Aspleniaceae

Asplenium L.

Edited by Julie F. Barcelona, 8 February 2013

1. Asplenium affine Sw., Schrad. J. Bot. 1800 (2) (1801) 56. --Asplenium spathulinum J Sm. ex Hook. --Asplenium gracilifolium Copel. Photos
2. Asplenium amboinense Willd., Sp. Pl. 5 (1810) 303; 
3. Asplenium apoense Copel. Photos
4. Asplenium callipteris (Fée) E.Fourn. --Asplenium epiphyticum Copel. Photos
5. Asplenium caudatum G Forst., Prodr. (1786) 80. --? Asplenium chamissoanum C.Presl. Photos
6. Asplenium cheilosorum Kunze ex Mett., Aspl., 133, n. 104 (1859) t. 5, Fig. 12-13. Photos
7. Asplenium colubrinum Christ 
8. Asplenium crinicaule Hance
9. Asplenium cuneatum Lam. Photos
10. Asplenium cymbifolium Christ, Bull. Boiss. 2, 6 (1906) 999. Photos  
11. Asplenium elmeri Christ, PJS 2 c (1907) Bot. 164 Photos
12. Asplenium excisum C.Presl. --Asplenium obscurum Blume. Photos
13. Asplenium exiguum Bedd. Photos
14. Asplenium griseum Copel.
15. Asplenium haenkei Copel. Photos
16. Asplenium insiticium Brackenr.
17. Asplenium lepturus J Sm. ex C.Presl. --Asplenium contiguum Kaulf. 
18. Asplenium lobulatum Mett., Linnaea 36 (1869) 100. Photos
19. Asplenium longissimum Blume. --Asplenium acutiusculum Blume. --? Asplenium anisodontum C.Presl. Photos  
20. Asplenium mantalingahanum PM Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp. 32 (1980) 46, t. 3.
21. Asplenium nidus L., Sp. Pl. (1753) 1079. Photos
    • var. nidus
    • var. curtisorum (Christ) Holtt., Gard. Bull. Sing. 27 (1974) 148; --Asplenium curtisorum Christ, Ann. Jard. Bot. Buitenzorg II. 4 (1904) 39.
22. Asplenium nigrescens Blume, Enum. Pl. Javae 2 (1828) 180. --Asplenium militare Copel. Photos
23. Asplenium normale D Don, Prodr. Fl. Nepal. (1825) 7. Photos  
24. Asplenium ofeliae Salgado, Amer. Fern J. 93(4) (2003) 193. –-Type: Merrill Phil. Plts. 700. (US, holo; iso: PRC). Luzon: Benguet prov., May 1911. Photos  
25. Asplenium pellucidum Lam., Encycl. 2 (1786) 305. Photos  
26. Asplenium persicifolium J Sm. ex Mett., Aspl. 97 (1859) no. 32. --Asplenium tripinnatifidum Copel. Photos
27. Asplenium phyllitidis D Don
    
    • ssp. malesicum Holttum, Gard. Bull. Sing. 27 (1974) 153; --Asplenium nidus L. var. phyllitidis (D. Don) Alderw., Handb. Malayan Ferns Suppl. (1917) 282.  --Probably incl. Asplenium colubrinum. Photos
28. Asplenium polyodon G Forst., Prodr. (1786) 80; --Asplenium falcatum Lam. Photos  
29. Asplenium robustum Blume. --Asplenium laserpitiifolium Lam.
30. Asplenium scandens J Sm. ex Mett. Photos
31. Asplenium schizocarpum Copel.
32. Asplenium scolopendrioides J Sm. ex Hook., Icon. Pl. (1854) t. 930
33. Asplenium semivarians Viane & Reichst., Pteridol. New Millennium (2003) 78 (-81; fig. 3). Tamil Nadu (India, Indian Subcontinent, Asia-Tropical); Yunnan (China South-Central, China, Asia-Temperate); West-Central Tropical Africa (Africa); Manipur (Assam, Indian Subcontinent, Asia-Tropical); Philippines (Malesia, Asia-Tropical); Sri Lanka (Indian Subcontinent, Asia-Tropical); Uganda (East Tropical Africa, Africa); Zimbabwe (South Tropical Africa, Africa); --Type: Collector(s): Fraser-Jenkins 9063. Locality: Shevaroy Hills, NW of Yercaud, NE of Salem, c. 1550m. 1978-12-13. holotype GENT, isotype Herb. R.Viane 
34. Asplenium setisectum Blume, Enum. Pl. Jav. (1828) 187; 
35. Asplenium stantoni Copel.
36. Asplenium steerei Harrington 
37. Asplenium subnormale Copel. in Perkins, Fragm. Fl. Philip. (1905) 183, t. 4, Fig. B. Photos
38. Asplenium subresectum Copel. 
39. Asplenium tenerum G Forst., Prodr. (1786) 80. Photos  
40. Asplenium tenuifolium Don
41. Asplenium thunbergii Kunze, Linnaea 10 (1836) 517; --Asplenium decorum Kunze. Photos
42. Asplenium unilaterale Lam., Encycl. 2 (1786) 305; --Asplenium filipes Copel., PJS 3 c (1908) Bot. 304. Photos  
43. Asplenium vittiforme Cav., Descr. (1802) 255. Photos
44. Asplenium vulcanicum Blume, Enum. Pl. Jav. (1828) 176; Possibly conspecific with Asplenium salignum Blume
45. Asplenium yoshinagae Makino var. planicaule (Wall. ex Mett.) C Morton, Contr. US Nat’l. Herb. 38 (1973) 227; --Asplenium planicaule Wall. ex Mett., Abhandl. Senckenb. Naturf. Gessell. 3 (1859) 201, non Lowe (1858).

