Dryopteridaceae

Elaphoglossaceae, Peranemataceae, Bolbitidaceae, Aspidiaceae

Arachniodes Blume

Last edited by Julie F. Barcelona, 11 July 2021

Arachniodes amabilis (Blume) Tindale, Contr. New South Wales Natl. Herb. 3 (1961) 90; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22045; ≡ Rumohra amabilis (Blume) Ching, Sinensia 5 (1934) 41; ≡ Aspidium amabile Blume, Enum. Pl. Javae 2 (1828) 165. Distribution: China, India, Japan, Java, Korea, Lesser Sunda Isls, Myanmar, Philippines, Sri Lanka, Sulawesi, Taiwan, Vietnam. BILIRAN, CAMIGUIN, LUZON: Benguet, Ilocos Norte, Mountain Province, Pampanga, Quezon, Sorsogon, MINDANAO: Bukidnon, Davao del Sur, Sarangani, South Cotabato, NEGROS, PANAY. Native.
Arachniodes carvifolia (Kunze) Ching, Acta Bot. Sin. 10 (1962) 256; ≡ Aspidium carvifolium Kunze, Linn. 24 (1851) 292. = Arachniodes aristata auct. non (G.Forst.) Tindale; Rufila et al., Ruhuna J. Science 13 (2022) 157. Distribution: Australia, Bhutan, Borneo, China, India, Japan, Korea, Lesser Sunda Isls, Malay Peninsula, Myanmar, Nepal, New Guinea, Philippines, Ryukyu Isls, Sri Lanka, Sulawesi, Vietnam. LUZON: Bataan, Cagayan, Camarines Norte, Ilocos Norte, Laguna, Mountain Province, Pangasinan, Quezon, Rizal, Zambales, MINDANAO: Bukidnon, ?Davao del Sur, Zamboanga del Sur, MINDORO, NEGROS. Native.
Arachniodes cornu-cervi (D.Don) Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 323; ≡ Aspidium cornu-cervi D.Don, Prodr. Fl. Nepal. (1825) 5. Distribution: Bhutan, China, India, Korea, Myanmar, Nepal, ?Pacific Ocean, Philippines, ?Thailand, Vietnam. Native.
Arachniodes hasseltii (Blume) Ching, Acta Bot. Sin. 10 (1962) 258; ≡ Rumohra hasseltii (Blume) Ching, Sinensia 5 (1934) 61; ≡ Polypodium hasseltii Blume, Fl. Jav. Fil. 195 (1829) t.92. Distribution: Australia, Borneo, China, India, Java, ?Lesser Sunda Isls, Malay Peninsula, Moluccas, New Guinea, Pacific Ocean, Philippines, Ryukyu Isls, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam. Native.
Arachniodes tripinnata (Goldm.) Sledge, Bull. Brit. Mus. (Nat. Hist.) Bot. 5 (1973) 41; ≡ Polystichum tripinnatum Goldm., Nova Acta 19, Suppl. 1 (1843) 463. Distribution: Borneo, Java, Pacific Ocean, Philippines, Sulawesi. LUZON: Bataan, Batangas, Cagayan, Ilocos Norte, Laguna, NCR, Nueva Ecija, Pampanga, Quezon, NEGROS, PALAWAN. Native.

Possibly present

Arachniodes coniifolia (T.Moore) Ching, Acta Bot. Sin. 10 (1962) 257; ≡ Lastrea coniifolia T.Moore, Ind. Fil. (1857) 88. = Rumohra wallichii Ching, Sinensia 5 (1934) 56. Distribution: Bhutan, China, India, Myanmar, Nepal, ?Philippines, Thailand, Vietnam.
Arachniodes sledgei Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 323. Distribution: India, ?Philippines, Sri Lanka.

Arthrobotrya J.Sm.

Last edited by Julie F. Barcelona, 17 June 2019

Arthrobotrya articulata (J.Sm. ex Fée) J.Sm., Hist. Fil. (1875) 141; ≡ Teratophyllum articulatum (J.Sm. ex Fée) Mett. ex Kuhn, Ann. Mus. Bot. Lugduno-Batavi 4 (1869) 297; ≡ Polybotrya articulata J.Sm. ex Fée, Acrost. (1845) 74; J.Sm. in Hook., J. Bot. 3 (1841) 401, nom. inval., nom. nud. Distribution: Bismarck Arch, Moluccas, New Guinea, Philippines, Solomon Isls, Sulawesi. LEYTE, LUZON, MINDANAO, SAMAR. Native.

