Heliotropiaceae
Euploca
Last edited by Pieter B. Pelser, 29 May 2020
- Euploca bracteata (R.Br.) M.W.Frohl. & M.W.Chase, Phytotaxa 434 (2020) 15; ≡ Heliotropium bracteatum R.Br., Prodr. Fl. Nov. Holland. (1810) 493; Merrill, EPFP 3 (1923) 377. Distribution: Australia, Java, Pacific Ocean. LUZON: Ilocos Norte, MINDANAO: Davao. Notes: Open sites at low elevation. Native.
- Euploca strigosa (Willd.) Diane & Hilger, Bot. Jahrb. Syst. 125 (2003) 49; ≡ Heliotropium strigosum Willd., Sp. Pl., 4th Ed. 1 (1797) 743; Merrill, EPFP 3 (1923) 378. = Heliotropium brevifolium Wall., Fl. Ind. (Carey & Wallich) 2 (1824) 2; Riedl, Fl. Males. ser. 1, 13 (1997) 102. Distribution: Africa, Bangladesh, Bhutan, China, India, Laos, Middle East, Myanmar, Nepal, Pakistan, Philippines, Sri Lanka, Thailand, Vietnam. LUZON: Cagayan, Ilocos Norte (photos), NCR, Zambales. Notes: Open sandy places near the sea. Native.
Heliotropium
Last edited by Pieter B. Pelser, 30 March 2025
- Heliotropium arboreum (Blanco) Mabb., Mabberley's Pl.-Book, 4th Ed. (2017) 1101; ≡ Tournefortia arborea Blanco, Fl. Filip., 1st Ed. (1837) 129. = Heliotropium foertherianum Diane & Hilger, Bot. Jahrb. Syst. 125 (2003) 46. = Tournefortia argentea L.f., Suppl. Pl. (1782) 133; Merrill, EPFP 3 (1923) 376; Riedl, Fl. Males. ser. 1, 13 (1997) 117; Romeroso et al., Asian J. Biol. Life Sc. 10 (2021) 437; ≡ Argusia argentea (L.f.) Heine, Fl. Nouv.-Calédonie & Dépend. 7 (1976) 108; Dayang Awa in Tree Fl. Sabah & Sarawak 2 (1996) 95; ≡ Messerschmidia argentea (L.f.) I.M.Johnst., J. Arnold Arbor. 16 (1935) 164; J. Arnold Arb. 32 (1951) 121; Type: from Sri Lanka. Distribution: Africa, Andaman Isls, Australia, Borneo, China, Indian Ocean, Java, Lesser Sunda Isls, Madagascar, Malay Peninsula, Moluccas, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Ryukyu Isls, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Vietnam. BALABAC, BATANES, BORACAY (photos), BUYAYAO (photos), DANJUGAN (photos), DIGYO ISL (photos), HOMONHON, LUMBUCAN, LUZON: Aurora (photos), MANTIGUE, MINDANAO: Misamis Oriental (photos), MINILOC, PALAWAN, SIBUYAN (photos), SULU ARCHIPELAGO, TURTLE IS. Notes: Along sandy seashores. Sea level. 0-2m. Conservation status: Least Concern (IUCN 3.1). Native.
- Heliotropium biblianum Craven, Blumea 50 (2005) 379; ≡ Tournefortia tetrandra Blume, Bijdr. fl. Ned. Ind. (1826) 845; I.M.Johnst., J. Arnold Arb. 16 (1935) 150; Riedl, Fl. Males. ser. 1, 13 (1997) 123. Distribution: Borneo, Java, Malay Peninsula, New Guinea, Nicobar Isls, Philippines, Sri Lanka, Sumatra. Notes: Montane forests and scrubs. Native.
- Heliotropium curassavicum L., Sp. Pl., 1st Ed. (1753) 130; Robinson, PJS 4c (1909) 695; Riedl, Fl. Males. ser. 1, 13 (1997) 103. Distribution: America, Caribbean. Notes: Occasionally occurring as weed. Naturalized.
- Heliotropium indicum L., Sp. Pl., 1st Ed. (1753) 130; Merr., Fl. Manila (1912) 395; Merrill, EPFP 3 (1923) 377; I.M.Johnst., J. Arnold Arb. 32 (1951) 111; Riedl, Fl. Males. ser. 1, 13 (1997) 105. Distribution: ?Neotropics. CUYO, LUZON: Cagayan (photos), PANAY (photos). Notes: Now pantropical. A common weed in the settled areas throughout the Philippines at low elevation. Naturalized.
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Heliotropium luzonicum (I.M.Johnst.) Craven, Blumea 50 (2005) 379;
≡ Tournefortia luzonica I.M.Johnst., J. Arnold Arbor. 16 (1935) 156; Riedl, Fl. Males. ser. 1, 13 (1997) 118;
Type: Adduru 237 (A, K), Luzon.
Distribution:
LUZON: Benguet, Cagayan, Ilocos Sur, Zambales, MINDANAO: South Cotabato, MINDORO.
Native.
- ssp. luzonicum. Distribution: New Guinea, Philippines. LUZON. Notes: In mountainous regions. Native.
- ssp. angustissimum (Riedl) Craven, Blumea 50 (2005) 379; ≡ Tournefortia luzonica ssp. angustissima Riedl, Blumea 41 (1996) 443; Fl. Males. ser. 1, 13 (1997) 119; Type: Ridsdale 1477 (L, holo), Luzon: Zambales, Sta Cruz, Acoje Mine concession area. Distribution: Endemic to the Philippines. LUZON: Zambales. Notes: On ultramafic soil. Only known from type locality. Native.
- ssp. sublucens (I.M.Johnst.) Craven, Blumea 50 (2005) 379; ≡ Tournefortia luzonica ssp. sublucens (I.M.Johnst.) Riedl, Blumea 41 (1996) 443; Fl. Males. ser. 1, 13 (1997) 119; Type: BS 44553 Ramos & Edaño (A, BM, K), Luzon: Zambales prov. Distribution: Endemic to the Philippines. LUZON: Zambales. Native.
- Heliotropium muelleri (I.M.Johnst.) Craven, Blumea 50 (2005) 380; ≡ Tournefortia muelleri I.M.Johnst., J. Arnold Arbor. 16 (1935) 157; Riedl, Fl. Males. ser. 1, 13 (1997) 120; Type: from Australia. Distribution: Australia, ?Java, Lesser Sunda Isls, New Guinea, Philippines. CATANDUANES, ?LUZON: ?Cagayan, ?NEGROS. Notes: Along rivers and in tidal areas. Native.
- Heliotropium sarmentosum (Lam.) Craven, Blumea 50 (2005) 380; ≡ Tournefortia sarmentosa Lam., Tabl. Encycl. 1 (1792) 416; Merr., Fl. Manila (1912) 394; Merrill, EPFP 3 (1923) 377; I.M.Johnst., J. Arnold Arb. 16 (1935) 147; J. Arnold Arb. 32 (1951) 115; Riedl, Fl. Males. ser. 1, 13 (1997) 121. = Tournefortia horsfieldii Miq., Fl. Ned. Ind. 2 (1858) 927; Merrill, EPFP 3 (1923) 376. Distribution: Australia, Borneo, Java, Lesser Sunda Isls, Malay Peninsula, Moluccas, New Guinea, Philippines, Sulawesi, Sumatra, Taiwan. BALABAC, BASILAN, BILIRAN, LUBANG, LUZON: Aurora, Bataan, Benguet, Cagayan, Cavite, Ilocos Norte, Ilocos Sur, La Union, Laguna, Mountain Province, NCR, Pampanga, Pangasinan, Quezon, Rizal, Zambales, MASBATE, MINDANAO: Bukidnon, Davao del Sur, MINDORO, PALAWAN, PANAY (photos), POLILLO, PUJADA, TAWI-TAWI, UBIAN. Notes: 600m (photos). Conservation status: Least Concern (IUCN 3.1). Native.
References
- Craven, LA. 2005. Malesian and Australian Tournefortia transferred to Heliotropium and notes on delimitation of Boraginaceae. Blumea 50: 375-381
- Frohlich, MW, M Thulin & MW Chase. 2020. Ninety-three new combinations in Euploca for species of Heliotropium section Orthostachys (Boraginaceae sensu APG). Phytotaxa 434: 13-21.
- Hilger HH & N Diane. 2003. A systematic analysis of Heliotropiaceae (Boraginales) based on trnL and ITS1 sequence data. Botanische Jahrbücher 125: 19-51
- Johnston, IM. 1951. Studies in the Boraginaceae. Journal of Arnold Arboretum 32: 1-26; 99-122.
- Luebert, F, G Brokamp, J Wen, M Weigend & HH Hilger. 2011. Phylogenetic relationships and morphological diversity in Neotropical Heliotropium (Heliotropiaceae). Taxon 60: 663-680
- Riedl, H. 1997. Boraginaceae. Flora Malesiana series 1, 13: 43-144