Davalliaceae

Davallia J.Sm.

Last edited by Pieter B. Pelser, 8 July 2026
  1. Davallia angustata Wall. ex Hook. & Grev., Icon. Filic. 2 (1832) t.231; Nooteboom, Fl. Males. ser. 2, 3 (1998) 247; ≡ Humata angustata (Wall. ex Hook. & Grev.) J.Sm., J. Bot. (Hooker) 3 (1841) 416. = Humata microsora Copel., PJS 7c (1912) 55; Type: Weber 1146, Mindanao: Agusan del Norte, Mt Hilong-Hilong, Mar-1911. Distribution: Borneo, ?Cambodia, Malay Peninsula, Pacific Ocean, Philippines, Singapore, Sulawesi, Sumatra, Thailand, Vietnam. MINDANAO: Agusan, Agusan del Norte, Bukidnon (photos), Davao Oriental, Surigao del Sur, PALAWAN. Notes: 0-1300m. Native.
  2. Davallia brevipes Copel., PJS 1(Suppl.) (1906) 147; Hovenkamp & de Joncheere, Blumea 33 (1988) 408; Nooteboom, Fl. Males. ser. 2, 3 (1998) 248. Distribution: Bismarck Arch, Moluccas, New Guinea, Philippines, Solomon Isls, Sulawesi. MINDANAO: Agusan del Norte, South Cotabato, Zamboanga del Sur. Notes: 80-1100m. Native.
  3. Davallia denticulata (Burm.f.) Mett., Filic. Afr. (Kuhn) (1867) 27; Nooteboom, Fl. Males. ser. 2, 3 (1998) 250; Pescuela et al., PJS 153 (2024) 664; ≡ Wibelia denticulata (Burm.f.) M.Kato & Tsutsumi, Acta Phytotax. Geobot. 59 (2008) 13; ≡ Adiantum denticulatum Burm.f., Fl. Ind. (N. L. Burman) (1768) 236. = ?Davallia elegans Sw., J. Bot. (Schrader) 1800 (1801) 87. Distribution: Andaman Isls, Australia, Bismarck Arch, Borneo, Cambodia, China, India, Laos, Lesser Sunda Isls, Malay Peninsula, Myanmar, New Guinea, Nicobar Isls, Philippines, Singapore, Sri Lanka, Sulawesi, Sumatra, Thailand, Vietnam. BASIAO, BASILAN, BOHOL, GENERAL ISLAND, LEYTE, LUZON: Apayao, Aurora, Bataan, Benguet, Ifugao, Laguna, Mountain Province, Nueva Vizcaya, Pampanga, Quezon, Rizal, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao del Sur, Lanao del Sur, Misamis Oriental, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY, POLILLO, SAMAR, SIBUTU, TAMLANGON. Notes: 0-2200m. Native.
  4. Davallia divaricata Blume, Enum. Pl. Javae (1828) 237; Nooteboom, Fl. Males. ser. 2, 3 (1998) 252. = Davallia decurrens Hook., Sp. Fil. 1 (1845) 167. Distribution: Borneo, Cambodia, China, Hong Kong, India, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, New Guinea, Philippines, Sulawesi, Sumatra, Thailand, Vietnam. BATAN, BOHOL, CEBU, LUZON: Apayao, Benguet, Ilocos Norte, Isabela, Kalinga-Apayao, Mountain Province, MINDANAO: Agusan del Norte, Cotabato, Davao, Davao Oriental, Davao del Sur, Lanao del Sur, South Cotabato, Zamboanga del Sur, NEGROS, POLILLO, SAMAR. Notes: 0-1850m. Native.
    • var. divaricata. Distribution: Andaman Isls, Australia, Bismarck Arch, Borneo, Cambodia, China, India, Java, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. BATAN, CEBU, LUZON: Apayao, Camarines Norte, Ilocos Norte, Pampanga, Quezon, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao Oriental, Davao del Sur, Misamis Occidental, SAMAR. Notes: 0-1850m. Native.
  5. Davallia embolostegia Copel., PJS 1(Suppl.) (1906) 147; Nooteboom, Fl. Males. ser. 2, 3 (1998) 253; Rufila et al., Ruhuna J. Science 13 (2022) 156; ≡ Wibelia embolostegia (Copel.) M.Kato & Tsutsumi, Acta Phytotax. Geobot. 59 (2008) 13. Distribution: Borneo, Moluccas, Pacific Ocean, ?Pacific Ocean, Philippines, Sulawesi, Sumatra, Thailand. LUZON: Albay, Bataan, Batangas, Benguet, Camarines Sur, Kalinga, Laguna, Mountain Province, Pampanga, Quezon, Rizal, Sorsogon, Zambales, MINDANAO: Bukidnon (photos), Davao del Sur, MINDORO, NEGROS, PANAY, SAMAR. Notes: 0-2100m. Native.
