Davalliaceae

Davallia J.Sm.

Last edited by Julie F. Barcelona & Pieter B. Pelser, 5 February 2023

Davallia angustata Wall. ex Hook. & Grev., Ic. Fil. (1831) t.231; ≡ Humata angustata (Wall. ex Hook. & Grev.) J.Sm., J. Bot. 3 (1841) 416. = Humata microsora Copel., PJS 7 c (1912) 55; Type: Weber 1146, Mindanao: Agusan del Norte, Mt Hilong-Hilong, Mar-1911. Distribution: Borneo, ?Cambodia, Malay Peninsula, Pacific Ocean, Philippines, Singapore, Sulawesi, Sumatra, Thailand, Vietnam. MINDANAO: Agusan, Agusan del Norte, Davao Oriental, Surigao del Sur, PALAWAN. Native.
Davallia brevipes Copel., PJS 1 (Suppl. 2) (1906) 147; Hovenkamp & de Joncheere, Blumea 33 (1988) 408. Distribution: Bismarck Arch, Moluccas, New Guinea, Philippines, Solomon Isls, Sulawesi. MINDANAO: Agusan del Norte, Zamboanga del Sur. Native.
Davallia denticulata (Burm.f.) Mett. ex Kuhn, Fil. Deck. (1867) 27; ≡ Wibelia denticulata (Burm.f.) M.Kato & Tsutsumi, Acta Phytotax. Geobot. 59 (2008) 13; ≡ Adiantum denticulatum Burm.f., Fl. Ind. (1768) 236. Distribution: Andaman Isls, Australia, Bismarck Arch, Borneo, Cambodia, China, India, Laos, Lesser Sunda Isls, Malay Peninsula, Myanmar, New Guinea, Nicobar Isls, Philippines, Singapore, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam. BASIAO, BASILAN, BOHOL, GENERAL ISLAND, LEYTE, LUZON: Apayao, Bataan, Benguet, Ifugao, Mountain Province, Nueva Vizcaya, Pampanga, Quezon, Rizal, MINDANAO: Bukidnon, Cotabato, Davao, Davao del Sur, Lanao del Sur, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY, SAMAR, SIBUTU. Native.
Davallia divaricata Blume, Enum. Fil. (1828) 237. Distribution: Borneo, Cambodia, China, Hong Kong, India, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, New Guinea, Philippines, Sulawesi, Sumatra, Thailand, Vietnam. BOHOL, CEBU, LUZON: Ilocos Norte, MINDANAO: Davao del Sur, NEGROS, SAMAR. Native.
Davallia embolostegia Copel., PJS 1 (Suppl. 2) (1906) 147; Rufila et al., Ruhuna J. Science 13 (2022) 156; ≡ Wibelia embolostegia (Copel.) M.Kato & Tsutsumi, Acta Phytotax. Geobot. 59 (2008) 13. Distribution: Borneo, Moluccas, Pacific Ocean, ?Pacific Ocean, Philippines, Sulawesi, Sumatra, Thailand. LUZON: Albay, Bataan, Batangas, Benguet, Kalinga, Laguna, Mountain Province, Pampanga, Quezon, Rizal, Sorsogon, MINDANAO: Davao del Sur, MINDORO, NEGROS, PANAY, SAMAR. Conservation status: Other Threatened Species (DAO 2017-11). Native.
Davallia epiphylla (G.Forst.) Spreng., Schrad., J. Bot. 1799 (1799) 271; ≡ Trichomanes epiphyllum G.Forst., Prod. (1786) 85. = Davallia denticulata var. elata (G.Forst.) Mett. ex Kuhn, Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 288; ≡ Trichomanes elatum G.Forst., Prod. (1786) 86. Distribution: Bismarck Arch, Moluccas, New Caledonia, New Guinea, Pacific Ocean, Philippines, Solomon Isls, Sulawesi. Conservation status: Other Threatened Species (DAO 2017-11; as Davallia denticulata var. elata). Native.
Davallia falcinella C.Presl, Reliq. Haenk. 1 (1825) 66; ≡ Trogostolon falcinellus (C.Presl) Copel., PJS 84 (1927) 251. Distribution: Endemic to the Philippines. BOHOL, LEYTE, LUZON: Benguet, Camarines, Quezon, Sorsogon, MINDANAO: Agusan del Norte, Surigao del Sur, Zamboanga Sibugay, NEGROS, PANAY, SAMAR. Native.
Davallia heterophylla Sm., Mém. Acad. Turin 5 (1793) 415; ≡ Humata heterophylla (Sm.) Desv., Prodr. (1825) 323. = Humata ophioglossa Cav., Descr. Pl. 1 (1802) 273. Distribution: Bismarck Arch, Borneo, Cambodia, Java, Lesser Sunda Isls, Malay Peninsula, Moluccas, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam. BALABAC, BILIRAN, CAMIGUIN, CATANDUANES, LEYTE, LUZON: Apayao, Aurora, Cagayan, Laguna, Pampanga, Quezon, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Lanao del Sur, Zamboanga del Sur, NEGROS, PALAWAN, SAMAR, SIBUYAN. Native.
