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Thymelaeaceae

Including Gonostylaceae

Aquilaria Lam.

Photos
  1. Aquilaria apiculata Merr., PJS 20 (1922) 411; EPFP 3 (1923) 130; --Quisumb., J. Arnold Arb. 27   (1946) 403; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 11. MINDANAO: Bukidnon. Dry and mossy forests, 1100-1800m.
  2. Aquilaria brachyantha (Merr.) Hallier f., Med.Rijksherb. no. 44 (1922) 16; --Merr., EPFP 4 (1925) 244; --Quisumb., J. Arnold Arb. 27 (1946) 403; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 10, Fig. 1d; --Gyrinopsis brachyantha Merr., PJS 7 c (1912) Bot. 313; EPFP 3 (1923) 130. LUZON: Cagayan. Note by L.L.Co: Twice collected. Low altitude primary forest.
  3. Aquilaria citrinaecarpa (Elmer) Hallier f., Meded. Rijks Herb. 44 (1922) 18; --Gyrinopsis citrinaecarpa Elmer, LPB 5 (1913) 1631; --Merr., EPFP 3 (1923) 130; --Quisumb., J. Arnold Arb. 27 (1946) 405; --Type: Elmer 13566 (L, NY, U, US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta. Wooded ridges of Duros Peak, 4000 ft., Aug-1912. MINDANAO: Agusan del Norte (Mt Urdaneta). Note by L.L.Co: Known only from the type collection. Moist compact soil of forested ridges, c. 1300m.
  4. Aquilaria cumingiana (Decne.) Ridley, J. Str. Br. R. As. Soc, no. 30 (1901) 80; --Ding Hou, Fl. Males. ser. i, 6 (1960) 15, Fig. 1i; --Gyrinopsis cumingiana Decne., Ann. Sc. Nat. ii, 19 (1843) 41; --Merr., EPFP 3 (1923) 131; 4 (1925) 244;  --Quisumb., J. Arnold Arb. 27 (1946) 405; --Type: Cuming 1617 (NY), Mindanao: Precise locality not indicated. --Gyrinopsis cumingiana var. pubescens Elmer, LPB 5 (1913) 1629; --Type: Elmer 10981 (L, NY, US, iso), Mindanao: Davao, Mt Apo, Todaya. Along ridge of Baracatan Creek, 1750 ft. Jun-1909. --Gyrinopsis decemcostata Hallier f., Meded. Rijks Herb. 44 (1922) 17; --Type: BS 1114 Ramos (L,   lecto), Luzon: Laguna prov., Dahican River, Sep-1912. --Gyrinopsis pubifolia Quisumb., J. Arnold Arb. 27 (1946) 406; --Type: BS 75314 (A). Catanduanes: Mt   Abucay, 1600m (Note by L.L.Co: altitude most likely in error as no part of Catanduanes reach this altitude), 11-Sep-1928. S Borneo (Sampit region), Philippines (common), and Moluccas (Morotai and Halmahera). Primary forests at low and medium altitudes. Photos
  5. Aquilaria filaria (Oken) Merr., J. Arn. Arb. 31 (1950) 283, excl. syn. Gyrinopsis brachyantha Merr.; --Ding Hou, Fl. Males. Ser. 1, 6 (1960) 11, Fig. 2 (map). --Gyrinopsis acuminata Merr., PJS 17 (1920) 294; EPFP 3 (1923) 130; --Quisumb., J. Arnold Arb. 27 (1946) 403; --Type: BS 35158 Ramos & Pascasio (US, isotype). Dinagat. May 1919. Philippines, Moluccas (Morotai, Seram, Ambon), New Guinea (Sorong, Babo and Kapor). DINAGAT, BUCAS GRANDE.
  6. Aquilaria malaccensis Lam., Encycl. 1 (1783) 49; --Merr., EPFP 3 (1923) 130; --Quisumb., J. Arnold   Arb. 27 (1946) 403; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 9, Fig. 1h. India (Assam), Bangladesh, Tenasserim, Sumatra (Simalur, Sibolangit, Palembang, and Banka), Peninsular Malaysia, N & E Borneo, Philippines. LUZON.
  7. Aquilaria parvifolia (Quisumb.) Ding Hou, Fl. Males. ser. 1, 6 (1960) 12; --Gyrinopsis parvifolia Quisumb., J. Arnold Arb. 27 (1946) 405; --Type: BS 76441 Edaño (A, holo; L, type fragment), Luzon: Camarines Sur prov., Her-it River, Dec-1928. LUZON: Camarines Sur. Note by L.L.Co: A total of four specimens cited by Ding Hou, including Wenzel 1201 (possibly not from Luzon but Leyte or NE Mindanao). Forested slopes, 1000m.
  8. Aquilaria urdanetensis (Elmer) Hallier f., Med. Rijksherb. no. 44 (1922) 16; --Ding Hou, Fl. Males.   ser. 1, 6 (1960) 10; --Gyrinopsis urdanetensis.Elmer, LPB 5 (1913) 1630; --Merr., EPFP 3 (1923) 131; --Quisumb., J. Arnold Arb. 27 (1946) 405; --Type: Elmer 14195 (L, NY, U, US, isolecto), Mindanao: Agusan del Norte prov., Mt Urdaneta. Damp mossy forest, 5250 ft., Oct-1912. MINDANAO: Agusan del Norte (Mt Urdaneta). Mossy forest on exposed ridges, c. 1700m. Note by L.L.Co: Twice collected.