Athyriaceae

Edited by Julie F. Barcelona, 22 January 2013

Anisocampium Presl = Athyrium

Athyrium Roth

Edited by Pieter B. Pelser, 3 March 2012

1. Athyrium anisopterum Christ. Photos
2. Athyrium aristulatum Copel.
3. Athyrium brevipinnulum Copel. --Athyrium meansianum (Copel.) Alderw. --Athyrium philippinense Christ apud Copel.
4. Athyrium cumingianum (C.Presl.) Ching; --Anisocampium cumingianum C.Presl.
5. Athyrium drepanopteron (Kunze) A. Braun. Photos
6. Athyrium elmeri Copel. Photos
7. Athyrium oreopteris Copel.
8. Athyrium puncticaule (Blume) Moore. Photos
9. Athyrium ramosii Copel.
10. Athyrium stramineum Copel.

Callipteris Bory = Diplazium

Cornopteris Nakai

Edited by Pieter B. Pelser, 3 March 2012

1. Cornopteris banahaoensis (C Chr.) K Iwats. & MG Price. --Athyrium nudum Copel.
2. Cornopteris decurrenti-alata (Hook.) Nakai. Photos
3. Cornopteris opaca (D. Don) Tagawa. --Athyrium gymnocarpum Copel.

Deparia Hooker & Greville

Edited by Pieter B. Pelser, 3 March 2012

1. Deparia boryana (Willd.) M.Kato, Bot. Mag. Tokyo 90 (1977) 36; --Dryoathyrium boryanum (Willd.) Ching; --Ctenitis boryana (Willd.) Copel.
2. Deparia confluens (Kunze) M.Kato. Photos
3. Deparia formosana (Rosenst.) Sano, Act. Phytotax. Geobot. 51 (2000) 17; --Diplaziopsis formosana (Rosenst.) MG Price, Contrib. Univ. Mich. Herb. 17 (1990) 269; -- Diplazium formosanum Rosenst., Hedw. 56 (1915) 337.
4. Deparia japonica (Thunb.) M.Kato, Bot. Mag. Tokyo 90 (1977) 37; --Lunathyrium japonicum (Thunb.) Kurata; --Athyrium japonicum (Thunb.) Ching. --Athyrium grammitoides (C.Presl.) Milde. Photos
5. Deparia petersenii (Kunze) M.Kato. Photos
6. Deparia subfluviatilis (Hayata) M.Kato.

