
Gentianaceae
Canscora
Last edited by Pieter B. Pelser, 3 November 2017
- Canscora diffusa (Vahl) R.Br. ex Roem. & Schult., Syst. Veg. ed. 15 (1818) 301; Merr., Fl. Manila (1912) 367; EPFP 3 (1923) 318; Thiv., Blumea 48 (2003) 11; ≡ Gentiana diffusa Vahl, Symb. Bot. 3 (1794) 47. Distribution: Africa, Asia, Australia, Malesia. LUZON: Apayao, Bataan, Benguet, Bulacan, Cavite, Ilocos Norte, Mountain Province, NCR, Nueva Vizcaya, Pangasinan, Quezon, Rizal, Zambales, MINDORO, PANAY. Native.
- Canscora macrocalyx Miq., Fl. Ned.-ind. 2 (1856) 558; Thiv., Blumea 48 (2003) 15. Distribution: Java, Lesser Sunda Isls, Philippines. MINDANAO: Zamboanga del Norte. Notes: Moist open places in grasslands, fields; heavy soil. 0-600m. Fl. Jan-Aug. Native.
Cyrtophyllum
Edited by Pieter B. Pelser, 7 January 2015
- Cyrtophyllum fragrans (Roxb.) DC., Prodr. 9 (1845) 31; ≡ Fagraea fragrans Roxb., Fl. Ind. 2 (1824) 32; Merr., EPFP 3 (1923) 314; Leenh., Fl. Males. ser. 1, 6 (1962) 304; Wong & Sugau, Tree Fl. Sabah & Sarawak 2 (1996) 206. = ?Fagraea cochinchinensis (Lour.) A.Chev., Cat. Pl. Jard. Bot. Saigon (1919) 65; Merr., EPFP 3 (1923) 314. Distribution: Andaman Isls, Borneo, India, Java Sea, Malay Peninsula, Myanmar, New Guinea, Philippines, Sulawesi, Sumatra, Thailand, Vietnam. BALABAC, BUSUANGA, CULION, MINDORO, PALAWAN. Notes: Lowland forests, especially secondary and disturbed ones. Native.
Exacum
Last edited by Pieter B. Pelser, 24 August 2019
- Exacum loheri (H.Hara) Klack., Bot. Jahrb. Syst. 126 (2006) 478; ≡ Cotylanthera loheri Hara, J. Jap. Bot. 50 (1975) 327; Type: Loher 5189 (K, holo), Luzon: Montalban, 26-Oct-1895. Distribution: Endemic to the Philippines. LUZON: Rizal. Native.
- Exacum parviflorum Merr., PJS 7 c (1912) Bot. 96; EPFP 3 (1923) 317; Type: Vanoverbergh 758. Distribution: Endemic to the Philippines. LUZON: Mountain Province. Notes: Open slopes, c. 1200m. Native.
- Exacum tenue (Blume) Klack., Bot. Jahrb. Syst. 126(4) (2006) 478; ≡ Cotylanthera tenuis Blume, Bijdr. (1825) 707; Merr., PJS 1 (1906) Suppl. 223; EPFP 3 (1923) 317; Type: from Java. = Eophylon tenellum A.Gray, J. Linn. Soc., Bot. 11 (1869) 23. Distribution: Borneo, Java, Philippines. LUZON: Bataan, Laguna, Mountain Province, Rizal, Sorsogon, MANGSEE, MINDANAO, NEGROS, PALAWAN, PANAY. Notes: Low and medium elevation damp primary forests, ascending to 1700m. Native.
- Exacum tetragonum Roxb., Fl. Ind. 1 (1820) 413. = Exacum chironioides Griseb., Gen. Sp. Gent. (1839) 109; Fern.-Vill., Novis. App. (1880) 136; Merr., Sp. Blancoanae (1918) 307; EPFP 3 (1923) 317. Distribution: Malesia. CULION, LUZON: Albay, Cagayan, Ilocos Sur, NCR, Nueva Vizcaya, Pangasinan, Rizal, Zambales, MINDANAO: Bukidnon, Cotabato, Davao, Davao del Sur, MINDORO, PANAY. Notes: Low and medium elevation wet grasslands, swamps, etc. Native.