Daphne L.

Photos
  1. Daphne luzonica C.B.Rob., Bull. Torr. Bot. Club 35 (1908) 72, 75; --Merr., EPFP 3 (1923) 132; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 35, Fig. 14a-b. LUZON: Benguet. Mossy forests on the higher mountains, 2000-2500m. Photos

Enkleia Griff.

No Photos
  1. Enkleia paniculata (Merr.) Hallier f., Meded. Rijks Herb. 44 (1922) 26; --Merr., EPFP 4 (1925) 244; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 25, Fig. 10h-I; --Macgregorianthus paniculatus Merr., PJS 7 c (1912) Bot. 312; EPFP 3 (1923) 132. Philippines, New Guinea. LUZON.

Gonystylus Teijsm. & Binn.

Photos
  1. Gonystylus macrophyllus (Miq.) Airy Shaw, Kew Bull. (1947) 9; Fl. Males. ser. 1, 4 (1953) 354. --Gonystylus bancanus (Miq.) Gilg in Engl. & Prantl, Nat. Pflanzenfam. Nachtr. 1 (1897) 232; --Merr., EPFP 3 (1923) 21. --Gonystylus ovatus Merr., PJS 12 c (1917) Bot. 283; EPFP 3 (1923) 21; --Type: FB 26713 Peñas. --Gonystylus philippinensis Elmer, LPB 7 (1915) 2674; --Merr., EPFP 3 (1923) 21; --Type: Elmer 12520. Sumatra, Banka, Java, Philippines. BABUYAN ISLS (CALAYAN), LUZON: Cagayan, Isabela, Nueva Ecija, Bataan, Laguna, Quezon, Camarines, Sorsogon, MINDORO, SIBUYAN, SAMAR, MINDANAO: Surigao. Primary forests from low altitudes to 1500m. 
  2. Gonystylus reticulatus (Elmer) Merr., PJS 12 c (1917) Bot. 284; EPFP 3 (1923) 21; --Airy Shaw, Fl. Males. ser. 1, 4 (1953) 354; --Thea reticulata Elmer, LPB 8 (1915) 2838; --Type: Elmer 13478. MINDANAO: Zamboanga, Agusan del Norte. Primary forests at low and medium altitudes, ascending to 900m.

 Gyrinopsis Decne. =Aquilaria

Macgregorianthus Merr. =Enkeleia

Phaleria Jack

Photos
  1. Phaleria capitata Jack, Mal. Misc. (1822) 59; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 20, Fig. 3a-g, 6. --Phaleria cumingii (Meisn.) Fernandez-Villar, Novis. App. (1880) 183; --Merr., EPFP 3 (1923) 131. Sri Lanka, Sumatra, Peninsular Malaysia, Borneo, Philippines, Java, Sulawesi, Moluccas (Buru), New Guinea (Waigeo Is), Palau. Photos
  2. Phaleria coccinea (Gaudich.) F Muell., Desc. Not. 2 (1885) 9; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 21. --Phaleria platyphylla Merr., PJS 14 (1919) 429; EPFP 3 (1923) 131. Philippines, Moluccas (Sula; Ambon; Seram & Kei), New Guinea, New Britain. PANAY, MINDANAO.
  3. Phaleria perrottetiana (Decne.) Fernandez-Villar, Novis. App. (1880) 183; --Merr., EPFP 3 (1923) 131; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 18, Fig. 4 (map).  N Borneo (Banguey Is & Lahat Datu), Philippines, Moluccas (Kei; Seram), New Guinea, Louisiade Archipelago (Sudest Is).

Pimelea Banks & Sol.