Diplazium Swartz

1. Diplazium accedens Blume Polillo, PNBG 184 Sotalbo (PUH). New record for the Philippines? Most specimens in D. proliferum; the rest in D. vestitum & D. dolichosorum. Photos
2. Diplazium allantoideum MG Price.
3. Diplazium atratum Christ; --Athyrium atratum (Christ) Copel.
4. Diplazium bantamense Blume; --Athyrium bantamense (Blume) Milde.
5. Diplazium bolsteri Copel.; --Athyrium bolsteri (Copel.) Copel.
6. Diplazium brachysoroides Copel.; --Athyrium brachysoroides (Copel.) Copel.
7. Diplazium calliphyllum (Copel.) MG Price, Contrib. Univ. Mich. Herb. 17 (1990) 270; --Athyrium calliphyllum Copel.
8. Diplazium cordifolium Blume; --Athyrium cordifolium (Blume) Copel. --Athyrium parvum Copel. Photos
9. Diplazium costulisorum (Copel.) C Chr.; --Athyrium costulisorum Copel.
10. Diplazium crenato-serratum (Blume) T.Moore, Index Fil. (T. Moore) 121 (1859) 325; --Athyrium crenato-serratum (Blume) Mildez. Photos
11. Diplazium cultratum C.Presl.; --Athyrium cultratum (C.Presl.) Milde. Photos
12. Diplazium cumingii (C.Presl.) C Chr.; --Athyrium cumingii (C.Presl.) Milde. Photos
13. Diplazium davaoense Copel.; --Athyrium davaoense (Copel.) Copel. Photos
14. Diplazium deltoideum (C.Presl.) C.Presl.; --Athyrium deltoideum (C.Presl.) Milde.
15. Diplazium dilatatum Blume; --Athyrium dilatatum (Blume) Milde. Photos
16. Diplazium doederleinii (Luerssen) Makino. --Athyrium irayense Copel.
17. Diplazium dolichosorum Copel.; --Athyrium dolichosorum (Copel.) Copel.
18. Diplazium egenolfioides MG Price, Brit. Fern Gaz. 10 (1973) 258, t. 1 LUZON: Mt. Makiling (several collections from this type locality); Nueva Vizcaya prov., Sta. Fe, mtns N of Imugan, c. 1200m, L.L.Co 1640 (MICH, PNH, PUH).
19. Diplazium esculentum (Retz.) Sw.; --Athyrium esculentum (Retz.) Copel. --Asplenium manilense Sprengel.
20. Diplazium forbesii (Baker) C.Chr., Index Filic. (1905) 232. --Diplazium pariens (Copel.) C Chr.; --Athyrium pariens Copel. Photos
21. Diplazium fraxinifolium C.Presl.; --Athyrium fraxinifolium (C.Presl.) Milde. Photos
22. Diplazium fructuosum Copel.; --Athyrium fructuosum (Copel.) Copel.
23. Diplazium fuliginosum (Hook.) MG Price, Brit. Fern Gaz. 10 (1973) 260; --Asplenium fuliginosum Hook. --Diplazium longissimum (Copel.) C Chr.; --Athyrium longissimum Copel.
24. Diplazium geophilum (Copel.) Alderw.; --Athyrium geophilum Copel.
25. Diplazium griffithii T Moore. --Athyrium platyphyllum Copel., 1908,  non Diplazium platyphyllum Christ, 1906. --Diplazium mearnsii Hieron. --Athyrium platyphyllum Copel. Photos
26. Diplazium irigense (Copel.) MG Price, Contrib. Univ. Mich. Herb. 17 (1990) 273; --Athyrium irigense Copel.
27. Diplazium lanceum (Thunb.) C.Presl. --Diplazium subsinuatum (Wall. ex Hook. & Grev.) Tagawa. --Athyrium dubium (Don) Copel.
28. Diplazium lomariaceum (Christ) MG Price, Gard. Bull. Sing. 36 (1983) 27. --Diplazium altum (Copel.) C Chr.; --Athyrium altum Copel. --Diplazium merrillii Copel.; --Athyrium merrillii (Copel.) Copel. Photos
29. Diplazium macrosorum (Copel.) MG Price, Contrib. Herb. Univ. Mich. 17 (1990) 273; --Athyrium macrosorum Copel.
30. Diplazium magnificum (Copel.) MG Price, 1982; --Athyrium magnificum Copel.
31. Diplazium melanopodium Fée; --Athyrium melanopodium (Fée) Copel. Photos
32. Diplazium mollifrons (C Chr.) MG Price
33. Diplazium muricatum (Mett.) Alderw.; --Athyrium muricatum (Mett.) Copel.
34. Diplazium oligosorum Copel.; --Athyrium oligosorum (Copel.) Copel.
35. Diplazium pallidum (Blume) Moore; --Athyrium pallidum Blume. Photos
36. Diplazium petiolare C.Presl.; --Athyrium petiolare (C.Presl.) Milde.
37. Diplazium pinatubicum MG Price, Contrib. Herb. Univ. Michigan 17 (1990) 273.
38. Diplazium polypodioides Blume. --Athyrium blumei (Bergsmann) Copel. --Diplazium ophiodontum (Copel.) C Chr.; --Athyrium ophiodontum Copel. --Diplazium asperum Blume; --Athyrium asperum (Blume) Milde.
39. Diplazium proliferum (Lam.) Thouars; --Callipteris prolifera (Lam.) Bory. --Diplazium accedens Blume.
40. Diplazium propinquum (Copel.) Alderw.; --Athyrium propinquum Copel.
41. Diplazium pseudocyatheifolium Rosenst.; --Athyrium pseudocyatheifolium (Rosenst.) Copel. --Athyrium ebenirachis Copel. Photos
42. Diplazium sibuyanense (Copel.) Alderw.; --Athyrium sibuyanense Copel. Photos
43. Diplazium sorzogonense (C.Presl.) C.Presl.; --Athyrium sorzogonense (C.Presl.) Milde. Photo
44. Diplazium subsinuatum (Hook. & Grev.) Tagawa, Col. Ill Jap. Pterid. (1959) 203, f. 298; --Price, Kalikasan 1 (1972) 42; --Asplenium subsinuatum Hook. & Grev., Icon. Fil. (1827) t. 27. -- Athyrium lanceum (Thunb.) Milde (1870, non Moore 1860); --Copel., Philip. J. Sci. 7C (1912) 55; --Diplazium lanceum (Thunb.) C. Presl, Tent. Pterid. (1836) 113 (non Bory 1833). -- Athyrium dubium (Don) Ohwi (1956, non Ching 1949); --Copel., Fern Fl. Philip. 3 (1961) 412 (non Diplazium dubium Link (1833)). Sri Lanka, India to China, Indochina, Taiwan to Japan, south to the Philippines (Batanes, Luzon and Panay) and Borneo.
45. Diplazium symmetricum (Copel.) MG Price, Gard. Bull. Sing. 36 (1983) 26.; --Athyrium symmetricum Copel. SAMAR: Mt. Cabayanan, Edano PNH 15148, lectotype (MICH).
46. Diplazium tenuifolium (Copel.) Lellinger, Am. Fern J. 74 (1954) 57; --Athyrium tenuifolium Copel. Formally published new combination?
47. Diplazium tomentosum Blume. --Diplazium vestitum C.Presl.; --Athyrium vestitum (C.Presl.) Milde.
48. Diplazium whitfordii Copel.; --Athyrium whitfordii (Copel.) Copel.
49. Diplazium williamsii Copel.; --Athyrium williamsii (Copel.) Copel. --Athyrium banahaoense Copel.; --Diplazium banahaoense (Copel.) C Chr.
50. Diplazium woodwardioides (C.Presl.) Holttum. --Diplazium sylvaticum (Bory) Sw.; --Synopsis Filicum (1867). --Athyrium sylvaticum (Blume) Milde.
51. Diplazium xiphophyllum (Baker) Copel.; --Athyrium xiphophyllum (Baker) Copel.

