Ocean. LUZON: Ilocos Norte, MINDANAO: Davao. Notes: Open sites at low elevation. Native.
Euploca strigosa (Willd.) Diane & Hilger, Bot. Jahrb. Syst. 125 (2003) 49; ≡ Heliotropium strigosum Willd., Sp. Pl. 1 (1797) 743; Merr., EPFP 3 (1923) 378. = Heliotropium brevifolium Wall., Fl. Ind., ed. 1820 2 (1824) 2; Riedl, Fl. Males. ser. 1, 13 (1997) 102. Distribution: Africa, Bangladesh, Bhutan, China, India, Laos, Middle East, Myanmar, Nepal, Pakistan, Philippines, Sri Lanka, Thailand, Vietnam. LUZON: Cagayan, NCR, Zambales. Notes: Open sandy places near the sea. Native.

Heliotropium L.

Last edited by Pieter B. Pelser, 14 July 2021

Heliotropium arboreum (Blanco) Mabb., Mabberley's Pl.-Book, ed. 4 (2017) 1101; ≡ Tournefortia arborea Blanco, Fl. Filip. (1837) 129. = Heliotropium foertherianum Diane & Hilger, Bot. Jahrb. Syst. 125 (2003) 46. = Tournefortia argentea L.f., Suppl. (1781) 133; Merr., EPFP 3 (1923) 376; Riedl, Fl. Males. ser. 1, 13 (1997) 117; ≡ Argusia argentea (L.f.) Heine, Fl. New Caled. & Depend. 7 (1976) 108; Dayang Awa in Tree Fl. Sabah & Sarawak 2 (1996) 95; ≡ Messerschmidia argentea (L.f.) I.M.Johnst., J. Arnold Arb. 16 (1935) 164; J. Arnold Arb. 32 (1951) 121; Type: from Sri Lanka. Distribution: Africa, Andaman Isls, Australia, Borneo, China, Indian Ocean, Java, Lesser Sunda Isls, Madagascar, Malay Peninsula, Moluccas, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Ryukyu Isls, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Vietnam. BATANES, LUZON, MINDANAO, PALAWAN, SULU ARCHIPELAGO. Notes: Along sandy seashores. Native.
Heliotropium biblianum Craven, Blumea 50 (2005) 379; ≡ Tournefortia tetrandra Blume, Bijdr. (1826) 845; I.M.Johnst., J. Arnold Arb. 16 (1935) 150; Riedl, Fl. Males. ser. 1, 13 (1997) 123. Distribution: Borneo, Java, Malay Peninsula, New Guinea, Nicobar Isls, Philippines, Sri Lanka, Sumatra. Notes: Montane forests and scrubs. Native.
Heliotropium curassavicum L., Sp. Pl. (1753) 130; C.B.Rob., PJS 4 c (1909) 695; Riedl, Fl. Males. ser. 1, 13 (1997) 103. Distribution: America, Caribbean. Notes: Occasionally occurring as weed. Naturalized.
Heliotropium indicum L., Sp. Pl. (1753) 130; Merr., Fl. Manila (1912) 395; EPFP 3 (1923) 377; I.M.Johnst., J. Arnold Arb. 32 (1951) 111; Riedl, Fl. Males. ser. 1, 13 (1997) 105. Distribution: ?Neotropics. Notes: Now pantropical. A common weed in the settled areas throughout the Philippines at low elevation. Naturalized.
Heliotropium luzonicum (I.M.Johnst.) Craven, Blumea 50 (2005) 379; ≡ Tournefortia luzonica I.M.Johnst., J. Arnold Arb. 16 (1935) 156; Riedl, Fl. Males. ser. 1, 13 (1997) 118; Type: Adduru 237 (A, K), Luzon. Distribution: LUZON: Benguet, Cagayan, Ilocos Sur, Zambales, MINDANAO: South Cotabato, MINDORO. Native.
- ssp. luzonicum. Distribution: New Guinea, Philippines. LUZON. Notes: In mountainous regions. Native.
- ssp. angustissimum (Riedl) Craven, Blumea 50 (2005) 379; ≡ Tournefortia luzonica ssp. angustissima Riedl, Blumea 41 (1996) 443; Fl. Males. ser. 1, 13 (1997) 119; Type: Ridsdale 1477 (L, holo), Luzon: Zambales, Sta Cruz, Acoje Mine concession area. Distribution: Endemic to the Philippines. LUZON: Zambales. Notes: On ultramafic soil. Only known from type locality. Native.
- ssp. sublucens (I.M.Johnst.) Craven, Blumea 50 (2005) 379; ≡ Tournefortia luzonica ssp. sublucens (I.M.Johnst.) Riedl, Blumea 41 (1996) 443; Fl. Males. ser. 1, 13 (1997) 119; Type: BS 44553 Ramos & Edaño (A, BM, K), Luzon: Zambales prov. Distribution: Endemic to the Philippines. LUZON: Zambales. Native.
Heliotropium muelleri (I.M.Johnst.) Craven, Blumea 50 (2005) 380; ≡ Tournefortia muelleri I.M.Johnst., J. Arnold Arb. 16 (1935) 157; Riedl, Fl. Males. ser. 1, 13 (1997) 120; Type: from Australia. Distribution: Australia, ?Java, Lesser Sunda Isls, New Guinea, Philippines. CATANDUANES. Notes: Along rivers and in tidal areas. Native.
Heliotropium sarmentosum (Lam.) Craven, Blumea 50 (2005) 380; ≡ Tournefortia sarmentosa Lam., Tabl. Encycl. 1 (1797) 416; Merr., Fl. Manila (1912) 394; EPFP 3 (1923) 377; I.M.Johnst., J. Arnold Arb. 16 (1935) 147; J. Arnold Arb. 32 (1951) 115; Riedl, Fl. Males. ser. 1, 13 (1997) 121. = Tournefortia horsfieldii Miq., Fl. Ind. Bat. 2 (1858) 927; Merr., EPFP 3 (1923) 376. Distribution: Australia, Borneo, Java, Lesser Sunda Isls, Malay Peninsula, Moluccas, New Guinea, Philippines, Sulawesi, Sumatra, Taiwan. BASILAN, BILIRAN, LUBANG, LUZON: Aurora, Bataan, Benguet, Cagayan, Cavite, Ilocos Norte, Ilocos Sur, La Union, Laguna, Mountain Province, NCR, Pampanga, Pangasinan, Quezon, Rizal, Zambales, MASBATE, MINDANAO: Bukidnon, Davao del Sur, MINDORO, PALAWAN, POLILLO, PUJADA, TAWI-TAWI, UBIAN. Native.

References

Craven, LA. 2005. Malesian and Australian Tournefortia transferred to Heliotropium and notes on delimitation of Boraginaceae. Blumea 50: 375-381
Frohlich, MW, M Thulin & MW Chase. 2020. Ninety-three new combinations in Euploca for species of Heliotropium section Orthostachys (Boraginaceae sensu APG). Phytotaxa 434: 13-21.
Hilger HH & N Diane. 2003. A systematic analysis of Heliotropiaceae (Boraginales) based on trnL and ITS1 sequence data. Botanische Jahrbücher 125: 19-51
Johnston, IM. 1951. Studies in the Boraginaceae. Journal of Arnold Arboretum 32: 1-26; 99-122.
Luebert, F, G Brokamp, J Wen, M Weigend & HH Hilger. 2011. Phylogenetic relationships and morphological diversity in Neotropical Heliotropium (Heliotropiaceae). Taxon 60: 663-680
Riedl, H. 1997. Boraginaceae. Flora Malesiana series 1, 13: 43-144