  6. Davallia epiphylla (G.Forst.) Sw., Syn. Fil. (Swartz) (1806) 134; ≡ Davallia epiphylla (G.Forst.) Spreng., J. Bot. (Schrader) 1799 (1799) 271, nom. inval.; ≡ Trichomanes epiphyllum G.Forst., Fl. Ins. Austr. (1786) 85. = Davallia denticulata var. elata (G.Forst.) Mett. ex Kuhn, Ann. Mus. Bot. Lugduno-Batavi 4 (1869) 288; Nooteboom, Fl. Males. ser. 2, 3 (1998) 251; ≡ Trichomanes elatum G.Forst., Fl. Ins. Austr. (1786) 86. Distribution: Bismarck Arch, Moluccas, New Caledonia, New Guinea, Pacific Ocean, Philippines, Solomon Isls, Sulawesi. LUZON: Bataan, Camarines Norte, Ifugao, Zambales, MINDANAO: Davao. Notes: 0-1600m. Native.
  7. Davallia falcinella C.Presl, Reliq. Haenk. 1 (1825) 66; Nooteboom, Fl. Males. ser. 2, 3 (1998) 254; ≡ Trogostolon falcinellus (C.Presl) Copel., PJS 34 (1927) 251. Distribution: Endemic to the Philippines. BOHOL, LEYTE, LUZON: Benguet, Camarines, Camarines Norte, Quezon, Sorsogon, MINDANAO: Agusan del Norte, Surigao, Surigao del Sur, Zamboanga, Zamboanga Sibugay, NEGROS, PANAY, SAMAR. Notes: 0-800m. Native.
  8. Davallia heterophylla Sm., Mém. Acad. Roy. Sci. (Turin) 5 (1793) 415; Nooteboom, Fl. Males. ser. 2, 3 (1998) 254; ≡ Humata heterophylla (Sm.) Desv., Mém. Soc. Linn. Paris 6 (1827) 323. = Humata ophioglossa Cav., Descr. Pl. (Cavanilles) (1801) 273. Distribution: Bismarck Arch, Borneo, Cambodia, Java, Lesser Sunda Isls, Malay Peninsula, Moluccas, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sulawesi, Sumatra, Thailand, Vietnam. BALABAC, BILIRAN, CAMIGUIN, CATANDUANES, DINAGAT (photos), LEYTE, LUZON: Apayao, Aurora, Bataan, Cagayan, Isabela (photos), Laguna, Pampanga, Quezon, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Davao Oriental (photos), Lanao del Sur, Zamboanga del Sur, NEGROS, PALAWAN, PANAY (photos), POLILLO, SAMAR, SIBUYAN. Notes: 0-900(-2100)m. Native.
  9. Davallia lorrainii Hance, Ann. Sci. Nat. (Paris), 5th Sér. 5 (1866) 254; ≡ Davallia trichomanoides var. lorrainii (Hance) Holttum, Rev. Fl. Malaya 2 (1955) 361; Nooteboom, Fl. Males. ser. 2, 3 (1998) 265. Distribution: Borneo, Cambodia, India, Malay Peninsula, Myanmar, Philippines, Sulawesi, Sumatra, Thailand, Vietnam. LUZON: Abra, Benguet, Camarines Norte, Ilocos Norte, Kalinga, Kalinga-Apayao, Mountain Province, MINDANAO: Bukidnon (photos), Davao, Davao del Norte, Davao del Sur. Notes: 100-3500m. Native.
  10. Davallia pectinata Sm., Mém. Acad. Roy. Sci. (Turin) 5 (1793) 415; Nooteboom, Fl. Males. ser. 2, 3 (1998) 258; ≡ Humata pectinata (Sm.) Desv., Mém. Soc. Linn. Paris 6 (1827) 323. Distribution: Australia, Bismarck Arch, Borneo, ?Cambodia, Java, Malay Peninsula, Moluccas, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. BOHOL, CAMIGUIN, GENERAL ISLAND, LUZON: Bataan, Laguna, Quezon, MINDANAO: Agusan del Norte, Davao Oriental, Surigao, Zamboanga del Sur, NEGROS, PALAWAN, POLILLO, SAMAR, SIBUTU, SULU ARCHIPELAGO, TAMLANGON, TAWI-TAWI. Native.
  11. Davallia pusilloides (Copel. ex Alderw.) Parris, Fern Gaz. 20 (2018) 305; ≡ Pachypleuria pusilloides (Copel.) M.Kato, J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13 (1985) 573; ≡ Humata pusilloides Copel. ex Alderw., Bull. Jard. Bot. Buitenzorg, 2nd Sér. 28 (1918) 26. Distribution: Borneo, Philippines. LUZON: Benguet, MINDANAO: Zamboanga del Sur, SIBUYAN. Native.
  12. Davallia repens (L.f.) Kuhn, Filic. Afr. (Kuhn) (1867) 27, nom. cons.; Nooteboom, Fl. Males. ser. 2, 3 (1998) 259; Rufila et al., Ruhuna J. Science 13 (2022) 157; Pescuela et al., PJS 153 (2024) 664; ≡ Humata repens (L.f.) Diels, Nat. Pflanzenfam. (Engler & Prantl) 1(4) (1899) 209; ≡ Adiantum repens L.f., Suppl. Pl. (1782) 446. = Humata intermedia C.Chr., Index Filic. (C.Chr) (1905) 353. = Humata obtusata Alderw., Bull. Jard. Bot. Buitenzorg, 2nd Sér. 1 (1911) 8. = Humata vestita (Blume) T.Moore, Index Fil. (T.Moore) (1857) 92. Distribution: Africa, Andaman Isls, Australia, Borneo, Cambodia, China, India, Indian Ocean, Japan, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, New Guinea, Pacific Ocean, Philippines, Ryukyu Isls, Singapore, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. BATAN, BILIRAN, BOHOL, CAMIGUIN, JOLO, LEYTE, LUZON: Abra, Albay, Apayao, Aurora, Bataan, Benguet, Cagayan, Camarines Norte, Camarines Sur, Ifugao, Ilocos Norte, Isabela, Kalinga, Laguna, Mountain Province, NCR, Nueva Vizcaya (photos), Pampanga, Quezon, Rizal, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao Oriental, Davao de Oro (photos), Davao del Norte, Davao del Sur, Lanao del Sur, Misamis Oriental, Sarangani, South Cotabato, Surigao, Zamboanga del Sur, MINDORO, NEGROS, PALAWAN, PANAY, SAMAR, SIBUYAN. Notes: 0-3420m. Native.
  13. Davallia sessilifolia Blume, Enum. Pl. Javae (1828) 231; Nooteboom, Fl. Males. ser. 2, 3 (1998) 262. Distribution: Borneo, Java, Lesser Sunda Isls, Moluccas, New Guinea, Pacific Ocean, Philippines, Solomon Isls, Sulawesi, Sumatra. LUZON. Notes: 150-1770m. Native.
  14. Davallia solida (G.Forst.) Sw., J. Bot. (Schrader) 1800 (1801) 87; Nooteboom, Fl. Males. ser. 2, 3 (1998) 263; Coritico & Amoroso, Nature Conservation Research 5 (2020) 81; Raganas et al., Biodiversitas 26 (2025) 1195. = Davallia elmeri Copel., LPB 9 (1920) 3107. = Davallia robinsonii Copel., PJS 30 (1926) 326. Distribution: BASIAO, BATAN, BOHOL, CAMIGUIN, CAMIGUIN DE BABUYANES, CATANDUANES, CEBU, GENERAL ISLAND, LEYTE (photos), LUZON: Albay, Apayao, Bataan, Benguet, Cagayan, Camarines Norte, Ilocos Norte, Kalinga, Laguna, Mountain Province, Nueva Vizcaya, Pampanga, Quezon, Rizal, Sorsogon, Zambales, MINDANAO: Agusan, Agusan del Norte, Bukidnon, Cotabato, Davao, Davao Oriental, Davao del Norte, Davao del Sur, Lanao del Sur, Misamis Occidental, Misamis Oriental (photos), Surigao, Zamboanga del Sur, MINDORO, NEGROS, PANAY (photos), SABTANG, SAMAR, SIBUTU, TAWI-TAWI. Notes: 0-1500m. Native.
    • var. solida. Distribution: Andaman Isls, Australia, Bismarck Arch, Borneo, Cambodia, China, India, Java, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, New Caledonia, New Guinea, Nicobar Isls, Pacific Ocean, Philippines, Singapore, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. BATAN, CEBU, LUZON: Apayao, Camarines Norte, Ilocos Norte, Pampanga, Quezon, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao Oriental, Davao del Sur, Misamis Occidental, SAMAR. Notes: Epiphytic or epilithic. 0-1500m. Native.
  15. Davallia trichomanoides Blume, Enum. Pl. Javae (1828) 238; Nooteboom, Fl. Males. ser. 2, 3 (1998) 264; Odulio et al., Biodivers. Data J. 14 (2026) 15. ≠ Davallia bullata auct. non Wall. ex Hook. Distribution: Bhutan, China, India, Japan, Java, Korea, Laos, Lesser Sunda Isls, Malay Peninsula, Moluccas, Myanmar, New Guinea, Pacific Ocean, Philippines, Ryukyu Isls, Sulawesi, Sumatra, Taiwan, Thailand, Vietnam. LUZON: Benguet, Ifugao, Mountain Province, Nueva Vizcaya, Pampanga, MINDANAO: Bukidnon, Cotabato, Davao del Sur, Lanao del Sur, Surigao del Sur (photos), SAMAR. Notes: 100-3500m. Native.
  16. Davallia wagneriana Copel., Fragm. Fl. Philipp. (1905) 180; Nooteboom, Fl. Males. ser. 2, 3 (1998) 266; Pescuela et al., PJS 153 (2024) 664. Distribution: Borneo, Moluccas, Philippines, Sulawesi, Sumatra. CAMIGUIN, LEYTE, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao, Davao del Sur, Misamis Oriental, Sarangani, South Cotabato, NEGROS, PANAY. Notes: 450-1600m. Native.

Davallodes (Copel.) Copel.

Last edited by Julie F. Barcelona & Pieter B. Pelser, 10 Juni 2026
  1. Davallodes hirsuta (C.Presl) Copel., PJS 3c (1908) 33; Nooteboom, Fl. Males. ser. 2, 3 (1998) 270; ≡ Davallia pubescens C.W.Chen, Sol Amazing (2017) 450; Rufila et al., Ruhuna J. Science 13 (2022) 157; ≡ Microlepia hirsuta C.Presl, Abh. Königl. Böhm. Ges. Wiss., 5th Ser. 6 (1851) 457; ≡ Leucostegia hirsuta J.Sm., J. Bot. (Hooker) 3 (1841) 416, nom. inval., nom. nud. = Davallodes grammatosorum Copel., PJS 3c (1908) 34. = Davallodes gymnocarpa Copel., PJS 3c (1908) 34. = Davallodes laxa Copel., PJS 34 (1927) 246. = Davallodes congesta Copel., PJS 34 (1927) 247. Distribution: Bismarck Arch, Borneo, New Guinea, Philippines, Solomon Isls, Sulawesi, Sumatra. BOHOL, CAMIGUIN, LUZON: Bataan, Benguet, Camarines Norte, Ilocos Norte, Laguna, Mountain Province, Pampanga, Quezon, Rizal, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Davao, Davao del Sur, Lanao, Lanao del Sur, Sarangani, South Cotabato, Zamboanga del Sur, MINDORO, NEGROS, PANAY (photos), SAMAR. Notes: 350-1900m. Native.
  2. Davallodes hymenophylloides (Blume) M.Kato & Tsutsumi, Acta Phytotax. Geobot. 59 (2008) 12; Pescuela et al., PJS 153 (2024) 664; ≡ Davallia hymenophylloides (Blume) Kuhn, Ann. Mus. Bot. Lugduno-Batavi 4 (1869) 286, nom. illeg.; Nooteboom, Fl. Males. ser. 2, 3 (1998) 255; ≡ Araiostegia hymenophylloides (Blume) Copel., PJS 34 (1927) 241; ≡ Aspidium hymenophylloides Blume, Enum. Pl. Javae (1828) 172. Distribution: Borneo, India, Java, Laos, Lesser Sunda Isls, Malay Peninsula, Philippines, Sri Lanka, Sulawesi, Sumatra, Thailand. BILIRAN, CAMIGUIN, LUZON: Apayao (photos), Aurora, Bataan, Batangas, Benguet, Cagayan, Camarines Norte, Camarines Sur, Ifugao, Laguna, Mountain Province, Nueva Vizcaya, Quezon, Rizal, Sorsogon, Zambales, MINDANAO: Agusan del Norte, Bukidnon, Cotabato, Davao Oriental (photos), Davao del Sur, Lanao, Misamis Oriental, Sarangani, MINDORO. Notes: 500-2200m. Native.

References