Dryoathyrium Ching = Deparia

Lunathyrium Koidzumi = Deparia

Blechnaceae

Blechnum L.

1. Blechnum egregium Copel. in Perkins, Fragm. Fl. Philip. (1905) 187. Photos
2. Blechnum finlaysonianum Hook. & Grev., Icon. Fil., t. (1831) 225. Photos
3. Blechnum fraseri (A. Cunningham) Luerss., Flora (1876) 292. Photos
4. Blechnum melanocaulon (Brack.) T.C.Chambers & P.A.Farrant, Blumea 46(2) (2001) 318. MINDANAO: North Cotabato prov. Photos
5. Blechnum orientale L., Sp. Pl. (1753) 1077. Photos
6. Blechnum patersonii (R Br.) Mett., Fil. Lips. (1856) 64, t. 4, Fig. 4-10.
7. Blechnum vestitum (Blume) Kuhn, Ann. Lugd.-Bat. 4 (1869) 284. --Blechnum capense (L.) Schlechtr. Photos
8. Blechnum vulcanicum (Blume) Kuhn, Ann. Lugd.-Bat. 4 (1869) 284. Photos

Brainea J Smith

1. Brainea insignis (Hook.) J Sm.

Stenochlaena J Smith

1. Stenochlaena areolaris (Harrington) Copel.
2. Stenochlaena cumingii Holttum. --Stenochlaena laurifolia C.Presl.
3. Stenochlaena milnei Underw. --Stenochlaena juglandifolia C.Presl. Photos
4. Stenochlaena palustris (NL Burm.) Bedd., Ferns Brit. Ind. Suppl. (1876) 26. Photos

Woodwardia Smith

1. Woodwardia orientalis Sw. var. prolifera (Hook. & Arn.) Hook.
2. Woodwardia unigemmata (Makino) Nakai. Photos

Cibotiaceae 

Cibotium Kaulfuss

1. Cibotium barometz (L.) J Sm. Assam to S China, W Malesia. LUZON: Mountain Province (near Bontoc). Known in the Philippines by a single collection, CAHUP 16635 Pancho.
2. Cibotium cumingii Kunze. Photos