Davallia hymenophylloides (Blume) Kuhn, Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 286; ≡ Araiostegia hymenophylloides (Blume) Copel., PJS 34 (1927) 241; ≡ Aspidium hymenophylloides Blume, Enum. Pl. Jav. (1828) 172. Distribution: Borneo, India, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Philippines, Sri Lanka, Sulawesi, Sumatra, Thailand. CAMIGUIN, LUZON: Aurora, Batangas, Benguet, Cagayan, Camarines Norte, Camarines Sur, Ifugao, Laguna, Mountain Province, Quezon, Rizal, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao del Sur, Lanao, Sarangani, MINDORO. Native.
Davallia lorrainii Hance, Ann. Sci. Nat. 5, 5 (1866) 254; ≡ Davallia trichomanoides var. lorrainii (Hance) Holttum, Rev. Fl. Malaya ed. 1, 2 (1955) 361. Distribution: Borneo, Cambodia, India, Malay Peninsula, Myanmar, Philippines, Sulawesi, Sumatra, Thailand, Vietnam. Conservation status: Other Threatened Species (DAO 2017-11). Native.
Davallia pectinata Sm., Mém. Acad. Turin 5 (1793) 415; ≡ Humata pectinata (Sm.) Desv., Mem. Soc. Linn. Paris 6, 2 (1827) 323. Distribution: Australia, Bismarck Arch, Borneo, ?Cambodia, Java, Malay Peninsula, Moluccas, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. CAMIGUIN, GENERAL ISLAND, LUZON: Laguna, MINDANAO: Agusan del Norte, Davao Oriental, Zamboanga del Sur, NEGROS, PALAWAN, SAMAR, SIBUTU, SULU ARCHIPELAGO, TAMLANGON, TAWI-TAWI. Native.
Davallia pubescens C.W.Chen, Sol Amazing (2017) 450; Rufila et al., Ruhuna J. Science 13 (2022) 157; ≡ Davallodes hirsuta (J.Sm.) Copel., PJS 3 c (1908) 33, nom. illeg., non Sw.; ≡ Leucostegia hirsuta J.Sm., J. Bot. 3 (1841) 416. = Davallodes grammatosorum Copel., PJS 3c (1908) 34. = Davallodes gymnocarpa Copel., PJS 3c (1908) 34. = Davallodes laxa Copel., PJS 34 (1927) 246. = Davallodes congesta Copel., PJS 34 (1927) 247. Distribution: Bismarck Arch, Borneo, New Guinea, Philippines, Solomon Isls, Sulawesi, Sumatra. MINDANAO: Bukidnon, Davao del Sur, Lanao del Sur, South Cotabato, Zamboanga del Sur, SAMAR. Native.
Davallia pusilloides (Copel. ex Alderw.) Parris, Fern Gaz. 20 (2018) 305; ≡ Pachypleuria pusilloides (Copel.) M.Kato, J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13 (1985) 573; ≡ Humata pusilloides Copel. ex Alderw., Bull. Jard. Bot. Buitenzorg Ser. 2, 28 (1918) 26. Distribution: Borneo, Philippines. LUZON: Benguet, MINDANAO: Zamboanga del Sur. Native.
Davallia repens (L.f.) Kuhn, Fil. Deck. (1867) 27, nom. cons.; Rufila et al., Ruhuna J. Science 13 (2022) 157; ≡ Humata repens (L.f.) Diels in Engl. & Prantl, Nat. Pflanzenfam. 14 (1899) 209; ≡ Adiantum repens L.f., Suppl. Pl. (1781) 446. = Humata intermedia C.Chr., Index Filic. (1905) 353. = Humata obtusata Alderw., Bull. Jard. Bot. Buitenzorg Ser. 2, I (1911) 8. = Humata vestita (Blume) T.Moore, Index (1857) 92. Distribution: Africa, Andaman Isls, Australia, Borneo, Cambodia, China, India, Indian Ocean, Japan, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, New Guinea, Pacific Ocean, Philippines, Ryukyu Isls, Singapore, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. MINDANAO: Bukidnon, Davao del Sur, PALAWAN, SAMAR. Native.
Davallia sessilifolia Blume, Enum. Fil. (1828) 231. Distribution: Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific Ocean, Philippines, Solomon Isls, Sulawesi, Sumatra. LUZON. Native.
