Amaranthaceae

including Chenopodiaceae

Achyranthes L.

Last edited by Pieter B. Pelser, 18 May 2021

Photos
  1. Achyranthes aspera L., Sp. Pl. (1753) 204; --Merr., Fl. Manila (1912) 195; EPFP 2 (1923) 131; --Backer, Fl. Males. ser. 1, 4 (1949) 88. Afghanistan, Africa, Andaman Isls, Australia, Bangladesh, Bismarck Arch, Borneo, Cambodia, China, India, Indian Ocean, Java, Laos, Lesser Sunda Isls, Middle East, Moluccas, Myanmar, Pacific Ocean, Peninsular Malaysia, Nepal, New Guinea, Nicobar Isls, Pakistan, Ryukyu Isls, Solomon Isls, Sri Lanka,Sulawesi, Sumatra, Taiwan, Thailand, Vietnam, Philippines. Throughout the Philippines. LUZON: Pampanga. Not native to the Philippines. Weed in open waste places; probably introduced in the Philippines (Merrill 1923). Photos
  2. Achyranthes bidentata Blume, Bijdr. (1825) 545; --Merr., EPFP 2 (1923) 131; --Backer, Fl. Males. ser. 1, 4 (1949) 88. Bangladesh, Bonin Isls, Borneo, Cambodia, China, India, Japan, Java, Korea, Laos, Lesser Sunda Isls, Moluccas, Myanmar, Nepal, New Guinea, Nicobar Isls, Pakistan, Peninsular Malaysia, Ryukyu Isls, Solomon Isls, Sri Lanka, Sulawesi, Sumatra, Taiwan, Thailand, Tibet, Vietnam, Philippines.

Aerua = Aerva

Aerva Forssk.

Last edited by Pieter B. Pelser, 20 May 2018

Photos 
  1. Aerva lanata (L.) Juss., Ann. Mus. Paris 11 (1808) 131; --Merr., Fl. Manila (1912) 194; EPFP 2 (1923) 130; --Backer, Fl. Males. ser. 1, 4 (1949) 84; --Achyranthes lanata L., Sp. Pl. (1753) 204. Africa, Middle East, Australia, Madagascar, Java, Nepal, India, Sri Lanka, Sumatra, Moluccas, Lesser Sunda Isl (Timor), New Guinea, Cocos (Keeling) Isl., Nicobars, Pakistan, Vietnam, Lakshadweep Isl (Laccadives), Maldives, Philippines. Photos
  2. Aerva sanguinolenta (L.) Blume, Bijdr. (1825) 547; --Backer, Fl. Males. ser. 1, 4 (1949) 85; --Achyranthes sanguinolenta L., Sp. Pl. ed. 2 (1762) 294. --Aerva scandens Wall., Cat. (1829) 6911, nomen; --Moq. in DC., Prodr. 13, 2 (1849) 302; --Merr., EPFP 2 (1923) 130. China, Taiwan, Bhutan, Cambodia, India, Laos, Malaysia, Nepal, Sikkim, Vietnam, Java, Sulawesi, Moluccas (Ambon, Tanimbar), Lesser Sunda Isl. (Lombok, Sumbawa, Flores, Timor), Thailand, Myanmar, Pakistan, Pakistani Kashmir, Jammu & Kashmir, Philippines.  Photos

Allmania R.Br. ex Wight

Last edited by Pieter B. Pelser, 20 May 2018

No Photos
  1. Allmania nodiflora (L.) R.Br. ex Wight in Hook., London J. Bot. 1 (1834) 226; --R.Br. in Wall., Cat. (1832) 6890, nom. nud.; --Backer, Fl. Males. ser. 1, 4 (1949) 74; --Celosia nodiflora L., Sp. Pl. (1753) 205. --Allmania albida R.Br. in Wall., Cat. (1832) No. 6981, nom. nud.; --Merr., EPFP 2 (1923) 127. China, India, Sikkim, Vietnam, Java, Sri Lanka, Laos, Cambodia, Peninsular Malaysia, Singapore, Sumatra, Lesser Sunda Isl. (Sumba, Wetar), Thailand, Lakshadweep Isl. (Laccadives), Philippines.