Cyatheaceae

Cyathea Smith

1. Cyathea acuminata Copel.
2. Cyathea apoensis Copel. --Cyathea lobata Copel. 
3. Cyathea atropurpurea Copel.; --Gymnosphaera atropurpurea (Copel.) Copel.
4. Cyathea biliranensis Copel. 
5. Cyathea binuangensis Copel.
6. Cyathea callosa Christ. --Cyathea foxworthyi Copel.
7. Cyathea camiguinensis Copel. 
8. Cyathea caudata (J Sm.) Copel. --Cyathea arborescens Copel. --Cyathea brevipes Copel. --Cyathea dupaxensis Copel.
9. Cyathea christii Copel. 
10. Cyathea cinerea Copel.
11. Cyathea contaminans (Wall.) Copel., PJS 4 c (1909) Bot. 60; --Cyathea clementis Copel. New Guinea to India. Throughout the Philippines. Usually common in open places and light forests from near sea-level to 1300m. 
12. Cyathea curranii Copel.
13. Cyathea edanoi Copel. Photos  
14. Cyathea elmeri (Copel.) Copel. --Cyathea argyrolepis Copel. --Cyathea dimorphotricha Copel.
15. Cyathea fenicis Copel.
16. Cyathea ferruginea Christ. --Cyathea ferrugineoides Copel.
17. Cyathea fuliginosa (Christ) Copel. --Cyathea bicolor Copel. --Cyathea dura Copel. --Cyathea lanaensis Christ. --Cyathea lepidigera Copel. --Cyathea mindanaensis (Christ) Copel. --Cyathea squamicostata Copel. --Cyathea sulitii Copel. --Cyathea warihon Copel.
18. Cyathea halconensis Christ. --Cyathea mearnsii Copel. --Cyathea melanophlebia Copel.
19. Cyathea heterochlamydea Copel. --Cyathea merrillii Copel. Photos  
20. Cyathea integra J Sm. ex Hook. --Cyathea arguta Copel. --Cyathea breviloba Copel. --Cyathea bulusanensis Copel. Photos  
21. Cyathea latebrosa (Wall.) Copel., PJS 4 c (1909) Bot. 52;
22. Cyathea latipinnula Copel.
23. Cyathea lepifera (J Sm. ex Hook.) Copel. --Cyathea calocoma (Christ) Copel. --Cyathea pteridioides Copel.
24. Cyathea loheri Christ, Bull. Herb. Boiss. 2, 6 (1906) 1007. --Cyathea campbellii Copel. --Cyathea fructuosa Copel. --Cyathea indusiosa Copel. --Cyathea matthewii (Christ) Domin. --Cyathea mitrata Copel.
25. Cyathea lurida (Blume) Copel. --Gymnosphaera melanorhachis (Copel.) Copel.
26. Cyathea masapilidensis Copel.
27. Cyathea microchlamys Holttum
28. Cyathea negrosiana Christ
29. Cyathea obliqua Copel.; --Gymnosphaera obliqua (Copel.) Copel.
30. Cyathea philippinensis Baker, Ann. Bot. 5 (1891) 186; --Copel., FFP 2 (1960) 205; --Holttum,  Fl. Males. ser. 2, 1 (1963) 149. --Type: From a cultivated plant originating from the Philippines, ex Hort. Veitch., Feb. 1878), cult. Kew (K). --Cyathea bicolana Copel. in Elmer, LPB 9 (1920) 3108; --Copel., FFP 2 (1960) 204. –-Type: Elmer 16528  (L*, U*, US*, isotypes). Luzon: Sorsogon prov., Mt Bulusan, July 1916. --Cyathea bontocensis Copel., PJS 46 (1931) 209; FFP 2 (1960) 205. –-Type: Vanoverbergh 813. Luzon:  Mountain Province. --Cyathea heteroloba Copel., PJS 38 (1929) 134; FFP 2 (1960) 204. –-Type: BS 45633 Ramos & Edaño (NY*, isotype). Luzon: Quirino prov., Mt Alzapan, altitude 1500m, May-June 1925. --Cyathea ramosiana Alderw., Bull. Jard. Bot. Buitenz. iii, 2 (1920) 137; --Copel., FFP 2 (1960) 206. --Type: BS 23411 Ramos (L*, US*, isotypes). Luzon: Sorsogon prov., July-Aug. 1915. LUZON: Mountain Province, Quirino (Mt Alzapan), Aurora (Mt Camatis), Quezon (Mt Banahaw), Camarines Sur (Mt Isarog, apparently common), Sorsogon (Mt Bulusan), MINDORO: Mindoro Oriental (Mt Halcon). Montane forests, alt. 800-1500m.   Photos  
31. Cyathea polypoda Baker in Stapf, Trans. Linn. Soc. Lond. Bot. 4 (1894) 250. --Gymnosphaera dinagatensis (Copel.) Copel. --Gymnosphaera glabra sensu Copel. Fern Fl. Philip. 2 (1960) 235, non Blume
32. Cyathea robinsonii Copel. --Cyathea pseudoalbizzia Copel.
33. Cyathea rufopannosa Christ
34. Cyathea setulosa Copel.
35. Cyathea sibuyanensis Copel.
36. Cyathea squamulata (Blume) Copel., PJS 4 c (1909) Bot. 37; --Gymnosphaera squamulata Blume
37. Cyathea suluensis Baker. --Cyathea sessilipinnula Copel.
38. Cyathea trichophora Copel., PJS 6 c (1911) Bot. 363; --Gymnosphaera trichophora (Copel.) Copel.
39. Cyathea tripinnata Copel., PJS 1 (1906) Suppl. 251. --Cyathea densisora Alderw. --Cyathea leucostegia Copel. --Cyathea leytensis Copel.
40. Cyathea zamboangana Copel. --Cyathea urdanetensis Copel.

Gymnosphaera Blume = Cyathea

Cystopteridaceae

Currania Copeland = Gymnocarpium

Cystopteris Bernhardi

1. Cystopteris tenuisecta (Blume) Mett., Ann. Lugd. Bat. 1 (1864) 241; --Acystopteris tenuisecta (Blume) Tagawa. --Ctenitis tenerifrons (Christ) Copel.

Gymnocarpium Newman

1. Gymnocarpium oyamense (Baker) Ching. --Currania gracilipes Copel.

Davalliaceae

Araiostegia Copeland = Davallia

Davallia J Smith

1. Davallia angustata Hook. & Grev.; --Humata angustata (Wall.) J Sm. --Humata microsora Alderw.
2. Davallia brevipes Copel.; --Hovenkamp & de Joncheere, Blumea 33 (1988) 408. Philippines, Sulawesi, New Guinea.
3. Davallia denticulata (NL Burm.) Mett., ex Kuhn, Fil. Deck., (1867) 27. Photos
   