Bolbitis Schott

Last edited by Julie F. Barcelona & Pieter B. Pelser, 30 July 2022

Bolbitis angustipinna (Hayata) H.Ito, J. Jap. Bot. 14 (1938) 443; ≡ Leptochilus angustipinnus Hayata, Icon. Pl. Formosan. 5 (1915) 297. Distribution: Bhutan, China, India, Myanmar, Nepal, Philippines, Sri Lanka, Sulawesi, Taiwan, Thailand. LUZON: Ilocos Norte. Native.
Bolbitis appendiculata (Willd.) K.Iwats., Acta Phytotax. Geobot. 18 (1959) 48; ≡ Egenolfia appendiculata (Willd.) J.Sm., Ferns Brit. For. (1866) 111; ≡ Acrostichum appendiculatum Willd., Sp. Pl. 5 (1810) 114. Distribution: Bangladesh, Bhutan, Cambodia, China, India, Japan, Java, Laos, Malay Peninsula, Myanmar, Nicobar Isls, Philippines, Ryukyu Isls, Singapore, Sri Lanka, Sumatra, Taiwan, Thailand, Vietnam. BALABAC, LUZON: Aurora, Bataan, Batangas, Benguet, Laguna, Quezon, Rizal, Sorsogon, Zambales, PALAWAN. Native.
Bolbitis × arguta (Fée) Ching, Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 47. Notes: Hennipman doubts whether this fern warrants a separate treatment as a hybrid; it may be close to Edanyoa difformis (=Bolbitis heteroclita). Native.
Bolbitis heteroclita (C.Presl) Ching, Index Filic., Suppl. 3 (1934) 48; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22045; ≡ Poecilopteris heteroclita (C.Presl) C.Presl, Tent. Pterid. (1836) 242; ≡ Acrostichum heteroclitum C.Presl, Reliq. Haenk. 1 (1825) 15. = Bolbitis cuspidata (C.Presl) Ching, Index Filic., Suppl. 3 (1934) 48; ≡ Nephrodium cuspidatum C.Presl, Reliq. Haenk. 1 (1825) 31. = Bolbitis pseudoscalpturata (Copel.) Ching, Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 49; ≡ Campium pseudoscalpturatum Copel., PJS 37 (1928) 363. = Bolbitis foxworthyi (Copel.) Ching, Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 48; ≡ Campium foxworthyi Copel., PJS 37 (1928) 364. = Bolbitis tenuissima (Copel.) Copel., FFP 2 (1960) 259; ≡ Campium tenuissimum Copel., PJS 37 (1928) 364. = Bolbitis membranacea (Copel.) C.Chr., Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 198; ≡ Campium membranaceum Copel., PJS 40 (1929) 307. = Bolbitis edanyoi Copel., PJS 81 (1952) 22. = Edanyoa difformis Copel., PJS 81 (1952) 22. = Cyrtogonium acuminatum Brack., U.S. Expl. Exped., Filic. 16 (1854) 86. Distribution: Bangladesh, Borneo, China, India, Java, Malay Peninsula, Moluccas, Myanmar, Nepal, New Guinea, Pacific Ocean, Philippines, Ryukyu Isls, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. BABUYAN, BATAN, BOHOL, CAMIGUIN, GENERAL ISLAND, LEYTE, LUZON: Aurora, Bataan, Batangas, Benguet, Cagayan, Camarines Norte, Camarines Sur, Ilocos Norte, Isabela, Laguna, NCR, Nueva Vizcaya, Pampanga, Quezon, Rizal, Sorsogon, MINDANAO: Agusan del Norte, Bukidnon, Davao Oriental, Davao del Sur, Lanao del Sur, South Cotabato, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, ROMBLON, SAMAR, SULU ARCHIPELAGO, TAWI-TAWI. Native.
Bolbitis quoyana (Gaudich.) Ching in C.Chr., Index Filic., Suppl. Tert. (1934) 49; ≡ Acrostichum quoyanum Gaudich., Voy. Uranie (1827) 306. = Bolbitis enormis (Copel.) C.Chr., Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 197; ≡ Campium enorme Copel., PJS 40 (1929) 307. = Stenosemia dimorpha Copel., PJS 84 (1955) 161. Distribution: Australia, Bismarck Arch, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific Ocean, Philippines, Solomon Isls, Sulawesi. LEYTE, LUZON: Apayao, Cagayan, Camarines Sur, Isabela, Quezon, MINDANAO: Agusan del Sur, Bukidnon, South Cotabato, NEGROS, PANAY, SAMAR. Native.
Bolbitis repanda (Blume) Schott, Gen. Fil. ad (1834) t.14; ≡ Acrostichum repandum Blume, Enum. Pl. Jav. (1828) 104. Distribution: Borneo, Lesser Sunda Isls, ?Nepal, Philippines, Sulawesi. CAMIGUIN, LUZON: Quezon. Native.
Bolbitis rhizophylla (Kaulf.) Hennipman, Blumea 18 (1970) 148; ≡ Egenolfia rhizophylla (Kaulf.) Fée, Mém. Foug. 5 (1852) 48; ≡ Gymnogramma rhizophylla Kaulf., Enum. Fil. (1824) 78. = Egenolfia intermedia (J.Sm. ex Fée) Fée, Gen. (1852) 308; ≡ Polybotrya intermedia J.Sm. ex Fée, Mém. Foug. (1845) 76; J.Sm. in Hook., J. Bot. 3 (1841) 401, nom. inval., nom. nud. = Egenolfia fluviatilis Copel., PJS 38 (1929) 152. Distribution: Philippines, Taiwan. BALABAC, LUZON: Abra, Bataan, Batangas, Benguet, Bulacan, Cagayan, Cavite, Ilocos Norte, Isabela, La Union, Laguna, NCR, Pampanga, Pangasinan, Quezon, Rizal, Sorsogon, Zambales, MARINDUQUE, MINDORO, PALAWAN, PANAY. Native.
Bolbitis scalpturata (Fée) Ching, Index Filic., Suppl. 3 (1934) 50; ≡ Heteroneuron scalpturatum Fée, Mém. Foug. (1845) 95. Distribution: China, Java, Lesser Sunda Isls, Myanmar, Philippines, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. LUZON: Bataan, Batangas, Ilocos Norte, Laguna, Rizal, MINDANAO: Zamboanga Sibugay, PALAWAN. Native.
Bolbitis sinuata (C.Presl) Hennipman, Blumea 18 (1970) 148; ≡ Polypodium sinuatum C.Presl, Rel. Haenk. 1 (1825) 21. = Bolbitis diversifolia (Blume) Schott, Gen. ad (1834) t.14; ≡ Acrostichum diversifolium Blume, Enum. Pl. Jav. (1828) 103. = Bolbitis subsimplex (Fée) Ching, Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 50; ≡ Gymnopteris subsimplex Fée, (Acrost.) 2e mém. 83 (1845) t.40. = Bolbitis hydrophylla (Copel.) Ching, Index Filic., Suppl. Tertium pro Annis 1917-1933 (1934) 48; ≡ Leptochilus hydrophyllus Copel., PJS 1 (Suppl. 2) (1906) 146. = Leptochilus stolonifer Christ, Bull. Herb. Boiss. 2, 6 (1906) 1004; Type: from Luzon: Rizal, Angilog. Distribution: Borneo, Java, Malay Peninsula, Moluccas, New Guinea, Nicobar Isls, Philippines, Sulawesi, Sumatra, Thailand. ALABAT, CATANDUANES, CEBU, LEYTE, LUZON: Albay, Apayao, Batangas, Cagayan, Camarines, Camarines Norte, Camarines Sur, Laguna, Pampanga, Quezon, Rizal, Sorsogon, MINDANAO: Zamboanga, Zamboanga Sibugay, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY, SAMAR. Native.
Bolbitis × sinuosa (Fée) Copel., FFP 2 (1960) 262; ≡ Heteroneuron sinuosum Fée, Mém. Foug., 2. Hist. Acrostich. (1845) 95; ≡ Cyrtogonium sinuosum J.Sm., J. Bot. 8 (1841) 403, nom. inval., nom. nud.; Type: Cuming 105 (W, lecto; B, BM, K, MICH, US, isolecto), Luzon: Laguna and Tayabas, Mt Cristobal. Distribution: Endemic to the Philippines. LUZON: Laguna, Quezon, MINDANAO: Agusan del Sur, POLILLO. Native.
- var. foxii (Copel.) Hennipman, Monogr. Fern Gen. Bolbitis (Leiden Bot. Ser. 2) (1977) 299; ≡ Bolbitis × foxii Copel., PJS, Bot. 81 (1952) 22; Type: Fox 328 (PNH 8904) (PNH, iso), Polillo, Bordios. Distribution: Endemic to the Philippines. LUZON: Quezon, MINDANAO: Agusan del Sur, POLILLO. Native.
- var. sinuosa. Distribution: Endemic to the Philippines. LUZON: Laguna, Quezon. Native.