Fagraea
Last edited by Pieter B. Pelser, 19 March 2023
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Fagraea auriculata Jack, Malay Misc. 2, No. 7 (1822) 82; Elmer, LPB 2 (1909) 509; Merr., EPFP 3 (1923) 314; Leenh., Fl. Males. ser. 1, 6 (1962) 326; Coritico et al., J. Trop. Biol. & Cons. 19 (2022) 149.
= Fagraea epiphytica Elmer, LPB 8 (1915) 2743.
Distribution:
LEYTE, LUZON: Laguna, Quezon, MINDANAO: Agusan, Agusan del Norte, Davao del Sur, MINDORO, PALAWAN, PANAY, SAMAR.
Native.
- ssp. auriculata. Distribution: Borneo, Indochina, Java, Lesser Sunda Isls, Malay Peninsula, Myanmar, Philippines, Sumatra, Thailand. LEYTE, MINDANAO: Agusan del Norte, PANAY, SAMAR. Native.
- ssp. parviflora Leenh., Fl. Males. ser. 1, 6 (1962) 328. = Fagraea curranii Merr., PJS 13 c (1918) Bot. 50; EPFP 3 (1923) 314. Distribution: Moluccas, Philippines. LUZON: Laguna, Quezon, MINDORO. Native.
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Fagraea blumei G.Don, Gard. Dict. 4 (1837) 69, as ‘blumii’; Leenh., Fl. Males. ser. 1, 6 (1962) 320; Wong & Sugau, Tree Fl. Sabah & Sarawak 2 (1996) 202.
= Fagraea cuernosensis Elmer, LPB 2 (1909) 600; Merr., EPFP 3 (1923) 314.
= Fagraea negrosensis Elmer, LPB 2 (1909) 599; Merr., EPFP 3 (1923) 315.
≠ Fagraea obovata auct. non Wall.; Miq., Fl. Ind. Bat. 2 (1857) 369; Merr., EPFP 3 (1923) 315, pro parte.
Distribution:
CAMIGUIN, CATANDUANES, LEYTE, LUZON: Zambales, MINDANAO: Cotabato, Davao del Sur, Misamis Occidental, MINDORO, NEGROS, PANAY, SAMAR.
Native.
- ssp. blumei. Distribution: Borneo, Java, Malay Peninsula, Philippines, Sumatra. NEGROS. Native.
- ssp. plumeriiflora (DC.) Leenh., Fl. Males. ser. 1, 6 (1962) 323; ≡ Fagraea plumeriiflora DC., Prodr. 9 (1845) 29; Elmer, LPB 2 (1909) 598; LPB 3 (1910) 857; Merr., EPFP 3 (1923) 315, excl. specimens from Sulawesi. Distribution: Endemic to the Philippines. CATANDUANES, LEYTE, LUZON, MINDANAO, PANAY, SAMAR. Notes: Intermediate forms between ssp. plumeriiflora and ssp. blumei occur in Borneo, Negros and Mindanao. Native.
- Fagraea borneensis Scheff. in Hassk., Flora 32 (1869) 309; Wong & Sugau, Tree Fl. Sabah & Sarawak 2 (1996) 202; ≡ Fagraea auriculata ssp. borneensis (Scheff.) Leenh., Fl. Males. ser. 1, 6 (1962) 328; Type: from Sarawak. = Fagraea nonok Elmer, LPB 3(1910) 858; Merr., EPFP 3 (1923) 315. = Fagraea jackii Elmer, LPB 8 (1915) 2744. Distribution: Borneo, Lesser Sunda Isls, Philippines. MINDORO, PALAWAN, SIBUYAN. Notes: Lowland and montane forests. Native.