   • var. denticulata
   • var. elata (G Forst.) Kuhn
4. Davallia divaricata Blume var. divaricata.
5. Davallia embolostegia Copel., PJS 1 (1906) Suppl. 147, t. 3. Photos
6. Davallia falcinella (J Sm.) C.Presl.; --Trogostolon falcinellus (C.Presl.) Copel.
7. Davallia heterophylla Sm.; --Humata heterophylla (J Sm.) Desv., Prodr. (1825) 323. --Humata ophioglossa Cav. Photos
8. Davallia hymenophylloides (Blume) Kuhn; --Araiostegia hymenophylloides (Blume) Copel., PJS 34 (1927) 241. Photos
9. Davallia pectinata Sm.; --Humata pectinata (J Sm.) Desv. Photos
10. Davallia repens (L.f.) Kuhn.; Humata repens (L.f.) Diels in Engl. & Prant, Nat. Pflanzenfam. 14 (1899) 209. --Humata intermedia C Chr. --Humata obtusata Alderw. --Humata pusilloides Copel. --Humata trifoliata Cav. --Humata vestita (Blume) T Moore, Index (1857) 92. Photos
11. Davallia sessilifolia Blume
12. Davallia solida (Forst.) Sw., Schrad. J. Bot. 1800 (1801) 87. --Davallia elmeri Copel. --Davallia robinsonii Copel. Malesia to Polynesia. Throughout the Philippines. Epiphytic or epilithic, sea-level to c. 600m. Photos
13. Davallia trichomanoides Blume. Photos
    • var. trichomanoides
    • var. lorrainii (Hance) Holttum; --Davallia lorrainii Hance, Ann. Sci. Nat. 5, 5 (1866) 254. 
14. Davallia wagneriana Copel. Photos

Davallodes (Copeland) Copeland

1. Davallodes hirsutum (C.Presl.) Copel. --Davallodes grammatosorum Copel. --Davallodes gymnocarpum Copel. --Davallodes laxum Copel. --Davallodes congestum Copel. Photos

Humata Cavanilles = Davallia

Trogostolon Copeland = Davallia falcinella

Dennstaedtiaceae 

including Hypolepidaceae, Monachosoraceae, and Pteridiaceae  

Dennstaedtia Bernhardi

1. Dennstaedtia articulata Copel.
2. Dennstaedtia dennstaedtioides (Copel.) Copel.
3. Dennstaedtia elmeri Copel.
4. Dennstaedtia fusca Copel.
5. Dennstaedtia glabrata (Cesati) C Chr.
6. Dennstaedtia hooveri Christ. Photos  
7. Dennstaedtia macgregori Copel.
8. Dennstaedtia merrillii Copel.
9. Dennstaedtia philippinensis Copel.
10. Dennstaedtia resinifera (Blume) Mett. ex Kuhn. --Dennstaedtia cuneata (J Sm. ex Hook.) T Moore.
11. Dennstaedtia rufidula C Chr. in C Chr. & Holltum, Gard. Bull. Sing. 7 (1934) 226, t. 51.
12. Dennstaedtia scabra (Wall.) T Moore. Photos
13. Dennstaedtia scandens (Blume) T Moore.
14. Dennstaedtia smithii (Hook.) T Moore.
15. Dennstaedtia williamsii Copel.

Histiopteris (Agardh) J Smith

1. Histiopteris incisa (Thunb.) J Sm., Hist. Fil. (1875) 295. --Histiopteris montana Copel. Photos  

Hypolepis Bernhardi

1. Hypolepis alpina (Blume) Hook., Sp. Fil. 2 (1852) 63.
2. Hypolepis bamleriana Rosenst., Feddes Repert. 10 (1912) 325.
3. Hypolepis glandulifera Brownsey & Chinnock, J. Adelaide Bot. Gard. 10 (1987) 16.
4. Hypolepis malesiana Brownsey.
5. Hypolepis pallida (Blume) Hook.?, Sp. Fil. 2 (1852) 64; Doubtfully occurring in the Philippines (Negros: Amlan, MG Price 2479).
6. Hypolepis polypodioides (Blume) Hook.
7. Hypolepis punctata (Thunb.) Mett. ex Kuhn Doubtfully occurring in the Philippines (Luzon: Ilocos Norte, Mt. Bubungkayo, Iwatsuki et al. P375 (L).
8. Hypolepis tenuifolia (Forst. f.) Bernh. ex C.Presl. 

Microlepia C.Presl.

1. Microlepia herbacea Ching & C Chr. apud Tard. ex C Chr. --Microlepia trapeziformis (Roxb.) Kuhn. Photos
2. Microlepia platyphylla (D Don) J Sm.
3. Microlepia protracta Copel.
4. Microlepia speluncae (L.) T Moore. --Microlepia sablanensis Christ. --? Microlepia manilensis C.Presl. Photos  
5. Microlepia strigosa (Thunb.) C.Presl. Photos
6. Microlepia todayensis Christ
7. Microlepia trichotricha J Sm.

Monachosorum Kunze

1. Monachosorum henryi Christ Luzon: Cordillera Central Mtn. Range, Mt. Data, alt. 2220 m, Co 4086 (L, PNH, PUH). New to the Philippines. Previously known from Taiwan and Mainland China (Guangdong, Guangxi, Guizhou and NE & W Yunnan). Photos  
2. Monachosorum subdigitatum (Blume) Kuhn, Chaetopt. (1882) 345; 

Paesia St. Hilaire

1. Paesia elmeri Copel. Photos
2. Paesia luzonica Christ, Philipp. J. Sci., C 3 (1908) 275. Photos
3. Paesia radula (Baker) C.Chr.

Pteridium Gleditsch ex Scopoli

1. Pteridium aquilinum (L.) Kuhn Photos
   • var. wightianum (J Agardh) Tryon. 
2. Pteridium caudatum (L.) Maxon. --Pteridium aquilinum (L.) Kuhn var. yarrabense Domin

Dicksoniaceae

Calochlaena (Maxon) M. Turner & R. White

1. Calochlaena javanica Culcita javanica (Blume) Maxon Culcita copelandii (Christ) Maxon. Culcita straminea (Labill.) Maxon. Photos  

Dicksonia L’Heritier

1. Dicksonia mollis Holttum Dicksonia blumei (Kunze) T Moore. Photos  

Diplaziopsidaceae

Edited by Julie F. Barcelona, 22 January 2013

Diplaziopsis C.Chr.

1. Diplaziopsis javanica (Blume) C.Chr., Index Filic. (1905) 227; --Diplazium javanicum (Blume) Makino, Phan. Pter. Jap. Ic. I11. 1(1901) pl. 46. Photos

Dipteridaceae

Cheiropleuria C.Presl.

1. Cheiropleuria bicuspis (Blume) C.Presl. Photos  

Dipteris Reinw.

1. Dipteris conjugata Reinw. Photos  
2. Dipteris lobbiana (Blume) Moore. Photos

Dryopteridaceae 

Acrophorus C.Presl. = Dryopteris

Acystopteris Nakai = Cystopteris

Arachniodes Blume

1. Arachniodes amabilis (Blume) Tindale; --Rumohra amabilis (Blume) Ching. Photos
2. Arachniodes aristata (Forster) Tindale; --Rumohra aristata (Forst.) Ching.
3. Arachniodes coniifolia (Wall. ex T Moore) Ching. --Rumohra wallichii Ching.
4. Arachniodes hasseltii (Blume) Ching; --Rumohra hasseltii (Blume) Ching.
5. Arachniodes tripinnata (Goldm.) Sledge. --Rumohra carvifolia (Kunze) Ching.

 Arthrobotrya J Smith = Teratophyllum

Bolbitis Schott

1. Bolbitis angustipinna (Hayata) H Ito.
2. Bolbitis appendiculata (Willd.) K Iwats. ssp. appendiculata; --Egenolfia appendiculata (Willd.) J Sm.
3. Bolbitis heteroclita (C.Presl.) Ching. --Bolbitis cuspidata (C.Presl.) Ching. --Bolbitis pseudoscalpturata (Copel.) Ching. --Bolbitis foxworthyi (Copel.) Ching. --Bolbitis tenuissima (Copel.) Copel. --Bolbitis membranacea (Copel.) C Chr. --Bolbitis edanyoi Copel. --Edanyoa difformis Copel. Photos
4. Bolbitis quoyana (Gaudich.) Ching. --Bolbitis enormis (Copel.) C Chr. --Stenosemia dimorpha Copel.
5. Bolbitis repanda (Blume) Schott.
6. Bolbitis rhizophylla (Kaulf.) Hennipman. --Egenolfia intermedia (J Sm.) Fée. --Egenolfia rhizophylla (Kaulf.) Fée. --Egenolfia fluviatilis Copel. Photos
7. Bolbitis scalpturata (Fée) Ching. Photos
8. Bolbitis sinuata (C.Presl.) Hennipman. --Bolbitis diversifolia (Blume) Schott. --Bolbitis subsimplex (Fée) Ching. --Bolbitis hydrophylla (Copel.) Ching. Photos

Hybrids

1. Bolbitis X arguta (Fée) Ching.  Hennipman doubts whether this fern warrants a separate treatment as a hybrid; it may be close to Edanyoa difformis (= B. X heteroclita).
2. Bolbitis heteroclita X rhizophylla Hennipman, Leid. Bot. Ser. 2 (1977) 291, Fig. 831-o.
3. Bolbitis X sinuosa (Fée) Copel., two nothomorphs distinguished: nm. sinuosa nm. foxii (Copel.) Hennipman and Bolbitis X foxii Copel.

Dubious species

1. Leptochilus stolonifer Christ, Bull. Herb. Boiss. Ii, 6 (1906) 1004. Luzon: Rizal prov., Angilog (type locality). Possibly synonymous with B. heteroclita but the venation pattern as given by Christ is deviating. Type not traced (Hennipman, Fl. Males. Ser ii 1 (1978) 330).

Excluded taxa

1. Bolbitis presliana (Fée) Ching  Included in Copeland’s treatment in Fern Fl. Philipp. 2 (1960) 260. This taxon does not occur in the Philippines nor anywhere in Malesia.

Cyrtomium C.Presl.

1. Cyrtomium fortunei J.Sm. forma latipinna Ching

 Diacalpe Blume = Dryopteris

Dryopsis Holttum & PJ Edwards

1. Dryopsis manipurensis (Bedd.) Holttum & PJ Edwards, Kew Bull. 41 (1986) 200. --Ctenitis copelandii (Christ) Copel. --Ctenitis manipurensis (Bedd.) Ching.

Dryopteris Adanson

1. Dryopteris cacaina Tagawa.
2. Dryopteris cochleata (Don) C.Chr.
3. Dryopteris chrysocoma C.Chr.
4. Dryopteris formosana (Christ) C.Chr. Photos
5. Dryopteris hendersonii (Bedd.) C.Chr., Index Filic. (1905) 270; --Nothoperanema hendersonii (Bedd.) Ching. --Ctenitis mearnsii Copel. As Ct. mearnsii: LUZON: Benguet (Pauai; Mt Pulag). Terrestrial in mossy forest, altitude 2100-2750m. Photos
6. Dryopteris hirtipes (Blume) Kuntze. Photos
7. Dryopteris nodosa (C. Presl) Li Bing Zhang; --Acrophorus nodosus C.Presl. --Acrophorus stipellatus (Wall.) T Moore. Photos
8. Dryopteris permagna MG Price.
9. Dryopteris polita Rosenst.
10. Dryopteris pseudocaenopteris (Kunze) Li Bing Zhang, Taxon 61(6) (2012) 1209; --Peranema aspidioides (Blume) Mett.; --Diacalpe aspidioides Blume. Photos
11.  Dryopteris pulvinulifera (Bedd.) Kuntze.
12.  Dryopteris purpurascens (Blume) Christ. Photos
13. Dryopteris sparsa (Don) Kuntze. Photos
14. Dryopteris subarborea (Baker) C.Chr.
15. Dryopteris uropinna MG Price. --Dryopteris erythrosora sensu Copel. (1960), non (Eaton) Kuntze. Photos
16. Dryopteris varia (L.) Kuntze.
17. Dryopteris wallichiana (Sprengel) Alston & Bonner. --Dryopteris paleacea (Sw.) C.Chr., nom. illeg. Photos
18. Dryopteris zhuweimingii Li Bing Zhang, Taxon 61(6) (2012) 1211; --Peranema cyatheoides D Don var. luzonicum (Copel.) Ching & SH Wu, Acta Phytotax. Sin. 21 (1983) 373; --Peranema luzonica Copel.

 Edanyoa Copeland = Bolbitis heteroclita

 Egenolfia Schott = Bolbitis

Elaphoglossum Schott ex J Smith

1. Elaphoglossum angulatum (Blume) T Moore.
2. Elaphoglossum apoense Holttum.
3. Elaphoglossum blumeanum (Fée) J Sm. var. blumeanum. --Elaphoglossum copelandii Christ var. philippinense Christ ex Holttum. Photos
4. Elaphoglossum calanasanicum Holttum.
5. Elaphoglossum callifolium (Blume) T Moore.
6. Elaphoglossum luzonicum Copel. --Elaphoglossum elmeri Copel.
7. Elaphoglossum melanostictum (Blume) T Moore. --Elaphoglossum basilanicum Copel.
8. Elaphoglossum negrosensis Holttum. --Elaphoglossum angulatum sensu Copel. (1960), pro parte, non (Blume) T Moore.
9. Elaphoglossum ophioglossoides (Goldm.) Holttum. --Elaphoglossum merrillii Christ.

Hemigramma Christ = Tectaria

Lastreopsis Ching

1. Lastreopsis tenera (R Br.) Tindale; --Ctenitis recedens (J Sm.) Copel.

Lomagramma J Smith

1. Lomagramma brooksii Copel.
2. Lomagramma copelandii Holttum.
3. Lomagramma merrillii Holttum. Photos
4. Lomagramma pteroides J Sm. --Lomagramma subcoriacea Copel. --Lomagramma pedicellata Copel. --Lomagramma cordata Copel.

Nothoperanema (Tagawa) Ching = Dryopteris

Peranema D Don = Dryopteris

Phanerophlebia C.Presl. = Cyrtomium

Polystichum Roth

1. Polystichum biaristatum (Blume) T Moore. --Polystichum aquilinum Copel.
2. Polystichum copelandii Christ.
3. Polystichum deltodon (Baker) Diels.
4. Polystichum doianum Tagawa. --Polystichum pulogense Copel.
5. Polystichum elmeri Copel. Photos
6. Polystichum fuscum Copel.
7. Polystichum horizontale C.Presl. Photos
8. Polystichum nepalense (Sprengel) C Chr.
9. Polystichum nudum Copel. Photos
10. Polystichum obliquum (D Don) T Moore.
11. Polystichum obtusum J Sm. ex C.Presl. --Polystichum edanoi Copel.
12. Polystichum (Cyrtomium) cf. caryotideum (Wall.) Presl.; --Phanerophlebia caryotidea (Wall. ex Hook. & Grev.) Copel. Philippine record based on a sterile specimen by Mearns from Mt. Sto. Tomas [Copeland, Fern Fl. Philipp. 2 (1960) 248].
13. Polystichum (Cyrtomium) cf. falcatum (L.f.) C.Presl.; --Phanerophlebia falcata (L. f.) Copel. Philippine record based on a Loher collection from Mt. Data [Copeland, