Dennstaedtiaceae

Pteridiaceae, Hypolepidaceae, Monachosoraceae

Dennstaedtia Bernh.

Last edited by Julie F. Barcelona & Pieter B. Pelser, 12 September 2023

Dennstaedtia articulata Copel., LPB 2 (1908) 396. Distribution: Endemic to the Philippines. LUZON: Aurora, NEGROS. Conservation status: Endangered (DAO 2017-11). Native.
Dennstaedtia dennstaedtioides (Copel.) Copel., PJS 2 c (1907) 126; ≡ Microlepia dennstaedtioides Copel., PJS 1 (Suppl. 2) (1906) 148; Type: Copeland 1475, Mindanao: Davao, monte Apo. Distribution: Philippines, Sulawesi. LUZON: Benguet, Laguna, Quezon, MINDANAO: Cotabato, Davao, Davao del Sur, Sarangani. Native.
Dennstaedtia elmeri Copel., LPB 1 (1907) 233. Distribution: Endemic to the Philippines. LUZON: Batangas, Benguet, Laguna, Quezon, MINDANAO: Bukidnon, PANAY. Native.
Dennstaedtia fusca Copel., PJS 81 (1952) 4. Distribution: Endemic to the Philippines. PANAY. Conservation status: Endangered (DAO 2017-11). Native.
Dennstaedtia glabrata (Ces.) C.Chr., Index Filic. (1905) 217; ≡ Dicksonia glabrata Ces., Rend. Ac. Napoli 16 (1877) 24. = Dennstaedtia erythrorachis (Christ) Diels in Engler & Prantl, Nat. Pflanzenfam. 1 (1899) 218. Distribution: ?Bismarck Arch, New Guinea, Pacific Ocean, Philippines, Solomon Isls, Sulawesi. BASILAN, LUZON: Quezon, MINDANAO: Zamboanga, Zamboanga del Sur, NEGROS. Native.
Dennstaedtia hirsuta (Sw.) Mett. ex Miq., Ann. Mus. Bot. Lugduno-Batavi 3 (1867) 181; ≡ Davallia hirsuta Sw., J. Bot. (Schrader) 1800 (1801) 87. Distribution: China, E Asia, Japan, Korea, Philippines, Taiwan. LUZON: Ilocos Norte. Notes: J.F. Barcelona (12-Sep-23): Known from the Philippines from Iwatsuki, Murata, & Gutierrez P514 (L), Iwatsuki, Murata, & Gutierrez P538 (L, PNH), Iwatsuki, Murata, & Gutierrez P585 (L), Price 3360 (L, PNH). Native.
Dennstaedtia hooveri Christ, PJS 2 c (1907) 169. Distribution: Moluccas, Philippines. LEYTE, LUZON: Quezon, MINDANAO: Agusan del Norte, Bukidnon, Davao Oriental, Davao del Sur, South Cotabato, Zamboanga del Sur. Native.
Dennstaedtia macgregorii Copel., PJS 81 (1952) 4. Distribution: Endemic to the Philippines. LUZON: Ifugao, Ilocos Norte, Kalinga. Native.
Dennstaedtia merrillii Copel., PJS 2 c (1907) 126. Distribution: Endemic to the Philippines. LUZON: Albay, Benguet, Ilocos Norte, Kalinga, Quezon, MINDANAO: Davao, Davao del Sur, MINDORO, NEGROS. Native.
Dennstaedtia philippinensis Copel., LPB 9 (1920) 3107. Distribution: Endemic to the Philippines. BASILAN, LUZON: Batangas, Laguna, Quezon, Sorsogon, MINDANAO: Davao del Sur, SULU ARCHIPELAGO. Native.
Dennstaedtia resinifera (Blume) Mett. ex Kuhn, Ann. Mus. Bot. Lugduno-Batavi 4 (1869) 290; ≡ Cheilanthes resinifera Blume, Enum. Fil. (1828) 138. = Dennstaedtia cuneata (J.Sm.) T.Moore, Index Fil. (1857) 97; ≡ Sitolobium cuneatum J.Sm., J. Bot. 3 (1841) 418. Distribution: Borneo, Java, Moluccas, Philippines. ALABAT, BASILAN, CATANDUANES, LEYTE, LUZON: Apayao, Bataan, Batangas, Benguet, Cagayan, Laguna, Pampanga, Quezon, Sorsogon, MINDANAO: Davao del Sur, Zamboanga del Sur, MINDORO. Native.
Dennstaedtia rufidula C.Chr. in C.Chr. & Holltum, Gard. Bull. Sing. 7 (1934) 226. Distribution: Borneo, Philippines. MINDANAO: Davao Oriental. Native.
Dennstaedtia scandens (Blume) T.Moore, Index Fil. (1861) 307; ≡ Dicksonia scandens Blume, Enum. Pl. Javae (1828) 240. Distribution: Lesser Sunda Isls, Moluccas, New Guinea, Pacific Ocean, Philippines, Solomon Isls, Sulawesi, Taiwan. LEYTE, LUZON: Benguet, Mountain Province, Nueva Vizcaya, Quezon, MINDANAO: Agusan del Norte, Bukidnon, Davao, Davao del Sur, Lanao del Sur, MINDORO. Native.
Dennstaedtia smithii (Hook.) T.Moore, Index Fil. (1861) 308; ≡ Dicksonia smithii Hook., Sp. Fil. 1 (1844) 80. Distribution: Philippines, Sulawesi, Taiwan. CATANDUANES, LUZON: Apayao, Bataan, Batangas, Benguet, Cagayan, Laguna, Mountain Province, NCR, Nueva Ecija, Nueva Vizcaya, Pampanga, Quezon, Rizal, Sorsogon, Zambales, MINDANAO: Bukidnon, Davao, Davao del Sur, Lanao del Sur, Zamboanga del Sur, NEGROS, PANAY. Native.
Dennstaedtia williamsii Copel., PJS 1 (Suppl. 2) (1906) 148. Distribution: Endemic to the Philippines. LUZON: Sorsogon, MINDANAO: Bukidnon, Davao del Sur (photos), Zamboanga del Sur. Native.
Dennstaedtia zeylanica (Sw.) Zink ex Fraser-Jenk. & Kandel, Ferns and Fern-allies of Nepal 1 (2015) 161; ≡ Dicksonia zeylanica Sw., J. Bot. (Schrader) 1800 (1801) 91. = Dennstaedtia scabra (Wall. ex Hook.) T.Moore, Index Fil. (1861) 307; ≡ Dicksonia scabra Wall. ex Hook., Sp. Fil. 1 (1844) 80. Distribution: Bhutan, Borneo, China, India, Japan, Korea, Laos, Malay Peninsula, Myanmar, Nepal, Philippines, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam. LUZON: Abra, Benguet, Ifugao, Ilocos Norte, Kalinga-Apayao, Mountain Province, Nueva Vizcaya, Quezon, MINDANAO: Cotabato, Davao del Sur, Sarangani, MINDORO. Native.