Excluded

Bolbitis feeiana (Copel.) Fraser-Jenk. & Gandhi, Indian Fern J. 35(1-2): 114 (2018). = Bolbitis presliana (Fée) Ching, Ind. Fil., Suppl. 3 (1934) 49. Notes: This taxon from India does not occur in the Philippines nor anywhere in Malesia. Included in Copeland’s treatment in FFP 2 (1960) 260 as Bolbitis presliana.

Cyrtomium C.Presl

Last edited by Julie F. Barcelona, 20 July 2019

Cyrtomium caryotideum (Wall. ex Hook. & Grev.) C.Presl, Tent. Pterid. (1836) 86; ≡ Polystichum falcatum var. caryotideum (Wall. ex Hook. & Grev.) Matsum., Index Pl. Jap. 1 (1904) 342; ≡ Phanerophlebia caryotidea (Wall. ex Hook. & Grev.) Copel., Gen. Fil. (1947) 111; ≡ Aspidium caryotideum Wall. ex Hook. & Grev., Ic. Fil. (1828) t.69. Distribution: Bhutan, China, India, Japan, Korea, Myanmar, Nepal, Pacific Ocean, Pakistan, Philippines, Taiwan, Tibet, Vietnam. Notes: Philippine record based on a sterile specimen by Mearns from Mt Sto. Tomas (Copeland, FFP 2 (1960) 248). Native.
Cyrtomium falcatum (L.f.) C.Presl, Tent. Pterid. (1836) 86; ≡ Polystichum falcatum (L.f.) Diels, Nat. Pflanzenfam. 1 (1899) 194; ≡ Phanerophlebia falcata (L.f.) Copel., Gen. Fil. (1947) 111; ≡ Polypodium falcatum L.f., Suppl. Pl. (1781) 446. Native.
- ssp. falcatum. Distribution: China, Japan, Korea, Philippines, Ryukyu Isls, Taiwan, Vietnam. Notes: Philippine record based on a Loher collection from Mt Data [Copeland, FFP 2 (1960) 248] at the turn of the 1900s. Native.

Possibly present

Cyrtomium fortunei J.Sm., Ferns Brit. For. (1866) 286. = Cyrtomium fortunei forma latipinna Ching, Ic. Fil. Sin. 3 (1935) t.126. Distribution: Bhutan, China, India, Korea, ?Philippines, Taiwan, Thailand, Vietnam.

Dryopteris Adans.

Last edited by Julie F. Barcelona & Pieter B. Pelser, 5 February 2023

Dryopteris alpina Rosenst., Repert. Spec. Nov. Regni Veg. 12 (1913) 173. = Stenolepia tristis (Blume) Alderw., Bull. Dept. Agric. Indes Neerl. 27 (1909) 46 t. 7; ≡ Aspidium triste Blume, Enum. Pl. Javae 2 (1828) 169; Type: from Java. Distribution: Borneo, Java, New Guinea, Philippines, Sulawesi. MINDANAO: Bukidnon, Cotabato. Notes: 2300-2900m. Conservation status: Other Threatened Species (DAO 2017-11 as Stenolepia tristis). Native.
Dryopteris cacaina Tagawa, Acta Phytotax. Geobot. 6 (1937) 155. Distribution: Japan, Philippines, Taiwan. Native.
Dryopteris chrysocoma (Christ) C.Chr., Index Filic. 5 (1905) 257; ≡ Aspidium filix-mas var. chrysocoma Christ, Bull. Herb. Boissier 6 (1898) 966. Distribution: Bhutan, China, India, Myanmar, Nepal, Pakistan, Philippines, Tibet, Vietnam. Conservation status: Endangered (DAO 2017-11). Native.
Dryopteris cochleata (Don) C.Chr., Index Filic. 5 (1905) 258; ≡ Nephrodium cochleatum D.Don, Prodr. Fl. Nepal. (1825) 6. Distribution: Bangladesh, Bhutan, China, India, Java, Lesser Sunda Isls, Malay Peninsula, Myanmar, Nepal, Pakistan, Philippines, Thailand, Vietnam. LUZON: Abra, Benguet, Mountain Province. Native.
Dryopteris formosana (Christ) C.Chr., Index Filic. 5 (1905) 266; Rufila et al., Ruhuna J. Science 13 (2022) 157; ≡ Aspidium formosanum Christ, Bull. Herb. Boissier, sér. 2, 4 (1904) 615. Distribution: China, Japan, Korea, Philippines, Ryukyu Isls, Taiwan. LUZON: Ilocos Norte, MINDANAO: Davao del Sur. Native.
Dryopteris hasseltii (Blume) C.Chr., Index Filic. 5 (1905) 269; ≡ Polypodium hasseltii Blume, Fl. Jav. Fil. (1829) 195. Distribution: Australia, Borneo, China, India, Java, ?Lesser Sunda Isls, Malay Peninsula, Moluccas, New Guinea, Pacific Ocean, Philippines, Ryukyu Isls, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. MINDANAO: Bukidnon, Davao del Sur. Native.
Dryopteris hendersonii (Bedd.) C.Chr., Index Filic. 5 (1905) 270; Chen et al., Phytotaxa 443 (2020) 132; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22045; ≡ Nothoperanema hendersonii (Bedd.) Ching, Acta Phytotax. Sin. 11 (1966) 28; ≡ Lastrea hendersonii Bedd., Suppl. Ferns S. Ind. (1876) 17; Type: from India. = Ctenitis mearnsii Copel., PJS 81 (1952) 24; Type: Mearns BS 4216 (MICH, lecto), Luzon: Benguet, Haight’s in the Oaks, 1907. Distribution: Bhutan, Borneo, China, India, Japan, Myanmar, Nepal, Philippines, Taiwan, Thailand, Vietnam. LUZON: Benguet, Ifugao, Ilocos Norte, Nueva Vizcaya, MINDANAO: Bukidnon, South Cotabato. Notes: Terrestrial in mossy forest, 2100-2750m. Native.
Dryopteris hirtipes (Blume) Kuntze, Revis. Gen. Pl. 2 (1891) 813; ≡ Aspidium hirtipes Blume, Enum. Pl. Javae 2 (1828) 148. Distribution: Borneo, India, Java, Lesser Sunda Isls, Malay Peninsula, Myanmar, Pacific Ocean, Philippines, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam. LUZON: Benguet, Mountain Province, MINDANAO: Davao del Sur, Sarangani. Native.
Dryopteris manipurensis (Bedd.) C.Chr., Index Filic. (1905) 276; ≡ Dryopsis manipurensis (Bedd.) Holttum & P.J.Edwards, Kew Bull. 41 (1986) 200; ≡ Ctenitis manipurensis (Bedd.) Ching, Bull. Fan Mem. Inst. Biol. Bot. 8 (1938) 297; ≡ Polypodium manipurense Bedd., J. Bot. (1888) 235. = Ctenitis copelandii (Christ) Copel., Gen. Fil. (1947) 124; ≡ Dryopteris copelandii Christ, PJS 2 c (1907) 216. Distribution: India, Nepal, New Guinea, Philippines. LUZON: Benguet, Ilocos Sur, Mountain Province. Native.
Dryopteris nodosa (C.Presl) Li Bing Zhang, Taxon 61 (2012) 1208; ≡ Acrophorus nodosus C.Presl, Tent. Pterid. (1836) 94. = Acrophorus stipellatus T.Moore, Gard. Chron. (1854) 135. Distribution: Bismarck Arch, Borneo, Japan, Lesser Sunda Isls, Myanmar, Pacific Ocean, Philippines, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam. CAMIGUIN, LUZON: Benguet, Ifugao, Kalinga, Mountain Province, Nueva Vizcaya, Quezon, MINDANAO: Bukidnon, Cotabato, Davao del Sur, MINDORO, NEGROS, PANAY. Native.
Dryopteris peranema Li Bing Zhang, Taxon 61 (2012) 1211. = Dryopteris zhuweimingii Li Bing Zhang, Taxon 61(6) (2012) 1211; ≡ Peranema cyatheoides var. luzonicum (Copel.) Ching & S.H.Wu, Acta Phytotax. Sin. 21 (1983) 373; ≡ Peranema luzonicum Copel., PJS 4 (1909) 111, as 'luzonica'; Type: Curran, Merritt & Zschokke 16280 (US, lecto), Luzon: Benguet: Mt Pulag, Jan-1909. Distribution: Bhutan, China, India, Myanmar, Nepal, Philippines, Taiwan, Tibet, Vietnam. LUZON: Benguet, Ifugao, Mountain Province, Nueva Vizcaya. Conservation status: Critically Endangered (DAO 2017-11, as Dryopteris zhuweimingii). Native.
Dryopteris permagna M.G.Price, Gard. Bull. Singapore 30 (1977) 249. Distribution: Endemic to the Philippines. LUZON: Benguet, Kalinga, Mountain Province, MINDANAO: South Cotabato. Conservation status: Endangered (DAO 2017-11). Native.
Dryopteris polita Rosenst., Repert. Spec. Nov. Regni Veg. 13 (1914) 218. Distribution: Borneo, Cambodia, China, Japan, Malay Peninsula, New Guinea, Philippines, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. MINDANAO: Bukidnon. Conservation status: Vulnerable (DAO 2017-11). Native.
Dryopteris pseudocaenopteris (Kunze) Li Bing Zhang, Taxon 61 (2012) 1209; Rufila et al., Ruhuna J. Science 13 (2022) 157; ≡ Diacalpe pseudocaenopteris Kunze, Bot. Zeitung (Berlin) 4 (1846) 457. = Peranema aspidioides (Blume) Mett., Fil. Lechl. 2 (1859) 33; ≡ Diacalpe aspidioides Blume, Enum. Pl. Javae 2 (1828) 241. Distribution: Bhutan, Borneo, China, India, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Myanmar, Nepal, New Guinea, Philippines, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam. LUZON: Benguet, Ifugao, Kalinga, Laguna, Mountain Province, Quezon, Zambales, MINDANAO: Bukidnon, Davao del Sur, Sarangani, NEGROS. Native.
Dryopteris pulvinulifera (Bedd.) Kuntze, Revis. Gen. Pl. 2 (1891) 213; ≡ Lastrea pulvinulifera Bedd., Ferns Brit. India (1870) t.333. Distribution: Bhutan, China, India, Nepal, Philippines, Sri Lanka. LUZON: Abra, Benguet, Ilocos Norte. Native.
Dryopteris purpurascens (Blume) Christ, PJS 2 c (1907) 213; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22045; ≡ Aspidium purpurascens Blume, Enum. Fil. (1828) 169. Distribution: Borneo, Philippines, Sulawesi. LUZON: Benguet, Laguna, MINDANAO: Cotabato, Davao, Davao del Sur, South Cotabato. Native.
Dryopteris sparsa (D.Don) Kuntze, Revis. Gen. Pl. 2 (1891) 813; ≡ Nephrodium sparsum D.Don, Prodr. Fl. Nepal. (1825) 6. Distribution: CAMIGUIN, LUZON: Batangas, Benguet, Ifugao, Kalinga, Laguna, Mountain Province, Pampanga, Quezon, MINDANAO: Bukidnon, Cotabato, Davao del Sur, Sarangani, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN. Native.
- ssp. rectipinnula Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes (2008) 320. Distribution: Australia, Bhutan, Borneo, ?Cambodia, China, India, Japan, Java, ?Laos, Lesser Sunda Isls, Malay Peninsula, Myanmar, Nepal, New Guinea, Pacific Ocean, Philippines, ?Solomon Isls, Sulawesi, Sumatra, Thailand, Tibet, Vietnam. MINDANAO: Bukidnon, PALAWAN. Native.
Dryopteris subarborea (Baker) C.Chr., Index Filic. (1905) 295; ≡ Polypodium subarboreum Baker, J. Linn. Soc. 24 (1887) 259. Distribution: Borneo, Malay Peninsula, New Guinea, Pacific Ocean, Philippines, Sulawesi, Sumatra. CATANDUANES, LUZON: Batangas, Benguet, Laguna, Mountain Province, Quezon, MINDANAO: Cotabato, Davao, Davao del Sur, PANAY, SAMAR. Native.
Dryopteris subtriangularis (C.Hope) C.Chr., Index Filic. 5 (1905) 296; ≡ Nephrodium subtriangulare C.Hope, J. Bot. 28 (1890) 327. = Dryopteris uropinna M.G.Price, Gard. Bull. Singapore 30 (1977) 244. = Dryopteris erythrosora auct. non (D.C.Eaton) Kuntze; Copel. (1960). Distribution: China, India, Myanmar, Philippines, Ryukyu Isls, Taiwan, Thailand, Tibet, Vietnam. LUZON: Abra, Ilocos Norte. Native.
Dryopteris varia (L.) Kuntze, Revis. Gen. Pl. 2 (1891) 814; ≡ Polypodium varium L., Sp. Pl. 2 (1753) 1090. Distribution: China, India, Japan, Korea, Myanmar, Pacific Ocean, Philippines, Ryukyu Isls, Sulawesi, Taiwan, Vietnam. LUZON: Benguet, Ifugao, Ilocos Norte, Kalinga, Pampanga, Quezon. Native.
Dryopteris wallichiana (Spreng.) Hyl., Bot. Not. (1953) 352; ≡ Aspidium wallichianum Spreng., Syst. Veg. 4 (1827) 104. Distribution: LUZON: Benguet, Ifugao, Nueva Vizcaya. Native.
- ssp. wallichiana. = Dryopteris paleacea (Lag. ex Sw.) C.Chr., Amer. Fern J. 1 (1911) 94, nom. illeg., non Hand.-Mazz. Distribution: Africa, America, Bhutan, Borneo, China, India, Japan, Java, Middle East, Myanmar, Nepal, New Guinea, Pacific Ocean, Philippines, Sri Lanka, Sulawesi, Taiwan, Vietnam. Native.

Elaphoglossum Schott ex J.Sm.

Last edited by Julie F. Barcelona, 8 September 2020

Elaphoglossum angulatum (Blume) T.Moore, Index Fil. 5 (1857) 351; ≡ Acrostichum angulatum Blume, Enum. Pl. Javae 2 (1828) 101. Distribution: Africa, Borneo, China, India, Indian Ocean, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific Ocean, Philippines, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. Native.
Elaphoglossum apoense Holttum, Blumea 14 (1966) 320. Distribution: New Guinea, Philippines, Sulawesi. MINDANAO: Cotabato, Davao del Sur. Native.
Elaphoglossum blumeanum (Fée) J.Sm., Ferns Brit. For. (1866) 106. = Elaphoglossum copelandii Christ, PJS 2 (1907) 176. = Elaphoglossum blumeanum var. philippinense Christ ex Holttum, Flora Malesiana Series II, 1 (1978) 311. = ?Elaphoglossum petiolatum auct. non (Sw.) Urb.; Coritico et al., PJS 149 (2020) 778. Distribution: Bismarck Arch, Borneo, Java, Malay Peninsula, Moluccas, Philippines, Solomon Isls, Sulawesi, Sumatra. CAMIGUIN, LEYTE, LUZON: Albay, Batangas, Benguet, Ilocos Norte, Kalinga, Laguna, Mountain Province, Pampanga, Quezon, Rizal, Sorsogon, MINDANAO: Bukidnon, Cotabato, Davao, Davao del Sur, Lanao del Sur, Sarangani, NEGROS. Native.
Elaphoglossum calanasanicum Holttum, Kalikasan 3 (1974) 196. Distribution: Endemic to the Philippines. LUZON: Apayao, Benguet. Notes: Only known from the type. Native.
Elaphoglossum callifolium (Blume) T.Moore, Index Fil. (1857) 7; ≡ Acrostichum callifolium Blume, Enum. Pl. Javae 2 (1828) 100. Distribution: Australia, Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific Ocean, Philippines, Sulawesi, Sumatra, Taiwan, ?Vietnam. LUZON: Apayao, Benguet, Camarines Sur, Ilocos Norte, Kalinga, Quezon, MINDANAO: Bukidnon, Cotabato, Davao Oriental, Davao del Sur, NEGROS. Native.
Elaphoglossum luzonicum Copel., LPB 1 (1907) 235. = Elaphoglossum elmeri Copel., LPB 3 (1910) 849. Distribution: Borneo, Malay Peninsula, New Guinea, Philippines, Taiwan. LUZON: Bataan, Batangas, Benguet, Camarines Sur, Ilocos Norte, Kalinga, Laguna, Mountain Province, Pampanga, Quezon, Zambales, MINDANAO: Davao del Sur, NEGROS, PANAY, SAMAR. Native.
Elaphoglossum melanostictum (Blume) T.Moore, Index Fil. (1862) 361; ≡ Acrostichum melanostictum Blume, Fl. Javae (1829) 26. = Elaphoglossum basilanicum Copel., PJS 11 (1916) 41. Distribution: Borneo, Java, Malay Peninsula, Philippines, Sumatra, Thailand. BASILAN, MINDANAO: Agusan del Sur, NEGROS, SAMAR. Native.
Elaphoglossum negrosensis Holttum, Blumea 14 (1966) 323. = Elaphoglossum angulatum auct. non (Blume) T.Moore; Copel. (1960), pro parte. Distribution: Endemic to the Philippines. NEGROS. Notes: Only known from the type. Native.
Elaphoglossum ophioglossoides (Goldm.) Holttum, Kalikasan 3 (1974) 197; ≡ Acrostichum ophioglossoides Goldm., Nov. Actorum Acad. Caes. Leop.-Carol. German. Nat. Cur. 19 Suppl. 1 (1843) 451. = Elaphoglossum merrillii Christ, PJS 3 (1908) 275. Distribution: Endemic to the Philippines. LUZON: Apayao, Batangas, Camarines Sur, Ilocos Norte, NCR, Quezon, MINDORO, NEGROS, SAMAR. Native.