- Fagraea ceilanica Thunb., Vet. Acad. Handl. Stockh. 3 (1782) 132; Leenh., Fl. Males. ser. 1, 6 (1962) 315. = Fagraea obovata Wall. in Roxb., Fl. Ind. 2 (1824) 33; Merr., EPFP 3 (1923) 315, pro parte. = Fagraea loheri Merr., PJS 27 (1925) 49; EPFP 4 (1925) 254; Type: Loher. Distribution: China, Hong Kong, India, Indochina, Myanmar, Solomon Isls, Sri Lanka, Taiwan, Thailand. BATAN, BILIRAN, CAMIGUIN, CATANDUANES, LEYTE, LUZON: Albay, Bataan, Benguet, Camarines Sur, Ifugao, Isabela, Laguna, Mountain Province, Pampanga, Pangasinan, Quezon, Rizal, Sorsogon, Zambales, MALAPACKUN, MINDANAO: Agusan del Norte, Bukidnon, Davao, Surigao del Norte, Zamboanga, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY, ROMBLON, SAMAR, SIBUYAN, SULU ARCHIPELAGO, TAWI-TAWI. Notes: Throughout Malesia, throughout the Philippines; high-climbing woody hemi-epiphyte in forests, 0-2100m. Native.
- Fagraea gitingensis Elmer, LPB 3 (1910) 859; Merr., EPFP 3 (1923) 315; Leenh., Fl. Males. ser. 1, 6 (1962) 334. Distribution: Moluccas, Philippines. SIBUYAN. Native.
- Fagraea longiflora Merr., PJS 3 c (1908) Bot. 260; Elmer, LPB 2 (1909) 597; Merr., EPFP 3 (1923) 315. = Fagraea macgregorii Merr., PJS 13 c (1918) Bot. 51. Distribution: Endemic to the Philippines. CATANDUANES, LUZON: Apayao, Aurora, Camarines Sur, Ifugao, Isabela, Laguna, Quezon, MINDANAO: Bukidnon. Notes: A dubious specimen from Mindanao (BS 38650). Native.
Gentiana
Last edited by Pieter B. Pelser, 24 May 2018
- Gentiana diversifolia Merr., PJS 1 (1906) Suppl. 224; Merr., EPFP 3 (1923) 319; Type: Merrill 4443 (PNH, holo, ?lost), Luzon: Benguet, Kabayan. On damp, rocky banks, in open lands, rare, c. 1300m, 27-Oct-1905. Distribution: Endemic to the Philippines. LUZON: Benguet. Notes: Damp rocky banks among grasses, 1200-1900m. Native.
- Gentiana elmeriana Halda, Acta Mus. Richnov., Sect. Nat. 3(1) (1995) 46; ≡ Gentiana copelandii Elmer, LPB 7 (1915) 2671, nom. illeg., non Greene; Merr., EPFP 3 (1923) 319; Type: Elmer 11692. Distribution: Endemic to the Philippines. MINDANAO: Bukidnon, Davao, Davao del Sur. Notes: In mosses at the summit, c. 2500m. Native.
- Gentiana quadrifaria Blume, Bijdr. Fl. Ned. Ind. 14 (1826) 847. = Gentiana apoensis Merr., PJS 1 (1906) Suppl. 223; Merr., EPFP 3 (1923) 318; Type: Copeland 1161 & 1432 (PNH, ?lost, syn), Mindanao: Davao, Mt Apo, summit area, c. 3000m, occasional in turf, but inconspicuous, not common, Apr/Oct-1904; de Vore & Hoover 371 (PNH, ?lost, syn), Mindanao: Davao, Mt Apo, May-1903. = Gentiana luzoniensis Merr., PJS 1 (1906) Suppl. 224; Merr., EPFP 3 (1923) 319; Type: Merrill 4558 (NY, US, iso), Luzon: Mountain Province, Mt Data, on dry, open grassy slopes in thin pine forests at c. 2100m, 4-Nov-1905. Distribution: India, Java, Myanmar, Philippines, Sri Lanka, Taiwan. LUZON: Benguet, Ifugao, Laguna, Mountain Province, Nueva Vizcaya, Quezon, MINDANAO: Cotabato, Davao, Davao del Sur, NEGROS. Notes: Open grasslands and exposed ridges on the higher mountains, 1700-2800m. Native.
Hoppea (Not native)
- Hoppea dichotoma Willd., Ges. Naturf. Freunde Berlin Neue Schriften 3 (1801) 434; Merr., EPFP 3 (1923) 318; Thiv, Blumea 48 (2003) 28. Distribution: India. LUZON: Abra, Benguet, Ilocos Norte, Pangasinan. Notes: Thiv argues that outside of India, this plant is non indigenous in occurrence. In the Philippines reported from open places at low and medium elevation, ascending to 1800m in Benguet. Rare or local. Naturalized.
Microrphium
Last edited by Pieter B. Pelser, 24 May 2018
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Microrphium pubescens C.B.Clarke, J. Asiat. Soc. Bengal 74 (1906) 88.
Distribution:
Malay Peninsula, Philippines, Thailand.
MINDORO, PALAWAN.
Native.
- ssp. elmerianum (Regalado & Soejarto) Thiv, Blumea 48 (2003) 33; ≡ Microrphium elmerianum Regalado & Soejarto, Novon 7 (1997) 77; Type: Soejarto et al. 6499 (F, holo; A, PNH, iso), Palawan: Puerto Princesa, Bacungan Hills, slopes above Nagtabon beach, 9°51’N, 118°37’E. Distribution: Endemic to the Philippines. PALAWAN. Notes: Fl. Feb-Aug. Native.
Schenkia (Not native)
Last edited by Pieter B. Pelser, 30 October 2019
- Schenkia spicata (L.) G.Mans., Taxon 53 (2004) 726; ≡ Centaurium spicatum (L.) Fritsch, Mitt. Naturwiss. Vereins Univ. Wien n.s., 5 (1907) 97; ≡ Erythraea spicata (L.) Pers., Syn. 1 (1905) 283; Merr., EPFP 3 (1923) 317; ≡ Gentiana spicata L., Sp. Pl. 1 (1753) 230. Distribution: Europe. BATAN. Notes: Introduced into Taiwan by early Portuguese colonists, extending thereafter southward to the Batanes Isls Group. Open places at low elevation. Naturalized.
Swertia
Last edited by Pieter B. Pelser, 24 May 2018
- Swertia decurrens C.B.Rob., PJS 3 c (1909) Bot. 214; Merr., EPFP 3 (1923) 319; Type: R.S. Williams 1529. Distribution: Endemic to the Philippines. LUZON: Benguet, Ifugao, Mountain Province. Notes: Open slopes, extending into the mossy forest, 1200-2300m. Native.
Tripterospermum
Last edited by Pieter B. Pelser, 1 August 2017
- Tripterospermum luzoniense (S.Vidal) J.Murata, J. Fac. Sci. Univ. Tokyo 14 (1989) 305; Wang & Chen in Huang, Fl. Taiwan, ed. 2, 4 (1998) 186; ≡ Crawfurdia luzoniensis S.Vidal, Rev. Pl. Vasc. Filip. (1886) 191; Merr., EPFP 3 (1923) 318; Type: Vidal 1228 (MA), Luzon: Lepanto. Distribution: Java, Philippines, Taiwan. LUZON: Benguet, Laguna, Mountain Province, Quezon, MINDANAO: Bukidnon. Notes: Mossy forests in very damp shaded ravines, 1100-2400m. Native.
Utania
Last edited by Pieter B. Pelser, 31 October 2023
- Utania cuspidata (Blume) K.M.Wong, Sugumaran & Sugau, Gard. Bull. Singapore 65 (2013) 237; ≡ Fagraea cuspidata Blume, Ann. Mus. Bot. Lugd.-Bat. 1 (1850) 170; Merr., EPFP 3 (1923) 314; Wong & Sugau, Tree Fl. Sabah & Sarawak 2 (1996) 204. Distribution: Borneo, Philippines. BALABAC. Notes: Mixed dipterocarp forests, in gaps and clearings, also secondary forests and forest edges. Native.
- Utania philippinensis (K.M.Wong & Sugau) K.M.Wong, Sugumaran & Sugau, Gard. Bull. Singapore 65 (2013) 237; ≡ Fagraea philippinensis K.M.Wong & Sugau, Sandakania 8 (1996) 35; Type: FB 3270 Ahern’s Collector (K, holo). ≠ Fagraea racemosa auct. non Jack ex Wall.; Leenh., Fl. Males. ser. 1, 6 (1962) 311, pro parte. Distribution: Borneo, Philippines. LUZON: Rizal, MINDANAO: Misamis Occidental. Native.
- Utania spicata (Baker) K.M.Wong, Sugumaran & Sugau, Gard. Bull. Singapore 65 (2013) 238; Malabrigo et al., Flora of Mt Mantalingahan Protected Landscape (2023) 120; ≡ Fagraea spicata Baker, Kew Bull. (1896) 25; Wong & Sugau, Tree Fl. Sabah & Sarawak 2 (1996) 214. = Fagraea congestiflora Elmer, LPB 8 (1915) 2741; Merr., EPFP 3 (1923) 316. ≠ Fagraea racemosa auct. non Jack ex Wall.; Leenh., Fl. Males. ser. 1, 6 (1962) 311. Distribution: Borneo, Philippines. PALAWAN. Notes: Lowland to montane forests, frequently in gaps and also secondary forests. Native.
- Utania volubilis (Wall.) Sugumaran, Gard. Bull. Singapore 64 (2012) 493; ≡ Fagraea volubilis Wall. in Roxb., Fl. Ind. 2 (1824) 36; Wong & Sugau, Tree Fl. Sabah & Sarawak 2 (1996) 215. = Fagraea scholaris Blanco, Fl. Filip. ed. 2 (1845) 93; Fl. Filip. ed. 3, 1 (1877) 171. ≠ Fagraea racemosa auct. non Jack ex Wall.; Merr., EPFP 3 (1923) 315; Leenh., Fl. Males. ser. 1, 6 (1962) 311. Distribution: Andaman Isls, Borneo, Java, Malay Peninsula, Moluccas, New Guinea, Nicobar Isls, Philippines, Sulawesi, Sumatra. LUZON: Rizal, MINDANAO: Agusan del Norte, MINDORO, PANAY. Notes: Lowland forests, including secondary forests and forest edges. Native.
References
- Backer, CA & Bakhuizen van den Brink. 1965. Flora of Java 2: 437-441.
- Geesink, R. 1973. A synopsis of the genus Swertia (Gent.) in Malesia. Blumea 21: 179-183
- Klackenberg, J. 2006. Cotylanthera transferred to Exacum (Gentianaceae). Botanische Jahrbücher 126: 477-481.
- Leenhouts, PW. 1962. Loganiaceae. Flora Malesiana, Series 1 6: 293-387 (includes Fagraea)
- Mansion, G. 2004. A new classification of the polyphyletic genus Centaurium Hill (Chironiinae, Gentianaceae): Description of the New World endenic Zeltnera, and reinstatement of Gyrandra Griseb. and Schenkia Griseb. Taxon 53: 719-740
- Murata, J. 1989. A synopsis of Tripterospermum (Gentianaceae). Journal of the Faculty of Science, Univ. Tokyo Sec. III, 16 (4): 273-339.
- Royen, P van. 1964. Nova Guinea Bot. 17: 369-416.
- Sugumaran, M & KM Wong. 2012. Studies in Malesian Gentianaceae I: Fagraea sensu lato - complex genus or several genera? A molecular phylogenetic study. Gardens' Bulletin Singapore 64: 301-332
- Sugumaran, M & KM Wong. 2014. Studies in Malesian Gentianaceae VI: A revision of Utania in the Malay Pensinsula with two new species. Plant Ecology and Evolution 147: 213-223
- Thiv, M. 2003. A taxonomic revision of Canscora, Cracosna, Duplipetala, Hoppea, Microrphium, Phylloccyclus and Schinziella (Gentianaceae-Canscorinae). Blumea 48: 1-46
- Wong, KM & JB Sugau. 1996. A revision of Fagraea (Loganiaceae) in Borneo, with notes on related Malesian species and 21 new species. Sandakania 8: 1-93.
- Wong, KM & M Sugumaran. 2012. Studies in Malesian Gentianaceae II: A taxonomic framework for the Fagraea complex, including the new genus Limahlania. Gardens' Bulletin Singapore 64: 481-495
- Wong, KM & M Sugumaran. 2012. Studies in Malesian Gentianaceae III: Cyrtophyllum reapplied to the Fagraea fragrans alliance. Gardens' Bulletin Singapore 64: 497-510
- Wong, KM & M Sugumaran. 2016. Studies in Malesian Gentianaceae, VII. A revision of Fagraea in the Malay Peninsula with five new species. Sandakania 21: 65-130.
- Wong, KM, M Sugumaran & JB Sugau. 2013. Studies in Malesian Gentianaceae V: The Fagraea complex in Borneo: New generic assignments and recombinations. Gardens' Bulletin Singapore 65; 235-239.