Photos
  1. Pimelea cornucopiae Vahl, En. Pl. 1 (1804) 305; --Ding Hou, Fl.  Males. ser. 1, 6 (1960) 46, Fig. 19a-g. --Pimelea philippinensis C.B.Rob., PJS 6 c (1911) Bot. 345; --Merr., EPFP 3 (1923) 134. Philippines, New Guinea (Western & Central Divisions), Australia (Queensland), D’Entrecastaux Isls (Ferguson Is), New Britain, Louisiade Archieplago. LUZON: Cagayan (near Sanches-Mira). Note by L.L.Co: once collected. 

Wikstroemia Endl.

Edited by James V. LaFrankie & Pieter B. Pelser, 23 August 2016

Photos
  1. Wikstroemia androsaemifolia Decne., Ann. Sci. Nat., Bot., sér. 2, 20 (1843) 50. Indonesia, New Guinea, Philippines. LUZON: Bulacan, PANAY: Antique. Photos
  2. Wikstroemia brachyantha Merr., PJS 13 c (1918) Bot. 313; EPFP 3 (1923) 132; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 31, Fig. 12a-b. Borneo (Mt Kinabalu), Philippines. LUZON, CATANDUANES. Primary forests at low and medium elevation. Photos
  3. Wikstroemia indica (L.) C.A.Mey., Bull. Acad. Imp. Sc. St. Petersb. 1 (1843) 357; --Merr., EPFP 3 (1923) 132; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 34. --Wikstroemia linearifolia Elmer, LPB 2 (1910) 680; --Merr., EPFP 3 (1923) 133. --Wikstroemia ovata Vidal, Sinopsis (1883) 229, non C.A.Mey.. --Wikstroemia pachyphylla Merr., PJS 12 c (1917) Bot. 297; EPFP 3 (1923) 133. --Wikstroemia pulgarensis Elmer, LPB 5 (1913) 1844; --Merr., EPFP 3 (1923) 133. India, SE Asia, through Malesia (absent in the seasonal parts such as the Lesser Sunda Isls) to Australia (N Australia, Queensland, New South Wales) and Melanesia (as far east as Fiji). In thickets and secondary growth, also occurring on sand as well as bluffs along the coast, on limestone, ultramafics, and marginal soils. Photos
  4. Wikstroemia lanceolata Merr. Govt. Lab. Publ. (Philipp.) 29 (1905) 31; --Merr., EPFP 3 (1923) 133; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 30. --Wikstroemia angustissima Merr., PJS 7 c (1912) Bot. 92; --Merr., EPFP 3 (1923) 132. LUZON, MINDORO, PALAWAN. Common on forested slopes at low and medium elevation, up to 1300m. Photos
  5. Wikstroemia meyeniana Warb. in Perkins, Fragm. Fl. Philipp. (1905) 171; --Merr., EPFP 3 (1923) 133; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 33. --Wikstroemia fenicis Merr., PJS 13 c (1918) Bot. 312; EPFP 3 (1923) 132. Indochina, Philippines. LUZON to MINDANAO. Primary humid forests at low and medium elevation, up to c. 400m. Photos
  6. Wikstroemia ovata C.A.Mey., Acad. Imp. Sc. St. Petersb. 1 (1843) 357; --Merr., EPFP 3 (1923) 133; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 31. Borneo (Sabah, small isls off E coast), Philippines. LUZON, MINDORO, PALAWAN, NEGROS, MINDANAO. Thickets, primary and secondary forests at low and medium elevation up to 800m. Photos
  7. Wikstroemia polyantha Merr., PJS 10 c (1915) Bot. 332; --Merr., EPFP 3 (1923) 133; --Ding Hou, Fl. Males. ser. 1, 6 (1960) 32. Peninsular Malaysia, Java (W part), Borneo (Sabah), Philippines. LUZON. Photos
  8. Wikstroemia retusa A.Gray, J. Bot. Brit & For. 3 (1865) 303; Hatusima, Mem. Fac. Agr. Kagoshima Univ. 5 (1966) 41. --Wikstroemia indica sensu Merr., EPFP 3 (1923) 132, non C.A.Mey., as to BATAN plants only. Ryukyus, Lanyu (?), Philippines: BATAN. Rocky littoral meadow, Hatusima 28482, 28754. Locally frequent. Photos
  9. Wikstroemia tenuiramis Miq., Fl. Ned. Ind., Eerste Bijv. 3 (1861) 354. Indonesia, Philippines. LUZON: Albay.

Literature

Airy Shaw, HK. 1953. Thymelaeaceae-Gonystyloideae. Flora Malesiana series 1, 4: 349-365

Ding Hou. 1960. Thymelaeaceae. Flora Malesiana series 1, 6: 1-48. Gonystylus (Gonystylaceae) is treated separately (q.v.).

Ding Hou. 1964. Notes on some Asiatic species of Aquilaria (Thymelaeaceae). Blumea 12: 285-288

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Last updated 24 May 2017