Lastreopsis Ching

Last edited by Pieter B. Pelser & Julie F. Barcelona, 11 September 2022

Lastreopsis tenera (R.Br.) Tindale, Victoria Naturalist 73 (1957) 181; ≡ Nephrodium tenerum R.Br., Prodr. (1810) 149. = Ctenitis recedens (J.Sm. ex T.Moore) Copel., Gen. Fil. (1947) 124; ≡ Polypodium recedens J.Sm. in Hook., J. Bot. 3 (1841) 394, nom. inval., nom. nud; ≡ Lastrea recedens J.Sm. ex T.Moore, Gard. Chr. (1855) 708. Distribution: Australia, India, Lesser Sunda Isls, Malay Peninsula, New Caledonia, Pacific Ocean, Philippines, Sri Lanka, Sumatra, Taiwan. LEYTE, LUZON: Batangas, Ilocos Norte, Laguna, Nueva Vizcaya, Quezon, Rizal. Native.

Lomagramma J.Sm.

Last edited by Julie F. Barcelona, 7 September 2020

Lomagramma brooksii Copel., PJS 3 (1908) 345. Distribution: Borneo, Moluccas, Pacific Ocean, Philippines, Sulawesi. BATAN, LUZON: Quezon, MINDORO. Native.
Lomagramma copelandii Holttum, Gard. Bull. Straits Settlem. 9 (1937) 201. Distribution: Moluccas, Philippines. BOHOL, CATANDUANES, LUZON: Laguna, Quezon, Sorsogon, MINDANAO: Agusan del Sur, Surigao, NEGROS, SAMAR. Native.
Lomagramma merrillii Holttum, Gard. Bull. Straits Settlem. 9 (1937) 208. Distribution: Philippines, Sulawesi. MINDANAO: Zamboanga del Sur, NEGROS. Native.
Lomagramma pteroides J.Sm., J. Bot. (Hooker) 3 (1841) 402. = Lomagramma subcoriacea Copel., PJS 40 (1929) 308. = Lomagramma pedicellata Copel., PJS 81 (1952) 23. = Lomagramma cordata Copel., PJS 84 (1955) 162. Distribution: Moluccas, Philippines. BILIRAN, BOHOL, CAMIGUIN, CEBU, JOLO, LUZON: Batangas, Laguna, Quezon, Sorsogon, MINDANAO: Bukidnon, South Cotabato, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY. Native.

Polystichum Roth

Last edited by Julie F. Barcelona, 12 April 2022

Polystichum biaristatum (Blume) T.Moore, Index Fil. (1858) 86; ≡ Aspidium biaristatum Blume, Enum. Pl. Javae 2 (1828) 164. = Polystichum aquilinum Copel., PJS 81 (1952) 20. Distribution: Bangladesh, Java, Lesser Sunda Isls, Malay Peninsula, Myanmar, Philippines, Sri Lanka, Sulawesi, Taiwan, Thailand, Vietnam. LUZON: Benguet, Ilocos Norte, Kalinga, Mountain Province, Pampanga, Quezon, Rizal, Zambales. Native.
Polystichum copelandii Christ, PJS 2 c (1907) 157. Distribution: Endemic to the Philippines. LUZON: Benguet, Mountain Province. Native.
Polystichum deltodon (Baker) Diels, Nat. Pflanzenfam. 1 (1899) 191; ≡ Aspidium deltodon Baker, Gard. Chr. n. s. 14 (1810) 494. Distribution: China, Japan, Myanmar, Philippines, Ryukyu Isls, Taiwan, Vietnam. LUZON: Bataan, Benguet, PANAY (photos). Native.
Polystichum elmeri Copel., PJS 81 (1952) 21; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22045. Distribution: Endemic to the Philippines. LUZON: Benguet, MINDANAO: Bukidnon, Davao, Davao del Sur, South Cotabato. Native.
Polystichum fuscum Copel., PJS 81 (1952) 20. Distribution: Endemic to the Philippines. LUZON: Abra, MINDANAO: Bukidnon. Native.
Polystichum moluccense T.Moore, Index Fil. (1858) 84; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22045. = Polystichum horizontale C.Presl, Epim. Bot. (1849) 57. = Polystichum nudum Copel., PJS 1 (Suppl. 2) (1906) 145. Distribution: Bismarck Arch, Borneo, Moluccas, New Guinea, ?Pacific Ocean, Philippines, Solomon Isls, Sulawesi. CAMIGUIN, LEYTE, LUZON: Batangas, Benguet, Camarines Norte, Camarines Sur, Ifugao, Kalinga, Laguna, Mountain Province, Quezon, Sorsogon, MINDANAO: Bukidnon, Davao del Sur, South Cotabato, Zamboanga del Sur, NEGROS, PANAY. Conservation status: Endangered (DAO 2017-11, as Polystichum nudum Copel.). Native.
Polystichum nepalense (Spreng.) C.Chr., Index Filic. 10 (1905) 84; ≡ Aspidium nepalense Spreng., Syst. 4 (1827) 97. Distribution: Bhutan, China, India, Myanmar, Nepal, Philippines, Taiwan, Tibet, Vietnam. LUZON: Benguet, Ifugao, Nueva Vizcaya. Native.
Polystichum obliquum (D.Don) T.Moore, Index Fil. 87 (1858) 98; ≡ Aspidium obliquum D.Don, Prodr. Fl. Nepal. (1825) 3. Distribution: Bhutan, China, India, Myanmar, Nepal, Philippines, Taiwan. LUZON: Benguet. Native.
Polystichum obtusum J.Sm. ex C.Presl, Epim. Bot. (1849) 53; J.Sm., J. Bot. 3 (1841) 412, nom. inval., nom. nud. = Polystichum edanoi Copel., PJS 83 (1954) 97. Distribution: Borneo, Philippines. LUZON: Bataan, Batangas, Benguet, Nueva Ecija, Rizal. Native.
Polystichum piceopaleaceum Tagawa, Acta Phytotax. Geobot. 5 (1936) 255. = Polystichum doianum Tagawa, Acta Phytotax. Geobot. 5 (1936) 256. = Polystichum pulogense Copel., 81 PJS (1952) 19. Distribution: Bhutan, China, India, Japan, Myanmar, Nepal, Pakistan, Philippines, Sri Lanka, Taiwan, Tibet. LUZON: Benguet, Ifugao, Nueva Vizcaya. Native.

Possibly present

Polystichum grandifrons C.Chr., Index Filic., Suppl. 3 (1934) 163. Distribution: China, India, Japan, ?Philippines, Taiwan, Vietnam.
Polystichum manmeiense (Christ) Nakaike, Misc. Publ. Nat. Sci. Mus. Tokyo (1982) 141; ≡ Aspidium manmeiense Christ, Bull. Herb. Boissier 6 (1898) 965. Distribution: Bhutan, China, India, Myanmar, Nepal, ?Philippines, Taiwan, Tibet.

Teratophyllum Mett.

Last edited by Julie F. Barcelona, 7 September 2020

Teratophyllum aculeatum (Blume) Mett. ex Kuhn, Ann. Mus. Bot. Lugduno-Batavi 4 (1869) 296; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22045; ≡ Lomaria aculeata Blume, Enum. Pl. Javae (1828) 205. Distribution: BOHOL, GENERAL ISLAND, LUZON: Bataan, Benguet, Kalinga, Laguna, Quezon, Rizal, MINDANAO: Bukidnon, South Cotabato, Zamboanga, MINDORO, PALAWAN, SULU ARCHIPELAGO, TAWI-TAWI. Native.
- var. aculeatum. Distribution: Borneo, Java, Moluccas, Myanmar, New Guinea, Philippines, Singapore, Sri Lanka, Sulawesi, Sumatra, Thailand. MINDANAO: Bukidnon. Native.
Teratophyllum arthropteroides (Christ) Holttum, Gard. Bull. Straits Settlem. 5 (1932) 303; ≡ Stenochlaena arthropteroides Christ, Bull. Herb. Boissier, sér. 2, 6 (1906) 998. Distribution: Endemic to the Philippines. LUZON: Laguna. Native.
Teratophyllum koordersii Holttum, Gard. Bull. Straits Settlem. 5 (1932) 301. Distribution: Philippines, Sulawesi. LEYTE, LUZON: Camarines Norte, Laguna, Quezon, MINDANAO: Surigao del Norte, SAMAR. Native.
Teratophyllum leptocarpum (Fée) Holttum, Gard. Bull. Straits Settlem. 9 (1937) 143; ≡ Lomariopsis leptocarpa Fée, Mém. Foug. (1845) 69. Distribution: Pacific Ocean, Philippines, Sulawesi. BASILAN, LUZON: Bataan, Ilocos Norte, Pampanga, Quezon, PANAY, SAMAR, SIBUYAN. Native.
Teratophyllum luzonicum Holttum, Gard. Bull. Straits Settlem. 5 (1932) 297. Distribution: Endemic to the Philippines. LUZON: Apayao, Aurora, Bataan, Pampanga, MINDANAO: Agusan del Norte, NEGROS, PALAWAN. Native.

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