Davallia solida (G.Forst.) Sw., J. Bot. (Schrader) 1800 (1801) 87; Coritico & Amoroso, Nature Conservation Research 5 (2020) 81. = Davallia elmeri Copel., LPB 9 (1920) 3107. = Davallia robinsonii Copel., PJS 30 (1926) 326. Distribution: BASIAO, CAMIGUIN DE BABUYANES, CATANDUANES, CEBU, GENERAL ISLAND, LUZON: Albay, Apayao, Bataan, Benguet, Ilocos Norte, Kalinga, Laguna, Mountain Province, Pampanga, Quezon, Rizal, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao, Davao Oriental, Davao del Sur, Lanao del Sur, Misamis Occidental, Surigao, Zamboanga del Sur, MINDORO, NEGROS, SAMAR, SIBUTU, TAWI-TAWI. Native.
- var. solida. Distribution: Andaman Isls, Australia, Bismarck Arch, Borneo, Cambodia, China, India, Java, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. LUZON: Apayao, Ilocos Norte, Pampanga, Quezon, Sorsogon, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao Oriental, Davao del Sur, Misamis Occidental, SAMAR. Notes: Epiphytic or epilithic, sea-level to c. 600m. Conservation status: Other Threatened Species (DAO 2017-11). Native.
Davallia trichomanoides Blume, Enum. Fil. (1828) 238. = Davallia bullata auct. non Wall. ex Hook. Distribution: Bhutan, China, India, Japan, Java, Korea, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, New Guinea, Pacific Ocean, Philippines, Ryukyu Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. LUZON: Benguet, Ifugao, Mountain Province, Nueva Vizcaya, Pampanga, MINDANAO: Bukidnon, Davao del Sur, Lanao del Sur. Native.
Davallia wagneriana Copel., Fragm. Fl. Philipp. (1905) 180. Distribution: Borneo, Moluccas, Philippines, Sulawesi, Sumatra. CAMIGUIN, LEYTE, MINDANAO: Agusan del Norte, Bukidnon, Davao del Sur, Sarangani, South Cotabato, NEGROS, PANAY. Native.

References

📋 Alston, AHG. 1959. Isoetaceae. Flora Malesiana series 2, 1: 63-64
📋 Amoroso, VB, FP Coritico & PW Fritsch. 2020. Actinostachys minuta, a new species of grass fern from Mindanao, Philippines. PhytoKeys 151: 59-66
📋 Amoroso, VB, YLL Cariño, JC Nobleza & FP Coritico. 2020. Ophioderma subsessile (Ophioglossaceae), a new snake tongue fern species from Mindanao, Philippines. Philippine Journal of Science 150 (S1): 215-221
Amoroso, VB, YLL Cariño, JC Nobleza & FP Coritico. 2021. A new species of Actinostachys (Schizaeaceae) from Mindanao, Philippines. Phytotaxa 508: 101-106.
Barcelona, JF & MG Price. 1999. A new Philippine Goniophlebium. Fern Gazette 15: 261-264.
Barcelona, JF & PB Pelser. 2014. Phanerosorus (Matoniaceae), a new fern genus record for the Philippines. Phytotaxa 170: 133-135.
📋 Barcelona, JF, B. Hernaez, & MG Price. 1996. Philippine Schizaea. Asia Life Sciences 5: 27-34
📋 Barcelona, JF, DL Nickrent, JV LaFrankie, JRC Callado & PB Pelser. 2013. Co’s Digital Flora of the Philippines: plant identification and conservation through cybertaxonomy. Philippine Journal of Science 142: 57-67
📋 Barcelona, JF, NE Dolotina, GS Madroñero, WG Granert & DD Sopot. 2006. The ferns and fern allies of the karst forests of Bohol Island, Philippines. American Fern Journal 96: 1-20
Barcelona, JF. 2002. Philippine pteridophyte collections as a resource for conservation planning. Proceedings of the “International Symposium on the Fern Flora Worldwide: Threats and Responses” sponsored by the British Pteridological Society and IUCN- Species Survival Commission.” Fern Gazette 16(6, 7 & 8): 307-312.
📋 Barcelona, JF. 2003. Preliminary report on the ferns and fern allies (pteridophytes) of Mt. Bali-it, Balbalasang-Balbalan National Park, Kalinga, Northern Luzon, Philippines. Sylvatrop: The Technical Journal of Philippine Ecosystems and Natural Resources 13: 81-92
Barcelona, JF. 2003. The taxonomy and ecology of the pteridophytes of Mt. Iraya & vicinity, Batan Island, Batanes Province, Northern Philippines, in Pteridology in the New Millenium (Chandra and Srivastava, eds). Kluwer Academy Publishers. The Netherlands, 299-325.
Barcelona, JF. 2004. Collection and conservation status of pteridophytes in Panay Island, Philippines. Philipp. Scient. 41:57-73.
📋 Barcelona, JF. 2005. Noteworthy fern discoveries in the Philippines at the turn of the 21st Century. Fern Gazette 17: 139-146
📋 Bosman, MTM. 1991. A monograph of the fern genus Microsorum (Polypodiaceae) including an attempt towards a reconstruction of the phylogenetic history of the microsoroids. Leiden Botanical Series 14: 1-161
📋 Brownsey PJ. 1987. A review of the fern genus Hypolepis (Dennstaedtiaceae) in the Malesian and Pacific regions. Blumea 32: 227-276
📋 CABI Invasive Species Compendium
📋 Chambers TC & PA Farrant. 2001. Revision of Blechnum (Blechnaceae) in Malesia. Blumea 46: 283-350
📋 Chang, Y-H, L-Y Kuo, W-L Chiou, C-W Chen, H-M Chang & VB Amoroso. 2013. Stenolepia Alderw. (Dryopteridaceae), a fern genus new to the Philippines. Philippine Journal of Science 142: 83-88
📋 Chao, Y-S, W-L Chiou, A Ebihara, T-C Hsu, Y-H Chang & C-Y Lin. 2021. Taxonomic and nomenclatural novelties in the Pteris fauriei group (Pteridaceae). Taiwania 66: 307-316
📋 Chen, C-W, CJ Rothfels, AMA Mustapeng, M Gubilil, DK Karger, M Kessler & Y-M Huang. 2018. End of an enigma: Aenigmopteris belongs in Tectaria (Tectariaceae: Polypodiopsida). Journal of Plant Research 131: 67-76
📋 Chen, C-W, LT Ngan, A Hidayat, L Evangelista, HP Nooteboom & W-L Chiou. 2014. First insights into the evolutionary history of the Davallia repens complex. Blumea 59: 49-58
📋 Chen, C-W, S Lindsay, CM Mynssen, L-Y Kuo, VB Amoroso & Y-M Huang. 2018. A new Haplopteris species from the Philippines and clarification of the status of H. amboinensis. Garden's Bulletin Singapore 70: 177-190
📋 Chen, C-W, S Lindsay, KT Yong, AMA Mustapeng, VB Amoroso, VD Dang & Y-M Huang. 2019. Clarification of two poorly known vittarioid ferns (Pteridaceae): Haplopteris angustissima and H. capillaris. Systematic Botany 44: 483-493
Chen, C-W, SK Tang, R Knapp, MAM Andi, P-F Lu, M Kessler & Y-M Huang. 2021. Leucostegia amplissima, the third species of the genus (Hypodematiaceae). Systematic Botany 46: 656-665.
Christenhusz, MJM, X-C Zhang & H Schneider. 2011. A linear sequence of extant families and genera of lycophytes and ferns. Phytotaxa 19: 7-54.
Copeland, E.B. 1961. Fern Flora of the Philippines 3: 547-553.
📋 Coritico FP & VB Amoroso. 2020. Threatened lycophytes and ferns in four protected areas of Mindanao, Philippines. Nature Conservation Research 5: 78-88
📋 Coritico FP, VB Amoroso & YC Liu. 2019. Athyrium nakanoi Makino (Athyriaceae), a new record from the Philippines and an identification key to the Malesian Athyrium sect. Polystichoides. Philippine Journal of Systematic Biology 13: 46-50
📋 Coritico, FP, VB Amoroso & M Lehnert. 2017. New records, names and combinations of scaly tree ferns (Cyatheaceae) in eastern Malesia. Blumea 62: 92-96
📋 Coritico, FP, VB Amoroso, FM Acma, YLL Cariño & PW Fritsch. 2020. Ferns and lycophytes of Mt. Tago Range, Bukidnon, southern Philippines: species richness, distribution, and conservation status. Philippine Journal of Science 149: 773-790
📋 Coritico, FP, YLL Cariño, MMM Guiang & VB Amoroso. 2022. New record of Ophioderma redactophylla (Ophioglossaceae) in the Philippines and new insights to its morphology. Philippine Journal of Science 151: 227-233
📋 Crane, E.H. 1997. A revised circumscription of the genera of the fern family Vittariaceae. Systematic Botany 22: 509-517
📋 DENR Administrative Order. 2017. Updated national list of threatened Philippine plants and their categories. DAO 2017-11
📋 Delos Angeles, MD, IE Buot Jr, L Hongmei & H Schneider. 2022. Pteridophyte diversity in the Samar Island Natural Park (SINP), Samar Island, Philippines. Philippine Journal of Science 151: 1929-1942
Ding, H-H, Y-S Chao & S-Y Dong. 2013. Taxonomic novelties in the fern genus Tectaria (Tectariaceae). Phytotaxa 122: 61-64.
Dong, S-H. 2014. Nomenclatural novelties in the fern genus Tectaria (Tectariaceae). Phytotaxa 178: 225-228.
📋 Ebihara A, CR Fraser-Jenkins, BS Parris, XC Zhang, YH Yang, WL Chiou, HM Chang, S Lindsay, D Middleton, M Kato, TN Praptosuwiryo, VB Amoroso, JF Barcelona, RHG Ranil, CH Park, N Murakami & A Hoya. 2012. Rare and threatened pteridophytes of Asia 1. An enumeration of narrowly distributed taxa. Bulletin of the National Museum of Nature and Science, Series B. 38: 93-119
📋 Ebihara, A, J-Y Dubuisson, K Iwatsuki, S Hennequin, & M Ito. 2006. A taxonomic revision of the Hymenophyllaceae. Blumea 51: 221-280
📋 Field, AR, W Testo, PD Bostock, JAM Holtum & M Waycott. 2016. Molecular phylogenetics and the morphology of the Lycopodiaceae subfamily Huperzioideae supports three genera: Huperzia, Phlegmariurus and Phylloglossum. Molecular Phylogenetics and Evolution 94: 635-657
Gruezo, WS. 2012. Tmesipteris zamorae: a new species of Tmesipteris Bernh. (Psilotaceae) from the Philippines. Asia Life Sciences 21: 653-670.
📋 Hassler, M & B Schmitt. 2019. Checklist of ferns and lycophytes of the world
📋 Hassler, M & B Schmitt. World ferns: checklist of ferns and lycophytes of the world
Hassler, M. 2019. World ferns: checklist of ferns and lycophytes of the world. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
Hauke, RI. 1963. A taxonomic monograph of the genus Equisetum subgenus Hippochaete. Beih. Nova Hedwigia 8: 1-123.
📋 Hennipman, E & MC Roos. 1983. Phylogenetic Systematics of the Polypodiaceae (Filicales). Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversiteit te Utrecht 525: 321-342
📋 Hennipman, E & WLA Hetterscheid. 1984. The emendation of the fern genus Christiopteris, including the transference of two taxa to the microsorioid Polypodiaceae. Mededelingen van het Botanisch Museum en Herbarium van de Rijksuniversiteit te Utrecht 539: 1-10
📋 Hennipman, E. 1970. Some novelties in Bolbitis from Asia and the Pacific (Filices). Blumea 18: 147-149
📋 Hennipman, E. 1974. The lectotypification of Acrostichum acuminatum Willd. and the reinstatement of the name Photinopteris speciosa (Bl.) Presl. Blumea 22: 51-52
📋 Hennipman, E. 1977. A monograph of the fern genus Bolbitis (Lomariopsidaceae). Leiden Botanical Series 2: 1-329
📋 Hennipman, E. 1987. A new species of Lecanopteris (Polypodiaceae) from the Philippines. Blumea 32: 311-312
Holltum, RE. 1975. Studies in the family Thelypteridaceae. IX. The genus Sphaerostephanos in the Philippines. Kalikasan 4: 47-68.
📋 Holttum, RE & E Hennipman. 1978. Lomariopsis group. Flora Malesiana series 2, 1: 255-330
📋 Holttum, RE, U Sen & D Mittra. 1970. Studies in the family Thelypteridaceae II. A comparative study of the type-species of Thelypteris Schmidel, Cyclosorus Link, and Ampelopteris Kunze. Blumea 18: 195-215
📋 Holttum, RE. 1959. Cyatheaceae. Flora Malesiana series 2, 1: 65-176
📋 Holttum, RE. 1959. Gleicheniaceae. Flora Malesiana series 2, 1: 1-36
📋 Holttum, RE. 1959. Keys to the genera of Pteropsida. Flora Malesiana series 2, 1: 15-19
📋 Holttum, RE. 1959. Schizaeaceae. Flora Malesiana series 2, 1: 37-61
📋 Holttum, RE. 1966. The genera Lomariopsis, Teratophyllum, and Lomagramma in the islands of the Pacific and Australia. Blumea 14: 215-223
📋 Holttum, RE. 1966. The genus Arthropteris J.Sm. in Malesia. Blumea 14: 225-229
📋 Holttum, RE. 1968. A re-definition of the fern-genus Taenitis Willd. Blumea 16: 87-95
📋 Holttum, RE. 1969. Studies in the family Thelypteridaceae. The genera Phegopteris, Pseudophegopteris, and Macrothelypteris. Blumea 17: 5-32
📋 Holttum, RE. 1971. Studies in the Family Thelypteridaceae. III. A new system of genera in the Old World. Blumea 19: 17-52
📋 Holttum, RE. 1972. Studies in the family Thelypteridaceae IV. The genus Pronephrium Presl. Blumea 20: 105-126
📋 Holttum, RE. 1973. Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai. Blumea 21: 293-325
Holttum, RE. 1975. A comparative account of the fern-genera Syngramma J.Sm. and Taenitis Willd., with discussion of their relationships to each other and to other genera. Kew Bulletin 30: 327-343.
📋 Holttum, RE. 1975. Studies in the family Thelypteridaceae VIII. The genera Mesophlebion and Plesioneuron. Blumea 22: 223-250
📋 Holttum, RE. 1976. Studies in the family Thelypteridaceae X. The genus Coryphopteris. Blumea 23: 18-47
📋 Holttum, RE. 1977. Studies in the family Thelypteridaceae. XII. The genus Amphineuron Holttum. Blumea 23: 205-218
📋 Holttum, RE. 1981. Thelypteridaceae. Flora Malesiana Ser. 2, 1: 331-599
📋 Holttum, RE. 1984. Studies in the fern-genera allied to Tectaria. III. Aenigmopteris and Ataxipteris, two new genera allied to Tectaria Cav., with comments on Psomiocarpa Presl. Blumea 30: 1-11
📋 Holttum, RE. 1985. Studies in the fern genera allied to Tectaria Cav. IV. The genus Ctenitis in Asia, Malesia and the Western Pacific. Blumea 31: 1-38
📋 Holttum, RE. 1991. New taxa in the Tectaria Group (Polypodiaceae) from Malesia. Blumea 35: 547-557
📋 Holttum, RE. 1991. Tectaria group. Flora Malesiana series 2, 2: 1-132
📋 Holttum. RE. 1966. Florae Malesianae Precursores XLIV. The fern-genus Elaphoglossum in Malesia, with descriptions of new species. Blumea 14: 317-326
📋 Hovenkamp, PH & F Miyamoto. 2005. A conspectus of the native and naturalized species of Nephrolepis (Nephrolepidaceae) in the world. Blumea 50: 279-322
📋 Hovenkamp, PH & F Miyamoto. 2012. Nephrolepidaceae. Flora Malesiana series 2, 4: 97-122
📋 Hovenkamp, PH & H Boon-Chuan. 2012. A revision of the fern genus Oleandra (Oleandraceae) in Asia. PhytoKeys 11: 1-37
📋 Hovenkamp, PH & NAP Franken. 1993. An account of the fern genus Belvisia Mirbel (Polypodiaceae). Blumea 37: 511-527
📋 Hovenkamp, PH, MTM Bosman, E Hennipman, HP Nooteboom, G. Rodl-Linder & MC Roos. 1998. Polypodiaceae. Flora Malesiana series 2, 3: 1-234
📋 Hovenkamp, PH. 1986. A monograph of the fern genus Pyrrosia (Polypodiaceae). Leiden Botanical Series 9: 1-280
📋 Hovenkamp, PH. 1998. An account of the Malay-Pacific species of Selliguea (Polypodiaceae). Blumea 43: 1-108
📋 Hovenkamp, PH. 1998. Lepisorus in Malesia. Blumea 43: 109-115
📋 Hovenkamp, PH. 2019. Flora of Singapore precursors, 13. New names and lectotypifications in Athyriaceae and Polypodiaceae. Gard. Bull. Singapore 71: 61-67
Hsu, T-C, Y-F Huang & Y-S Chao. 2019. Systematics of Hymenophyllum subgenus Mecodium (Hymenophyllaceae) in Taiwan. Systematic Botany 44: 753-767.
Johns, RJ. 1995. Lecanopteris lomarioides: Polypodiaceae. Curtis's Botanical Magazine 12: 89-95.
📋 Karger, DN, S Lehtonen, VB Amoroso, M Kessler. 2012. A new species of Lindsaea (Lindsaeaceae, Polypodiopsida) from Mt. Hamiguitan, Mindanao, Philippines. Phytotaxa 56: 15-20
📋 Kato, M, Y Yatabe, N Sahashi & N Murakami. 2001. Taxonomic studies of Cheiropleuria (Dipteridaceae). Blumea 46: 513-525
📋 Kato, M. 1993. A taxonomic study of the genus Matonia (Matoniaceae). Blumea 38: 167-172
📋 Kato, M. 1998. Matoniaceae. Flora Malesiana series 2, 3: 289-294
Kinosian, SP, WD Pearse & PG Wolf. 2020. Cryptic diversity in the model fern genus Ceratopteris (Pteridaceae). Molecular Phylogenetics and Evolution 152: 106938.
📋 Kramer, KU. 1967. The Lindsaeoid ferns of the Old World II. A revision of Tapeinidium. Blumea 15: 545-556
📋 Kramer, KU. 1967. The Lindsaeoid ferns of the Old World III. Notes on Lindsaea and Sphenomeris in the Flora Malesiana area. Blumea 15: 557-574
📋 Kramer, KU. 1971. Lindsaea Group. Flora Malesiana series 2, 1: 177-254
📋 Laferrière, JE. 1998. Cheiropleuriaceae. Flora Malesiana series 2, 3: 285-286
📋 Laferrière, JE. 1998. Equisetaceae. Flora Malesiana series 2, 3: 287-288
📋 Lehnert, M & FP Coritico. 2018. The genus Dicksonia (Dicksoniaceae - Cyatheales) in western Malesia. Blumea 63: 268-278
📋 Lin, YX & R Viane. 2013. Aspleniaceae. In: Pp. 267–316 in Wu, ZY, PH Raven & DY Hong (eds.), Flora of China, Vol. 2-3 (Pteridophytes). Beijing: Science Press; St. Louis: Missouri Botanical Garden Press
📋 Lindsay, S, PH Hovenkamp & DJ Middleton. 2017. New combinations and typifications in Aglaomorpha (Polypodiaceae). Gardens' Bulletin Singapore 69: 149-155
📋 Liu, H-M, R-H Jiang, J Guo, P Hovenkamp, LR Perrie, L Shepherd, S Hennequin & H Schneider. 2013. Towards a phylogenetic classification of the climbing fern genus Arthropteris. Taxon 62: 688-700
📋 Liu, Y-C, CR Fraser-Jenkins, VB Amoroso & W-L Chiou. 2008. Athyrium erythropodum (Woodsiaceae, Pteridophyta), a new Philippine record. Blumea 53: 447-452
📋 Luong, TT, PH Hovenkamp & MSM Sosef. 2015. Revision of the fern genus Orthiopteris (Saccolomataceae) in Malesia and adjacent regions. PhytoKeys 53: 39–71
📋 Masuyama, S & Y Watano. 2010. Cryptic species in the fern Ceratopteris thalictroides (L.) Brongn. (Parkeriaceae). IV. Taxonomic revision. Acta Phytotaxonomica et Geobotanica 61: 75-86
📋 Moody, K. 1989. Weeds reported in rice in South and Southeast Asia. IRRI, Philippines
📋 Murdock, AG. 2008. A taxonomic revision of the eusporangiate fern family Marattiaceae, with description of a new genus Ptisana. Taxon 57: 737-755
📋 Nooteboom, HP. 1992. Notes on Davalliaceae I. The genera Araiostegia, Davallodes, Leucostegia, and Gymnogrammitis. Blumea 37: 165-187
📋 Nooteboom, HP. 1994. Notes on Davalliaceae II. A revision of the genus Davallia. Blumea 39: 151-214
📋 Nooteboom, HP. 1997. The microsoroid ferns (Polypodiaceae). Blumea 42: 261-395
📋 Nooteboom, HP. 1998. Davalliaceae. Flora Malesiana series 2, 3: 235-276
📋 Paclibar, GCB & ER Tadiosa. 2020 Plant species diversity and assessment in Quezon Protected Landscape, Southern Luzon, Philippines. Philippine Journal of Systematic Biology 4: 1-19
Panigrahi, G.1993. Parahemionitis, a new genus of Pteridaceae. American Fern Journal 83: 90-92.
📋 Parris, BS. 2007. Five new genera and three new species of Grammitidaceae (Filicales) and the re-establishment of Oreogrammitis. Garden's Bulletin Singapore 59: 233-274
📋 Parris, BS. 2009. New genera of Malesian Grammitidaceae (Monilophyta). Blumea 54: 217-219
Parris, BS. 2021. Calymmodon pectinatus, a new species of grammitid fern (Polypodiaceae) from Malesia. Fern Gazette 21: 254-259.
📋 Perrie, LR, AR Field, DJ Ohlsen & PJ Brownsey. 2021. Expansion of the fern genus Lecanopteris to encompass some species previously included in Microsorum and Colysis (Polypodiaceae). Blumea 66: 242-248
📋 Petchsri, S, T Boonkerd, BR Baum, D Karladee, S Suriyong & A Lungkaphin. 2012. Phenetic study of the Microsorum punctatum complex (Polypodiaceae). ScienceAsia 38: 1-12
📋 Picasa albums of Taiwanese pterophytes and gynosperms
Price, MG. 1990. Philippine fern notes. Contr. Univ. Mich. Herb. 17: 267-278.
📋 Ravensberg, WJ & E Hennipman. 1986. The Pyrrosia species formerly referred to Drymoglossum and Saxiglossum (Filicales, Polypodiaceae). Leiden Botanical Series 9: 281-310
Reid, JD, GM Plunkett, GA Peters. 2006. Phylogenetic relationships in the heterosporous fern genus Azolla (Azollaceae) based on DNA sequence data from three noncoding regions. International Journal of Plant Sciences 167(3): 529-538.
📋 Rolleri, C. 1993. Revision of the genus Christensenia. American Fern Journal 83: 3-19
📋 Roos, MC. 1985. New combinations in Aglaomorpha Schott (Polypodiaceae). Blumea 31: 153
📋 Ruhfel, B, S Lindsay & CC Davis. 2008. Phylogenetic placement of Rheopteris and Monogramma (Pteridaceae, s.l.): evidence from Rbcl sequence data. Systematic Botany 33: 37-43
📋 Rödl-Linder, G. 1994. A monograph of the fern genus Thylacopteris (Polypodiaceae). Blumea 39: 351-364
📋 Rödl-Linder. G. 1990. A monograph of the fern genus Goniophlebium (Polypodiaceae). Blumea 34: 277-423
📋 Salgado, AE. 2017. Does Asplenium falcatum exist in the Philippines? Edinburgh Journal of Botany 74: 1-4
Salgado, AE. 2021. Taxonomic and nomenclatural notes on Philippine ferns. IV. Asplenium pahli (Polypodiales, Aspleniineae, Aspleniaceae), a new species from Leyte, Philippines. Phytotaxa 524: 71-84.
📋 Salgado, AE. 2022. Taxonomic and nomenclatural Notes on Philippine ferns. V. On the identity of Asplenium lepturus, A. militare, and A. nigrescens (Polypodiales, Aspleniaceae. Philippine Journal of Science 151: 291-301
📋 Saunders, RMK. 1998. Azollaceae. Flora Malesiana series 2, 3: 277-284
📋 Schuettpelz, E & KM Pryer. 2007. Fern phylogeny inferred from 400 leptosporangiate species and three plastid genes. Taxon 56: 1037-1050
📋 Schuettpelz, E, H Schneider, L Huiet, MD Windham & KM Pryer. 2007. A molecular phylogeny of the fern family Pteridaceae: assessing overall relationships and the affinities of previously unsampled genera. Molecular Phylogenetics and Evolution 44:1172-1185
📋 Senterre B, G Rouhan, C Morel & JY Dubuisson. 2017. New species and records in the fern genus Didymoglossum for the flora of Seychelles, with notes on the Southeast Asian D. motleyi complex (Hymenophyllaceae). Phytotaxa 292: 201-217
Shang, H, Z-L Liang & L-B Zhang. 2021. Taxonomy of the fern genus Didymochlaena (Didymochlaenaceae) from Asia and Pacific islands based on morphological and molecular evidence with the description of four new species and one new status. Phytotaxa 479: 71-82.
📋 Smith AR, Pryer KM, Schuettpelz E, Korall P, Schneider H & Wolf PG. 2006. A classification for extant ferns. Taxon 55: 705–731
📋 TPG. 2019. Updating Taiwanese pteridophyte checklist: a new phylogenetic classification. Taiwania 64: 367-395
📋 Tan, BC. 2013. The Selaginella flora, a good indicator of the Philippine pleistocene island groups. Philippine Journal of Science 142: 211-213
📋 Testo, WL, AR Field, AB Sessa & M Sundue. 2019. Phylogenetic and morphological analyses support the resurrection of Dendroconche and the recognition of two new genera in Polypodiaceae subfamily Microsoroideae. Systematic Botany 44: 1-16
Tryon. 1941. A revision of the genus Pteridium. Rhodora 43: 1-70.
📋 Wang, LI, Z-Q Wu, QP Xiang, J Heinrichs, H Schneider & XC Zhang. 2010. A molecular phylogeny and a revised classification of tribe Lepisoreae (Polypodiaceae) based on an analysis of four plastid DNA regions. Botanical Journal of the Linnean Society 162: 28-38
📋 Wardani, W & B Adjie. 2018. The genus Athyrium Roth in Malesia: a checklist with working names and distributions. Sibbaldia 16: 39-48
📋 Wieffering, JH. 1964. A preliminary revision of the Indo-Pacific species of Ophioglossum (Ophioglossaceae). Blumea 12: 321-337
Yesilyurt, JC & H Schneider. 2010. The new fern genus Calciphilopteris (Pteridaceae). Phytotaxa 7: 52-59.
📋 Zhang XC & HP Nooteboom. 1997. Some new combinations in Plagiogyria (Plagiogyriaceae, Pteridophyta). Blumea 42: 483-484
📋 Zhang XC & HP Nooteboom. 1998. A taxonomic revision of Plagiogyriaceae (Pteridophyta). Blumea 43: 401-469
📋 Zhang XC & HP Nooteboom. 1998. Plagiogyriaceae. Flora Malesiana series 2, 3: 295-316
📋 Zhang, L-B & L Zhang. 2012. The inclusion of Acrophorus, Diacalpe, Nothoperanema, and Peranema in Dryopteris: The molecular phylogeny, systematics, and nomenclature of Dryopteris subg. Nothoperanema (Dryopteridaceae). Taxon, 61: 1199-1216
📋 Zhang, L-B. 2012. Reducing the fern genus Dryopsis to Dryopteris and the systematics and nomenclature of Dryopteris subgenus Erythrovariae section Dryopsis (Dryopteridaceae). Phytotaxa 71: 17-27.
📋 Zhang, XC. 2003. New combinations in Haplopteris (Pteridophyta: Vittariaceae). Annales Botanici Fennici 40: 459-461
📋 Zhou, S, W Dong, X Chen, X Zhang, J Wen & H Schneider. 2014. How many species of bracken (Pteridium) are there? Assessing the Chinese brackens using molecular evidence. Taxon 63: 509-521
Zhou, X-M & L-B Zhang. 2019. A monograph of the fern genus Pteridrys (Pteridryaceae). Systematic Botany 44 (2019) 243-273.