Alternanthera Forssk.

Last edited by Pieter B. Pelser, 15 January 2021

Photos
 
  1. Alternanthera bettzickiana (Regel) G.Nicholson, Illust. Dict. Gard. 1 (1884) 59, as ‘bettzchiana’; --Veldk., Blumea 19 (1971) 169; --Telanthera bettzickiana Regel, Index Seminum (LE) 1862 (1862) 28. --Alternanthera ficoidea (L.) R.Br. ex Roemer & Schultes, Syst. 5 (1819) 555, nom. illeg., auct. non P.Beauv.; --Moody, Weeds reported in rice in South and Southeast Asia (1989) 273; --Gomphrena ficoidea L., Sp. Pl. (1753) 225. --Alternanthera ficoides var. versicolor (Regel) Backer, Fl. Males. ser. 1, 4 (1949) 93; --Alternanthera versicolor Regel, Gartenfl. (1869) 101; --Merr., Fl. Manila (1912) 192; EPFP 2 (1923) 132. Native of Brazil. Photos
  2. Alternanthera brasiliana (L.) Kuntze, Revis. Gen. Pl. 2 (1891) 537; --Backer, Fl. Males. ser. 1, 4 (1949) 94; --Gomphrena brasiliana L., Amoen. Acad. 4 (1759) 10. --Alternanthera dentata (Moench) Scheygr., Fl. Suriname 1 (1932) 39; --Gomphrena dentata Moench, Suppl. Meth. (1802) 273. Native to tropical America. Photos
  3. Alternanthera paronychioides A.St.-Hil., Voy. Brésil 2, 2 (1833) 43; --Veldk., Blumea 19 (1971) 167; --Bakh.f. & Veldk., Addenda & Corr., Fl. Males. ser. 1, 6 (1972) 916.
  4. Alternanthera sessilis (L.) DC., Cat. Hort. Monsp. (1813) 77; --R.Br. ex Sweet, Hort. Suburb. Lond. (1818) 48; --Roth in Roemer & Schultes, Syst. 5 (1819) 554; --Merr., Fl. Manila (1912) 191; EPFP 2 (1923) 132; --Backer, Fl. Males. ser. 1, 4 (1949) 92. Native of Old World. Photos

Possibly present:

  1. Alternanthera philoxeroides (Mart.) Griseb. in Goett, Abh. 24 (1879) 36; --Backer, Fl. Males. ser. 1 (1949) 93; --Bucholzia philoxeroides Mart., Beitr. Amarantac. (1825) 107. Native of Brazil. Note by P.B. Pelser (15-Jan-21): Reported from Java and Singapore, and listed for the Philippines in L.L. Co's original documents that formed the foundation for this checklist, but I have not seen any specimens.

Amaranthus L.

Last edited by Pieter B. Pelser, 24 May 2021

Photos 
  1. Amaranthus blitum L., Sp. Pl. (1753) 990. Native to the Mediterranean region.
  2. Amaranthus caudatus L., Sp. Pl. (1753) 990; --Backer, Fl. Males. ser. 1, 4 (1949) 80. Native to American tropics. MINDORO. Photos
  3. Amaranthus cruentus L., Syst. Nat. ed. 10 2 (1759) 1269; --Amaranthus hybridus L. ssp. cruentus (L.) Thell., Fl. Adv. Montpellier (1912) 205. --Amaranthus paniculatus L., Sp. Pl. ed. 2 (1763) 1406; --Merr., EPFP 2 (1923) 128; --Amaranthus hybridus ssp. cruentus var. paniculatus (L.) Thell. in Asch. & Gr. Syn. 5, 1 (1914) 247; --Backer, Fl. Males. ser. 1, 4 (1949) 79. --Amaranthus caudatus auct. non L.; --Merr., Fl. Manila (1912) 193. Native to the Americas. LUZON to MINDANAO. Naturalized in open waste places. Photos
  4. Amaranthus dubius Mart. ex Thell., Fl. Adv. Montpellier (1912) 203. Not native. Invasive species (CABI 2017).
  5. Amaranthus spinosus L., Sp. Pl. (1753) 991; --Merr., Fl. Manila (1912) 192; EPFP 2 (1923) 128; --Backer, Fl. Males. ser. 1, 4 (1949) 78. Pantropic ubiquist. Throughout Malesia and the Philippines where it is probably introduced. Weed in open waste lands, roadsides, fields, gardens, etc., gregarious and abundant along sand bars and margins of streams, 0-1400m. Photos .
  6. Amaranthus tricolor L., Sp. Pl. (1753) 989; --Merr., EPFP 2 (1923) 128; --Backer, Fl. Males. ser. 1, 4 (1949) 77. Native to South America. LUZON to MINDANAO. Naturalized in waste places.
  7. Amaranthus viridis L., Sp. Pl. ed. 2 (1763) 1405; --Merr., Fl. Manila (1912) 193; EPFP 2 (1923) 128. --Amaranthus gracilis Desf., Tabl. Enc. Bot. (1804) 43; --Backer, Fl.Males. ser. 1, 4 (1949) 76. Native to S & C America. Not native and naturalized in the Philippines. Waste places and cultivated grounds (Merrill 1912). Photos

Possibly present:

  1. Amaranthus hybridus L., Sp. Pl. (1753) 990. --Amaranthus hybridus ssp. incurvatus (Trimen ex Gren. & Gord.) Brenan, Watsonia 4 (1961) 268; --Amaranthus incurvatus Trimen ex Gren. & Gord., Fl. France Prosp. (1846) 8. Native to North America. Note by P.B. Pelser (15-Jan-21): Listed for the Philippines in L.L. Co's original documents that formed the foundation for this checklist, but I have not seen any specimens. Photos

Blutaparon Raf. [Not native]

Last edited by Pieter B. Pelser & Rex Sadaba, 26 April 2018

Photos 
  1. Blutaparon vermiculare (L.) Mears, Taxon 31 (1982) 113; --Gomphrena vermicularis L., Sp. Pl. 1 (1753) 224. Native to the Americas and Africa, introduced in Japan and the Philippines. CEBU (photos by Rex Sadaba). Coastal saline and mangrove habitats. Photos

Celosia L. [Not native]

Last edited by Pieter B. Pelser, 10 August 2019

Photos 
  1. Celosia argentea L., Sp. Pl. (1753) 205; --Merr., Fl. Manila (1912) 190; EPFP 2 (1923) 127; --Backer, Fl. Males. ser. 1, 4 (1949) 73. Pantropic ubiquitous weed, wild but non-native, possibly introduced from tropical Africa. Throughout the Philippines at low and medium elevation in the settled areas. Very common along sandy-gravelly river banks, open dry roadsides, fields, gardens, etc. Photos
  2. Celosia cristata L., Sp. Pl. (1753) 205; --Merr., EPFP 2 (1923) 127; --Celosia argentea var. cristata (L.) Kuntze, Revis. Gen. Pl. 1 (1891) 541. Pantropic in cultivation for ornamental purposes. Also sometimes running wild but not far from human habitation.

Chenopodium L.

Edited by Pieter B. Pelser, 26 June 2012

Photos
 
  1. Chenopodium acuminatum Willd., Ges. Nat. Fr. Neue Schr. 2 (1799) 124; --Merr., EPFP 2 (1923) 125; --Backer, Fl. Males. ser. 1, 4 (1949) 103. Siberia, China, Japan, Taiwan, N Philippines. BATAN. In waste places.
  2. Chenopodium album L., Sp. Pl. (1753) 219; --Backer, Fl. Males. ser. 1, 4 (1949) 103. Europe, Continental Asia, Africa, Australia, America. Introduced in Java and naturalized in the E part of the island, N Philippines. LUZON: Benguet. A casual weed in temperate vegetable gardens, c. 1300-1400m. Photos .

Cyathula Blume

Photos 
 
  1. Cyathula prostrata (L.) Blume, Bijdr. (1825) 549; --Merr., Fl. Manila (1912) 194; EPFP 2 (1923) 129; --Backer, Fl. Males. ser. 1, 4 (1949) 82. Photos

Deeringia R.Br.

Last edited by Pieter B. Pelser, 20 May 2018

Photos
  1. Deeringia amaranthoides (Lam.) Merr., Interp. Herb. Amb. (1917) 211; --Merr., EPFP 2 (1923) 126; --Backer, Fl. Males. ser. 1, 4 (1949) 71; --Achyranthes amaranthoides Lam., Encycl. 1 (1785) 548. Deeringia baccata Moq. in DC., Prodr. 132 (1849) 236; --Merr., Fl. Manila (1912) 190. India, Laos, Thailand, Vietnam, Cambodia, Australia, Java, Peninsular Malaysia, Sumatra, Sulawesi, Lesser Sunda Isl, New Guinea, Fiji, Solomons, Christmas Isl, Marianas, Tonga, China, Taiwan, Tibet, Ryukyu Isl, Bhutan, Indonesia, Nepal, Sikkim, Myanmar, Pakistani Kashmir, Philippines. Photos .
  2. Deeringia polysperma (Roxb.) Moq. in DC., Prodr. 132 (1849) 236; --Merr., EPFP 2 (1923) 126; --Backer, Fl. Males. ser. 1, 4 (1949) 72; --Celosia polysperma Roxb., Fl. Ind. 2 (1824) 511. --Deeringia polysperma var. pubescens (Schinz) Merr., EPFP 2 (1923) 126; --Deeringia indica var. pubescens Schinz. Sumatra, Peninsular Malaysia, Java, Philippines, Sulawesi, Kabaena, Moluccas, New Guinea. BATANES to MINDANAO and PALAWAN. Thickets, light forests, forest edges, low to medium elevation, ascending to 1800m in Benguet. Much rarer than Deeringia amaranthoides. Photos.

Dysphania R.Br. [Not native]

Last edited by Pieter B. Pelser, 6 July 2017

No Photos

  1. Dysphania ambrosioides (L.) Mosyakin & Clemants, Ukrayins'k. Bot. Zhurn. 59 (2002) 382; --Fuentes-Bazan et al. 2012; --Chenopodium ambrosioides L., Sp. Pl. (1753) 219; --Merr., Fl. Manila (1912) 189; EPFP 2 (1923) 125; --Backer, Fl. Males. ser. 1, 4 (1949) 101. Native of the neotropics, introduced in many other regions: Europe, Asia, Africa, Australia. In Malesia naturalized in Java, N Sulawesi, E New Guinea as well as ‘throughout’ the Philippines (although now seem to be common only in Benguet prov. of N LUZON, c. 1400m). Invasive species (CABI 2017).

Gomphrena L. [Not native]

Photos
  1. Gomphrena celosioides Mart., Beitr. Amar. (1825) 93. Native of tropical America, introduced in tropical and S Africa, India and Australia. Now widespread in Malesia. Throughout the Philippines. Gregarious mat-forming weed along roadsides, abandoned fields, wastelands, and other similar hot, dry and dusty sites at low elevation. Photos
  2. Gomphrena globosa L., Sp. Pl. (1753) 224; --Merr., Fl. Manila (1912) 191; EPFP 2 (1923) 132; --Backer, Fl. Males. ser. 1, 4 (1949) 95. Native of tropical America, now pantropic. Introduced long ago in Malesia, either cultivated or semi-naturalized. Throughout the settled areas of the Philippines in cultivation, often occurring as an escape. Lowlands to c.1300m. Photos

 Iresine P.Browne [Not native]

Last edited by Pieter B. Pelser, 8 September 2019

Photos
  1. Iresine diffusa Humb. & Bonpl. ex Willd., Sp. Pl., ed. 4, 4 (1806) 765.

Lipandra Moq. [Not native]

Last edited by Pieter B. Pelser, 26 June 2012

No Photos

  1. Lipandra polysperma (L.) S.Fuentes, Uotila & Borsch, Willdenowia 42 (2012) 14; --Chenopodium polyspermum L., Sp. Pl. (1753) 220; --Merr., EPFP 2 (1923) 126; --Backer, Fl. Males. ser. 1, 4 (1949) 101. Europe, Continental Asia; elsewhere locally introduced. Merrill (1923) records the plant as a casual weed in the Philippines.

Psilotrichum Blume

Last edited by Pieter B. Pelser, 20 May 2018

Photos
  1. Psilotrichum ferrugineum (Roxb.) Moq. in DC., Prodr. 13, 2 (1849) 279; --Backer, Fl. Males. ser. 1, 4 (1949) 90; --Achyranthes ferruginea Roxb., Fl. Ind. 2 (1824) 502. --Psilotrichum trichotomum Blume, Bijdr. (1825) 545; --Merr., EPFP 2 (1923) 130. China, India, Cambodia, Thailand, Vietnam, Java, Peninsular Malaysia, Sumatra, Borneo, Sulawesi, New Guinea, Palau Isl., Myanmar, Nepal, Laos, Philippines.

Pupalia Juss.

Last edited by Pieter B. Pelser, 20 May 2018

Photos
  1. Pupalia lappacea (L.) Juss., Ann. Mus. Paris 2 (1803) 132; --Merr., EPFP 2 (1923) 129; --Backer, Fl. Males. ser. 1, 4 (1949) 83; --Achyranthes lappacea L., Sp. Pl. (1753) 204. --Pupalia atropurpurea (Lam.) Moq. in DC., Prodr. 132 (1849) 331; --Merr., Fl. Manila (1912) 193; --Achyranthes atropurpurea Lam., Encycl. 1 (1785) 546. Africa, Middle East, Java, India, Sulawesi, Lesser Sunda Isls (Bali, Lombok, Sumba, Sumbawa), New Guinea, Sri Lanka, Afghanistan, Pakistan, Pakistani Kashmir, Jammu & Kashmir, Myanmar, Philippines.

  Possibly present:

  1. Digera muricata (L.) Mart., Beitr. Amar. (1825) 77; --Backer, Fl. Males. ser. 1, 4 (1949) 80; --Achyranthes muricata L., Sp. Pl. ed. 2 (1762) 295. Africa, Middle East, Java, Sulawesi, Moluccas, Lesser Sunda Isls (Sumba, Sumbawa), India, Lakshadweep Isl, Indochina, Sri Lanka, Afghanistan, Pakistan, Jammu & Kashmir, Myanmar, ?Philippines. Note by P.B. Pelser (15-Mar-20): included for the Philippines in the original, unpublished version of this checklist by L.L. Co, but it is not clear to me what the source of this information is.

Literature

Backer, CA. 1949. Amaranthaceae. Flora Malesiana series 1, 4: 69-98; idem, 1, 6 (1972) 915-917

Backer, CA. 1949. Chenopodiaceae. Flora Malesiana series 1, 4: 99-106


CABI Invasive Species Compendium

Chen, S-H & Y-C Ling. 2012. Remarks on the Species of Gomphrena (Amaranthaceae) of Taiwan. Taiwania 57: 312-317

Fuentes-Bazan, S, P Uotila & T Borsch. 2012. A novel phylogeny-based generic classification for Chenopodium sensu lato, and a tribal rearrangement of Chenopodioideae (Chenopodiaceae). Willdenowia 42: 5-24

Moody, K. 1989. Weeds reported in rice in South and Southeast Asia. IRRI, Philippines

Pedersen, TM. 2000. Studies in South American Amaranthaceae V. Bonplandia 10: 83-112.

Veldkamp, JF. 1971. Alternanthera paronychioides St Hil. (Amaranthaceae) in Indo-Malesia. Blumea 19: 167-169


Pelser, P.B., J.F. Barcelona & D.L. Nickrent (eds.). 2011 onwards. Co's Digital Flora of the Philippines. www.philippineplants.org

Copyright © 2011, Co's Digital Flora of the Philippines

Last updated 24 May 2021