Histiopteris (Agardh) J.Sm.

Last edited by Julie F. Barcelona, 7 September 2020

Histiopteris incisa (Thunb.) J.Sm., Hist. Fil. (1875) 295; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22044. = Histiopteris montana Copel., Fragm. Fl. Philipp. (1905) 193. Distribution: Africa, America, Australia, Bhutan, Borneo, Cambodia, China, India, Indian Ocean, Japan, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, Nepal, New Caledonia, Philippines, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Taiwan, Thailand, Tibet, Vietnam. BILIRAN, LEYTE, LUZON: Bataan, Benguet, Camarines Norte, Camarines Sur, Ifugao, Ilocos Norte, Kalinga, Laguna, Mountain Province, Nueva Ecija, Nueva Vizcaya, Quezon, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao, Davao del Norte, Davao del Sur, Lanao del Sur, Sarangani, South Cotabato, Surigao, MINDORO, NEGROS, PANAY. Native.

Hypolepis Bernh.

Last edited by Julie F. Barcelona, 7 September 2020

Hypolepis alpina (Blume) Hook., Sp. Fil. 2 (1852) 63; ≡ Cheilanthes alpina Blume, Enum. Pl. Javae 2 (1828) 138. Distribution: Borneo, Java, New Guinea, Pacific Ocean, Philippines, Ryukyu Isls, Taiwan. LUZON, MINDANAO: Davao del Sur. Native.
Hypolepis bamleriana Rosenst., Fedde Repert. 10 (1912) 325. Distribution: Borneo, Moluccas, New Guinea, Pacific Ocean, Philippines, Sulawesi. LUZON: Ilocos Norte, MINDANAO: Bukidnon, Davao del Sur, NEGROS. Native.
Hypolepis malesiana Brownsey, Blumea 32 (1987) 248. Distribution: New Guinea, Philippines, ?Sumatra. LUZON: Benguet, Mountain Province, Zambales, NEGROS. Native.
Hypolepis polypodioides (Blume) Hook., Sp. Fil. 2 (1852) 63; ≡ Cheilanthes polypodioides Blume, Enum. Pl. Javae 2 (1828) 139. Distribution: Bangladesh, Bhutan, Cambodia, China, India, Laos, Malay Peninsula, Myanmar, Nepal, Philippines, Taiwan, Tibet, Vietnam. LUZON: Benguet, Ifugao, Mountain Province, Nueva Vizcaya, MINDANAO: Bukidnon. Native.
Hypolepis resistens (Kunze) Hook., Sp. Fil. 2. (1852) 64; ≡ Cheilanthes resistens Kunze, Linn. 24 (1851) 275. = Hypolepis glandulifera Brownsey & Chinnock, J. Adelaide Bot. Gard. 10 (1987) 16. Distribution: Australia, Bhutan, Borneo, China, India, Malay Peninsula, Myanmar, New Caledonia, New Guinea, Philippines, Sri Lanka, Vietnam. LUZON: Benguet, MINDANAO: Davao del Norte, Davao del Sur. Native.
Hypolepis rugosula (Labill.) J.Sm., Companion Bot. Mag. 72 (1846) 8; ≡ Polypodium rugosulum Labill., Nov. Holl. Pl. 2 (1806) 92. Distribution: LUZON: Albay, MINDORO. Native.
- ssp. rudis (Kunze) Schwartsb., Acta Bot. Brasil. 28 (2014) 216; ≡ Cheilanthes rudis Kunze, Bot. Zeit. (Berlin) (1848) 213; Type: Cuming 140 (FI, K, RB), Luzon. Distribution: Endemic to the Philippines. LUZON. Native.
Hypolepis tenuifolia (G.Forst.) Bernh., Neues J. Bot. 1 (1805) 34; ≡ Lonchitis tenuifolia G.Forst., Fl. Ins. Austr. (1786) 80. Distribution: Australia, China, Lesser Sunda Isls, New Caledonia, New Guinea, Pacific Ocean, Philippines, Sulawesi, Taiwan, Vietnam. LUZON: Bataan, Batangas, Benguet, Ifugao, Kalinga, Laguna, Mountain Province, Nueva Ecija, Nueva Vizcaya, Quezon, Sorsogon, Zambales, MINDANAO: Bukidnon, Davao, Davao del Sur, Lanao del Sur, NEGROS. Native.

Possibly present

Hypolepis pallida (Blume) Hook., Sp. Fil. 2 (1852) 64; ≡ Cheilanthes pallida Blume, Enum. Fil. (1828) 139. Distribution: ?Borneo, Malay Peninsula, Pacific Ocean, ?Philippines, Sumatra, Taiwan, ?Vietnam. ?NEGROS. Notes: Doubtfully occurring in the Philippines (Negros: Amlan, M.G.Price 2479).
Hypolepis punctata (Thunb. ex Murray) Mett. ex Kuhn, Fil. Afr. (1868) 120; ≡ Polypodium punctatum Thunb., Fl. Jap. (1784) 337. Distribution: China, Korea, ?Philippines, Ryukyu Isls, Taiwan. LUZON: Albay, Benguet, Ilocos Norte, Quezon, MINDANAO. Notes: Doubtfully occurring in the Philippines.

Microlepia C.Presl

Last edited by Julie F. Barcelona & Pieter B. Pelser, 18 September 2023

Microlepia matthewii Christ, Not. syst. 1 (1909) 54. = Microlepia herbacea Ching & C.Chr. ex Tardieu & C.Chr., Notul. Syst. (Paris) 6 (1937) 6. Distribution: China, Laos, Philippines, Thailand, Vietnam. LUZON: Benguet, Ilocos, Ilocos Norte, Kalinga, Mountain Province, MINDANAO: Davao del Sur, Sarangani. Native.
Microlepia pilosiuscula (Sm.) C.V.Morton, Contr. U.S. Natl. Herb. 38 (1974) 313; ≡ Davallia pilosiuscula Sm. in A.Rees, Cycl. 11 (1808) n. 10. = Microlepia trapeziformis (Roxb.) Kuhn, Chaetopterides (1882) 27. Distribution: Java, Malay Peninsula, Moluccas, Philippines, Sulawesi, Sumatra. LUZON: Benguet, Mountain Province. Notes: J.F. Barcelona (18-Sep-23): Known from the Philippines from Price 414 (PNH) and Hernaez s.n. (PNH160901). Native.
Microlepia platyphylla (D.Don) J.Sm., Lond. J. Bot. 1 (1842) 427b; ≡ Davallia platyphylla D.Don, Prodr. Fl. Nepal. (1825) 10. Distribution: Bhutan, China, India, Laos, Myanmar, Nepal, Philippines, Sri Lanka, Taiwan, Thailand, Tibet, Vietnam. LUZON: Benguet, Mountain Province. Native.
Microlepia protracta Copel., PJS 81 (1952) 5; Rufila et al., Ruhuna J. Science 13 (2022) 157. Distribution: Endemic to the Philippines. BALUT, MINDANAO: Davao del Sur. Conservation status: Endangered (DAO 2017-11). Native.
Microlepia speluncae (L.) T.Moore, Ind. Fil. (1857) 93; ≡ Polypodium speluncae L., Sp. Pl. 2 (1753) 1093. = Microlepia hispidula Alderw., Malayan Ferns Fern Allies (1917) 233. = Microlepia sablanensis Christ, PJS 2 c (1907) 168. = Microlepia trichosticha (Hook.) J.Sm. ex Fée, Mém. Foug., 5. Gen. Filic. (1852) 326; ≡ Davallia trichosticha Hook., Sp. Fil. 1 (1845) 183; ≡ Microlepia trichosticha J.Sm., J. Bot. (Hooker) 3 (1841) 416, nom. inval., nom. nud.; Type: Cuming 328, Samar. = ?Microlepia manilensis C.Presl, Tent. Pterid. (1836) 125. Distribution: Africa, America, Andaman Isls, Australia, Bangladesh, Bhutan, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, ?Japan, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, Nepal, Nicobar Isls, Pacific Ocean, Philippines, Ryukyu Isls, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam. BALABAC, BASILAN, BATAN, BILIRAN, BOHOL, CAMIGUIN, CAMIGUIN DE BABUYANES, CULION, LEYTE, LUBANG, LUZON: Abra, Albay, Apayao, Aurora, Bataan, Batangas, Benguet, Bulacan, Cagayan, Camarines Norte, Camarines Sur, Ifugao, Ilocos Norte, Isabela, Kalinga, La Union, Laguna, Mountain Province, NCR, Nueva Ecija, Pampanga, Pangasinan, Quezon, Rizal, Sorsogon, Zambales, MINDANAO: Bukidnon, Davao, Davao Oriental, Davao del Sur, Lanao, Lanao del Sur, Maguindanao del Norte, Surigao, Zamboanga, Zamboanga Sibugay, Zamboanga del Sur, MINDORO, PALAWAN, PANAY, POLILLO, ROMBLON, SAMAR, SIARGAO, SIBUYAN. Native.
Microlepia strigosa (Thunb.) C.Presl, Abh. Kön. Böhm. Ges. Wiss. 5(6) (Epim. Bot.) (1851) 95; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22045; ≡ Trichomanes strigosum Thunb., Fl. Jap. (1784) 339. Distribution: Bangladesh, Bhutan, Borneo, China, India, Japan, Java, Korea, Lesser Sunda Isls, Malay Peninsula, Myanmar, Nepal, New Caledonia, Pacific Ocean, Philippines, Ryukyu Isls, Sri Lanka, Sulawesi, Thailand, Vietnam. BATAN, LUZON: Bataan, Benguet, Ifugao, Ilocos Norte, Ilocos Sur, Kalinga, Laguna, Mountain Province, Nueva Vizcaya, Pampanga, Quezon, Rizal, Zambales, MINDANAO: Bukidnon, Davao del Sur, Lanao del Sur, South Cotabato, Zamboanga del Sur, SABTANG. Native.
Microlepia todayensis Christ, PJS 3 (1908) 272. Distribution: China, Malay Peninsula, Myanmar, Philippines, Sumatra, Vietnam. CATANDUANES, LUZON: Sorsogon, MINDANAO: Cotabato, Davao, Davao del Sur, Lanao del Sur, South Cotabato, PANAY, POLILLO, SAMAR. Native.

Possibly present

Microlepia firma Mett. ex Kuhn, Linn. 36 (1869) 146. Distribution: Bhutan, ?Cambodia, China, India, Myanmar, Nepal, ?Philippines, Sri Lanka, Thailand, Tibet, Vietnam.
Microlepia rhomboidea (Wall. ex Kunze) Prantl, Arbeiten Königl. Bot. Gart. Breslau 1 (1892) 31; ≡ Davallia rhomboidea Wall. ex Kunze, Bot. Zeit. (Berlin) (1850) 158. Distribution: Bangladesh, Bhutan, China, India, Myanmar, Nepal, ?Philippines, Sri Lanka, Taiwan, Vietnam.

Monachosorum Kunze

Last edited by Julie F. Barcelona, 22 June 2019

Monachosorum henryi Christ, Bull. Herb. Boiss. 6 (1898) 869; Galope-Obemio et al., J. Threatened Taxa 14 (2022) 22045. Distribution: Bhutan, China, India, Myanmar, Nepal, Philippines, Taiwan, Thailand, Tibet, Vietnam. LUZON: Benguet, Kalinga, Mountain Province, MINDANAO: South Cotabato. Native.
Monachosorum subdigitatum (Blume) Kuhn, Chaetopt. (1882) 345; ≡ Aspidium subdigitatum Blume, Enum. Pl. Jav. (1828) 171. Distribution: Bhutan, Borneo, ?India, Java, Malay Peninsula, Moluccas, Myanmar, New Guinea, Philippines, Sulawesi, Sumatra, Vietnam. LUZON: Abra, Benguet, Ifugao, Ilocos Norte, Kalinga, Mountain Province, Nueva Vizcaya, MINDANAO: Cotabato, Davao, Davao del Sur, South Cotabato, MINDORO. Native.

Paesia A.St.-Hil.

Last edited by Julie F. Barcelona, 7 September 2020

Paesia elmeri Copel., LPB 3 (1910) 826. Distribution: Malay Peninsula, Philippines, Sumatra. LUZON: Benguet, Mountain Province, MINDANAO: Bukidnon, Cotabato, Davao, Davao del Sur, MINDORO. Native.
Paesia radula (Baker) C.Chr., Index Filic. (1906) 476; ≡ Pteris radula Baker, J. Bot. (1880) 211. = Paesia luzonica Christ, PJS 3 c (1908) 275. Distribution: Borneo, Java, Moluccas, New Guinea, Philippines, Sulawesi, Sumatra, Taiwan. LUZON: Benguet, Mountain Province, MINDANAO: Bukidnon. Native.

Pteridium Gled. ex Scop.

Last edited by Julie F. Barcelona, 7 September 2020

Pteridium aquilinum (L.) Kuhn, Bot. Ost-Afrika 3 (1879) 11; ≡ Pteris aquilina L., Sp. Pl. 2 (1753) 1075. Distribution: BOHOL, LUZON: Albay, Apayao, Bataan, Batangas, Benguet, Ifugao, Ilocos Norte, Ilocos Sur, Kalinga, Laguna, Mountain Province, NCR, Quezon, Rizal, MINDANAO: Bukidnon, Cotabato, Davao Oriental, Davao del Sur, Lanao del Sur, South Cotabato, Surigao, Zamboanga del Norte, MINDORO, NEGROS, SANTA CRUZ ISLS, TAWI-TAWI. Notes: Widespread. Native.
- var. wightianum (Wall. ex J.Agardh) R.M.Tryon, Rhodora 43 (1941) 22; ≡ Pteris recurvata var. wightiana Wall. ex J.Agardh, Recens. Spec. Pter. (1839) 50. = Pteridium revolutum (Blume) Nakai, Bot. Mag. (Tokyo) 39 (1925) 109; ≡ Pteris revoluta Blume, Enum. Pl. Javae 2 (1828) 214. Distribution: Australia, China, Indochina, Malesia, Philippines. BOHOL, LUZON: Albay, Apayao, Bataan, Benguet, Ifugao, Ilocos Norte, Ilocos Sur, Kalinga, Mountain Province, Quezon, Rizal, MINDANAO: Bukidnon, Cotabato, Lanao del Sur, South Cotabato, MINDORO, NEGROS, TAWI-TAWI. Native.

Possibly present

Pteridium rostratum (Burm.f.) Fraser-Jenk., Indian Fern J. 36 (2019) 28; ≡ Polypodium rostratum Burm.f., Fl. Ind. (1768) 235. = Pteridium aquilinum var. yarrabense Domin, Biblioth. Bot. 20 (1913) 161. Distribution: Australia, Cambodia, Malay Peninsula, Moluccas, Myanmar, ?Philippines, Sumatra, Thailand, Vietnam. BILIRAN, CAMIGUIN, LUZON: Zambales, MINDANAO: Bukidnon. Notes: J.F. Barcelona (23-Sep-23): Perhaps represented in the Philippines by BS Ramos 14782, Santos 6035, Santos 6175 and Sulit 5511.

References

Alston, AHG. 1959. Isoetaceae. Flora Malesiana series 2, 1: 63-64
Amoroso, VB, FP Coritico & PW Fritsch. 2020. Actinostachys minuta, a new species of grass fern from Mindanao, Philippines. PhytoKeys 151: 59-66
Amoroso, VB, YLL Cariño, JC Nobleza & FP Coritico. 2020. Ophioderma subsessile (Ophioglossaceae), a new snake tongue fern species from Mindanao, Philippines. Philippine Journal of Science 150 (S1): 215-221
Amoroso, VB, YLL Cariño, JC Nobleza & FP Coritico. 2021. A new species of Actinostachys (Schizaeaceae) from Mindanao, Philippines. Phytotaxa 508: 101-106.
Barcelona, JF & MG Price. 1999. A new Philippine Goniophlebium. Fern Gazette 15: 261-264.
Barcelona, JF & PB Pelser. 2014. Phanerosorus (Matoniaceae), a new fern genus record for the Philippines. Phytotaxa 170: 133-135.
Barcelona, JF, B. Hernaez, & MG Price. 1996. Philippine Schizaea. Asia Life Sciences 5: 27-34
Barcelona, JF, DL Nickrent, JV LaFrankie, JRC Callado & PB Pelser. 2013. Co’s Digital Flora of the Philippines: plant identification and conservation through cybertaxonomy. Philippine Journal of Science 142: 57-67
Barcelona, JF, NE Dolotina, GS Madroñero, WG Granert & DD Sopot. 2006. The ferns and fern allies of the karst forests of Bohol Island, Philippines. American Fern Journal 96: 1-20
Barcelona, JF. 2002. Philippine pteridophyte collections as a resource for conservation planning. Proceedings of the “International Symposium on the Fern Flora Worldwide: Threats and Responses” sponsored by the British Pteridological Society and IUCN- Species Survival Commission.” Fern Gazette 16(6, 7 & 8): 307-312.
Barcelona, JF. 2003. Preliminary report on the ferns and fern allies (pteridophytes) of Mt. Bali-it, Balbalasang-Balbalan National Park, Kalinga, Northern Luzon, Philippines. Sylvatrop: The Technical Journal of Philippine Ecosystems and Natural Resources 13: 81-92
Barcelona, JF. 2003. The taxonomy and ecology of the pteridophytes of Mt. Iraya & vicinity, Batan Island, Batanes Province, Northern Philippines, in Pteridology in the New Millenium (Chandra and Srivastava, eds). Kluwer Academy Publishers. The Netherlands, 299-325.
Barcelona, JF. 2004. Collection and conservation status of pteridophytes in Panay Island, Philippines. Philipp. Scient. 41:57-73.
Barcelona, JF. 2005. Noteworthy fern discoveries in the Philippines at the turn of the 21st Century. Fern Gazette 17: 139-146
Bosman, MTM. 1991. A monograph of the fern genus Microsorum (Polypodiaceae) including an attempt towards a reconstruction of the phylogenetic history of the microsoroids. Leiden Botanical Series 14: 1-161
Brownsey PJ. 1987. A review of the fern genus Hypolepis (Dennstaedtiaceae) in the Malesian and Pacific regions. Blumea 32: 227-276
CABI Invasive Species Compendium
Chambers TC & PA Farrant. 2001. Revision of Blechnum (Blechnaceae) in Malesia. Blumea 46: 283-350
Chang, Y-H, L-Y Kuo, W-L Chiou, C-W Chen, H-M Chang & VB Amoroso. 2013. Stenolepia Alderw. (Dryopteridaceae), a fern genus new to the Philippines. Philippine Journal of Science 142: 83-88
Chao, Y-S, W-L Chiou, A Ebihara, T-C Hsu, Y-H Chang & C-Y Lin. 2021. Taxonomic and nomenclatural novelties in the Pteris fauriei group (Pteridaceae). Taiwania 66: 307-316
Chen, C-W, CJ Rothfels, AMA Mustapeng, M Gubilil, DK Karger, M Kessler & Y-M Huang. 2018. End of an enigma: Aenigmopteris belongs in Tectaria (Tectariaceae: Polypodiopsida). Journal of Plant Research 131: 67-76
Chen, C-W, LT Ngan, A Hidayat, L Evangelista, HP Nooteboom & W-L Chiou. 2014. First insights into the evolutionary history of the Davallia repens complex. Blumea 59: 49-58
Chen, C-W, S Lindsay, CM Mynssen, L-Y Kuo, VB Amoroso & Y-M Huang. 2018. A new Haplopteris species from the Philippines and clarification of the status of H. amboinensis. Garden's Bulletin Singapore 70: 177-190
Chen, C-W, S Lindsay, KT Yong, AMA Mustapeng, VB Amoroso, VD Dang & Y-M Huang. 2019. Clarification of two poorly known vittarioid ferns (Pteridaceae): Haplopteris angustissima and H. capillaris. Systematic Botany 44: 483-493
Chen, C-W, SK Tang, R Knapp, MAM Andi, P-F Lu, M Kessler & Y-M Huang. 2021. Leucostegia amplissima, the third species of the genus (Hypodematiaceae). Systematic Botany 46: 656-665.
Christenhusz, MJM, X-C Zhang & H Schneider. 2011. A linear sequence of extant families and genera of lycophytes and ferns. Phytotaxa 19: 7-54.
Copeland, E.B. 1961. Fern Flora of the Philippines 3: 547-553.
Coritico FP & VB Amoroso. 2020. Threatened lycophytes and ferns in four protected areas of Mindanao, Philippines. Nature Conservation Research 5: 78-88
Coritico FP, VB Amoroso & YC Liu. 2019. Athyrium nakanoi Makino (Athyriaceae), a new record from the Philippines and an identification key to the Malesian Athyrium sect. Polystichoides. Philippine Journal of Systematic Biology 13: 46-50
Coritico, FP, VB Amoroso & M Lehnert. 2017. New records, names and combinations of scaly tree ferns (Cyatheaceae) in eastern Malesia. Blumea 62: 92-96
Coritico, FP, VB Amoroso, FM Acma, YLL Cariño & PW Fritsch. 2020. Ferns and lycophytes of Mt. Tago Range, Bukidnon, southern Philippines: species richness, distribution, and conservation status. Philippine Journal of Science 149: 773-790
Coritico, FP, YLL Cariño, MMM Guiang & VB Amoroso. 2022. New record of Ophioderma redactophylla (Ophioglossaceae) in the Philippines and new insights to its morphology. Philippine Journal of Science 151: 227-233
Crane, E.H. 1997. A revised circumscription of the genera of the fern family Vittariaceae. Systematic Botany 22: 509-517
DENR Administrative Order. 2017. Updated national list of threatened Philippine plants and their categories. DAO 2017-11
Ding, H-H, Y-S Chao & S-Y Dong. 2013. Taxonomic novelties in the fern genus Tectaria (Tectariaceae). Phytotaxa 122: 61-64.
Dong, S-H. 2014. Nomenclatural novelties in the fern genus Tectaria (Tectariaceae). Phytotaxa 178: 225-228.
Ebihara A, CR Fraser-Jenkins, BS Parris, XC Zhang, YH Yang, WL Chiou, HM Chang, S Lindsay, D Middleton, M Kato, TN Praptosuwiryo, VB Amoroso, JF Barcelona, RHG Ranil, CH Park, N Murakami & A Hoya. 2012. Rare and threatened pteridophytes of Asia 1. An enumeration of narrowly distributed taxa. Bulletin of the National Museum of Nature and Science, Series B. 38: 93-119
Ebihara, A, J-Y Dubuisson, K Iwatsuki, S Hennequin, & M Ito. 2006. A taxonomic revision of the Hymenophyllaceae. Blumea 51: 221-280
Field, AR, W Testo, PD Bostock, JAM Holtum & M Waycott. 2016. Molecular phylogenetics and the morphology of the Lycopodiaceae subfamily Huperzioideae supports three genera: Huperzia, Phlegmariurus and Phylloglossum. Molecular Phylogenetics and Evolution 94: 635-657
Gruezo, WS. 2012. Tmesipteris zamorae: a new species of Tmesipteris Bernh. (Psilotaceae) from the Philippines. Asia Life Sciences 21: 653-670.
Hassler, M & B Schmitt. 2019. Checklist of ferns and lycophytes of the world
Hassler, M & B Schmitt. World ferns: checklist of ferns and lycophytes of the world
Hassler, M. 2019. World ferns: checklist of ferns and lycophytes of the world. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
Hauke, RI. 1963. A taxonomic monograph of the genus Equisetum subgenus Hippochaete. Beih. Nova Hedwigia 8: 1-123.
Hennipman, E & MC Roos. 1983. Phylogenetic Systematics of the Polypodiaceae (Filicales). Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversiteit te Utrecht 525: 321-342
Hennipman, E & WLA Hetterscheid. 1984. The emendation of the fern genus Christiopteris, including the transference of two taxa to the microsorioid Polypodiaceae. Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversiteit te Utrecht 539: 1-10
Hennipman, E. 1970. Some novelties in Bolbitis from Asia and the Pacific (Filices). Blumea 18: 147-149
Hennipman, E. 1974. The lectotypification of Acrostichum acuminatum Willd. and the reinstatement of the name Photinopteris speciosa (Bl.) Presl. Blumea 22: 51-52
Hennipman, E. 1977. A monograph of the fern genus Bolbitis (Lomariopsidaceae). Leiden Botanical Series 2: 1-329
Hennipman, E. 1987. A new species of Lecanopteris (Polypodiaceae) from the Philippines. Blumea 32: 311-312
Holltum, RE. 1975. Studies in the family Thelypteridaceae. IX. The genus Sphaerostephanos in the Philippines. Kalikasan 4: 47-68.
Holttum, RE & E Hennipman. 1978. Lomariopsis group. Flora Malesiana series 2, 1: 255-330
Holttum, RE, U Sen & D Mittra. 1970. Studies in the family Thelypteridaceae II. A comparative study of the type-species of Thelypteris Schmidel, Cyclosorus Link, and Ampelopteris Kunze. Blumea 18: 195-215
Holttum, RE. 1959. Cyatheaceae. Flora Malesiana series 2, 1: 65-176
Holttum, RE. 1959. Gleicheniaceae. Flora Malesiana series 2, 1: 1-36
Holttum, RE. 1959. Keys to the genera of Pteropsida. Flora Malesiana series 2, 1: 15-19
Holttum, RE. 1959. Schizaeaceae. Flora Malesiana series 2, 1: 37-61
Holttum, RE. 1966. The genera Lomariopsis, Teratophyllum, and Lomagramma in the islands of the Pacific and Australia. Blumea 14: 215-223
Holttum, RE. 1966. The genus Arthropteris J.Sm. in Malesia. Blumea 14: 225-229
Holttum, RE. 1968. A re-definition of the fern-genus Taenitis Willd. Blumea 16: 87-95
Holttum, RE. 1969. Studies in the family Thelypteridaceae. The genera Phegopteris, Pseudophegopteris, and Macrothelypteris. Blumea 17: 5-32
Holttum, RE. 1971. Studies in the Family Thelypteridaceae. III. A new system of genera in the Old World. Blumea 19: 17-52
Holttum, RE. 1972. Studies in the family Thelypteridaceae IV. The genus Pronephrium Presl. Blumea 20: 105-126
Holttum, RE. 1973. Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai. Blumea 21: 293-325
Holttum, RE. 1975. A comparative account of the fern-genera Syngramma J.Sm. and Taenitis Willd., with discussion of their relationships to each other and to other genera. Kew Bulletin 30: 327-343.
Holttum, RE. 1975. Studies in the family Thelypteridaceae VIII. The genera Mesophlebion and Plesioneuron. Blumea 22: 223-250
Holttum, RE. 1976. Studies in the family Thelypteridaceae X. The genus Coryphopteris. Blumea 23: 18-47
Holttum, RE. 1977. Studies in the family Thelypteridaceae. XII. The genus Amphineuron Holttum. Blumea 23: 205-218
Holttum, RE. 1981. Thelypteridaceae. Flora Malesiana Ser. 2, 1: 331-599
Holttum, RE. 1984. Studies in the fern-genera allied to Tectaria. III. Aenigmopteris and Ataxipteris, two new genera allied to Tectaria Cav., with comments on Psomiocarpa Presl. Blumea 30: 1-11
Holttum, RE. 1985. Studies in the fern genera allied to Tectaria Cav. IV. The genus Ctenitis in Asia, Malesia and the Western Pacific. Blumea 31: 1-38
Holttum, RE. 1991. New taxa in the Tectaria Group (Polypodiaceae) from Malesia. Blumea 35: 547-557
Holttum, RE. 1991. Tectaria group. Flora Malesiana series 2, 2: 1-132
Holttum. RE. 1966. Florae Malesianae Precursores XLIV. The fern-genus Elaphoglossum in Malesia, with descriptions of new species. Blumea 14: 317-326
Hovenkamp, PH & F Miyamoto. 2005. A conspectus of the native and naturalized species of Nephrolepis (Nephrolepidaceae) in the world. Blumea 50: 279-322
Hovenkamp, PH & F Miyamoto. 2012. Nephrolepidaceae. Flora Malesiana series 2, 4: 97-122
Hovenkamp, PH & H Boon-Chuan. 2012. A revision of the fern genus Oleandra (Oleandraceae) in Asia. PhytoKeys 11: 1-37
Hovenkamp, PH & NAP Franken. 1993. An account of the fern genus Belvisia Mirbel (Polypodiaceae). Blumea 37: 511-527
Hovenkamp, PH, MTM Bosman, E Hennipman, HP Nooteboom, G. Rodl-Linder & MC Roos. 1998. Polypodiaceae. Flora Malesiana series 2, 3: 1-234
Hovenkamp, PH. 1986. A monograph of the fern genus Pyrrosia (Polypodiaceae). Leiden Botanical Series 9: 1-280
Hovenkamp, PH. 1998. An account of the Malay-Pacific species of Selliguea (Polypodiaceae). Blumea 43: 1-108
Hovenkamp, PH. 1998. Lepisorus in Malesia. Blumea 43: 109-115
Hovenkamp, PH. 2019. Flora of Singapore precursors, 13. New names and lectotypifications in Athyriaceae and Polypodiaceae. Gard. Bull. Singapore 71: 61-67
Hsu, T-C, Y-F Huang & Y-S Chao. 2019. Systematics of Hymenophyllum subgenus Mecodium (Hymenophyllaceae) in Taiwan. Systematic Botany 44: 753-767.
Johns, RJ. 1995. Lecanopteris lomarioides: Polypodiaceae. Curtis's Botanical Magazine 12: 89-95.
Karger, DN, S Lehtonen, VB Amoroso, M Kessler. 2012. A new species of Lindsaea (Lindsaeaceae, Polypodiopsida) from Mt. Hamiguitan, Mindanao, Philippines. Phytotaxa 56: 15-20
Kato, M, Y Yatabe, N Sahashi & N Murakami. 2001. Taxonomic studies of Cheiropleuria (Dipteridaceae). Blumea 46: 513-525
Kato, M. 1993. A taxonomic study of the genus Matonia (Matoniaceae). Blumea 38: 167-172
Kato, M. 1998. Matoniaceae. Flora Malesiana series 2, 3: 289-294
Kinosian, SP, WD Pearse & PG Wolf. 2020. Cryptic diversity in the model fern genus Ceratopteris (Pteridaceae). Molecular Phylogenetics and Evolution 152: 106938.
Kramer, KU. 1967. The Lindsaeoid ferns of the Old World II. A revision of Tapeinidium. Blumea 15: 545-556
Kramer, KU. 1967. The Lindsaeoid ferns of the Old World III. Notes on Lindsaea and Sphenomeris in the Flora Malesiana area. Blumea 15: 557-574
Kramer, KU. 1971. Lindsaea Group. Flora Malesiana series 2, 1: 177-254
Laferrière, JE. 1998. Cheiropleuriaceae. Flora Malesiana series 2, 3: 285-286
Laferrière, JE. 1998. Equisetaceae. Flora Malesiana series 2, 3: 287-288
Lehnert, M & FP Coritico. 2018. The genus Dicksonia (Dicksoniaceae - Cyatheales) in western Malesia. Blumea 63: 268-278
Lin, YX & R Viane. 2013. Aspleniaceae. In: Pp. 267–316 in Wu, ZY, PH Raven & DY Hong (eds.), Flora of China, Vol. 2-3 (Pteridophytes). Beijing: Science Press; St. Louis: Missouri Botanical Garden Press
Lindsay, S, PH Hovenkamp & DJ Middleton. 2017. New combinations and typifications in Aglaomorpha (Polypodiaceae). Gardens' Bulletin Singapore 69: 149-155
Liu, H-M, R-H Jiang, J Guo, P Hovenkamp, LR Perrie, L Shepherd, S Hennequin & H Schneider. 2013. Towards a phylogenetic classification of the climbing fern genus Arthropteris. Taxon 62: 688-700
Liu, Y-C, CR Fraser-Jenkins, VB Amoroso & W-L Chiou. 2008. Athyrium erythropodum (Woodsiaceae, Pteridophyta), a new Philippine record. Blumea 53: 447-452
Luong, TT, PH Hovenkamp & MSM Sosef. 2015. Revision of the fern genus Orthiopteris (Saccolomataceae) in Malesia and adjacent regions. PhytoKeys 53: 39–71
Masuyama, S & Y Watano. 2010. Cryptic species in the fern Ceratopteris thalictroides (L.) Brongn. (Parkeriaceae). IV. Taxonomic revision. Acta Phytotaxonomica et Geobotanica 61: 75-86
Moody, K. 1989. Weeds reported in rice in South and Southeast Asia. IRRI, Philippines
Murdock, AG. 2008. A taxonomic revision of the eusporangiate fern family Marattiaceae, with description of a new genus Ptisana. Taxon 57: 737-755
Nooteboom, HP. 1992. Notes on Davalliaceae I. The genera Araiostegia, Davallodes, Leucostegia, and Gymnogrammitis. Blumea 37: 165-187
Nooteboom, HP. 1994. Notes on Davalliaceae II. A revision of the genus Davallia. Blumea 39: 151-214
Nooteboom, HP. 1997. The microsoroid ferns (Polypodiaceae). Blumea 42: 261-395
Nooteboom, HP. 1998. Davalliaceae. Flora Malesiana series 2, 3: 235-276
Paclibar, GCB & ER Tadiosa. 2020 Plant species diversity and assessment in Quezon Protected Landscape, Southern Luzon, Philippines. Philippine Journal of Systematic Biology 4: 1-19
Panigrahi, G.1993. Parahemionitis, a new genus of Pteridaceae. American Fern Journal 83: 90-92.
Parris, BS. 2007. Five new genera and three new species of Grammitidaceae (Filicales) and the re-establishment of Oreogrammitis. Garden's Bulletin Singapore 59: 233-274
Parris, BS. 2009. New genera of Malesian Grammitidaceae (Monilophyta). Blumea 54: 217-219
Parris, BS. 2021. Calymmodon pectinatus, a new species of grammitid fern (Polypodiaceae) from Malesia. Fern Gazette 21: 254-259.
Perrie, LR, AR Field, DJ Ohlsen & PJ Brownsey. 2021. Expansion of the fern genus Lecanopteris to encompass some species previously included in Microsorum and Colysis (Polypodiaceae). Blumea 66: 242-248
Petchsri, S, T Boonkerd, BR Baum, D Karladee, S Suriyong & A Lungkaphin. 2012. Phenetic study of the Microsorum punctatum complex (Polypodiaceae). ScienceAsia 38: 1-12
Picasa albums of Taiwanese pterophytes and gynosperms
Price, MG. 1990. Philippine fern notes. Contr. Univ. Mich. Herb. 17: 267-278.
Ravensberg, WJ & E Hennipman. 1986. The Pyrrosia species formerly referred to Drymoglossum and Saxiglossum (Filicales, Polypodiaceae). Leiden Botanical Series 9: 281-310
Reid, JD, GM Plunkett, GA Peters. 2006. Phylogenetic relationships in the heterosporous fern genus Azolla (Azollaceae) based on DNA sequence data from three noncoding regions. International Journal of Plant Sciences 167(3): 529-538.
Rolleri, C. 1993. Revision of the genus Christensenia. American Fern Journal 83: 3-19
Roos, MC. 1985. New combinations in Aglaomorpha Schott (Polypodiaceae). Blumea 31: 153
Ruhfel, B, S Lindsay & CC Davis. 2008. Phylogenetic placement of Rheopteris and Monogramma (Pteridaceae, s.l.): evidence from Rbcl sequence data. Systematic Botany 33: 37-43
Rödl-Linder, G. 1994. A monograph of the fern genus Thylacopteris (Polypodiaceae). Blumea 39: 351-364
Rödl-Linder. G. 1990. A monograph of the fern genus Goniophlebium (Polypodiaceae). Blumea 34: 277-423
Salgado, AE. 2017. Does Asplenium falcatum exist in the Philippines? Edinburgh Journal of Botany 74: 1-4
Salgado, AE. 2021. Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524: 71-84.
Salgado, AE. 2022. Taxonomic and nomenclatural Notes on Philippine ferns. V. On the identity of Asplenium lepturus, A. militare, and A. nigrescens (Polypodiales, Aspleniaceae. Philippine Journal of Science 151: 291-301
Saunders, RMK. 1998. Azollaceae. Flora Malesiana series 2, 3: 277-284
Schuettpelz, E & KM Pryer. 2007. Fern phylogeny inferred from 400 leptosporangiate species and three plastid genes. Taxon 56: 1037-1050
Schuettpelz, E, H Schneider, L Huiet, MD Windham & KM Pryer. 2007. A molecular phylogeny of the fern family Pteridaceae: assessing overall relationships and the affinities of previously unsampled genera. Molecular Phylogenetics and Evolution 44:1172-1185
Senterre B, G Rouhan, C Morel & JY Dubuisson. 2017. New species and records in the fern genus Didymoglossum for the flora of Seychelles, with notes on the Southeast Asian D. motleyi complex (Hymenophyllaceae). Phytotaxa 292: 201-217
Shang, H, Z-L Liang & L-B Zhang. 2021. Taxonomy of the fern genus Didymochlaena (Didymochlaenaceae) from Asia and Pacific islands based on morphological and molecular evidence with the description of four new species and one new status. Phytotaxa 479: 71-82.
Smith AR, Pryer KM, Schuettpelz E, Korall P, Schneider H & Wolf PG. 2006. A classification for extant ferns. Taxon 55: 705–731
TPG. 2019. Updating Taiwanese pteridophyte checklist: a new phylogenetic classification. Taiwania 64: 367-395
Tan, BC. 2013. The Selaginella flora, a good indicator of the Philippine pleistocene island groups. Philippine Journal of Science 142: 211-213
Testo, WL, AR Field, AB Sessa & M Sundue. 2019. Phylogenetic and morphological analyses support the resurrection of Dendroconche and the recognition of two new genera in Polypodiaceae subfamily Microsoroideae. Systematic Botany 44: 1-16
Tryon, RM. 1941. A revision of the genus Pteridium. Rhodora 43: 1-70.
Wang, LI, Z-Q Wu, QP Xiang, J Heinrichs, H Schneider & XC Zhang. 2010. A molecular phylogeny and a revised classification of tribe Lepisoreae (Polypodiaceae) based on an analysis of four plastid DNA regions. Botanical Journal of the Linnean Society 162: 28-38
Wardani, W & B Adjie. 2018. The genus Athyrium Roth in Malesia: a checklist with working names and distributions. Sibbaldia 16: 39-48
Wieffering, JH. 1964. A preliminary revision of the Indo-Pacific species of Ophioglossum (Ophioglossaceae). Blumea 12: 321-337
Yesilyurt, JC & H Schneider. 2010. The new fern genus Calciphilopteris (Pteridaceae). Phytotaxa 7: 52-59.
Zhang XC & HP Nooteboom. 1997. Some new combinations in Plagiogyria (Plagiogyriaceae, Pteridophyta). Blumea 42: 483-484
Zhang XC & HP Nooteboom. 1998. A taxonomic revision of Plagiogyriaceae (Pteridophyta). Blumea 43: 401-469
Zhang XC & HP Nooteboom. 1998. Plagiogyriaceae. Flora Malesiana series 2, 3: 295-316
Zhang, L-B & L Zhang. 2012. The inclusion of Acrophorus, Diacalpe, Nothoperanema, and Peranema in Dryopteris: The molecular phylogeny, systematics, and nomenclature of Dryopteris subg. Nothoperanema (Dryopteridaceae). Taxon, 61: 1199-1216
Zhang, L-B. 2012. Reducing the fern genus Dryopsis to Dryopteris and the systematics and nomenclature of Dryopteris subgenus Erythrovariae section Dryopsis (Dryopteridaceae). Phytotaxa 71: 17-27.
Zhang, XC. 2003. New combinations in Haplopteris (Pteridophyta: Vittariaceae). Annales Botanici Fennici 40: 459-461
Zhou, S, W Dong, X Chen, X Zhang, J Wen & H Schneider. 2014. How many species of bracken (Pteridium) are there? Assessing the Chinese brackens using molecular evidence. Taxon 63: 509-521
Zhou, X-M & L-B Zhang. 2019. A monograph of the fern genus Pteridrys (Pteridryaceae). Systematic Botany 44 (2019) 243-273.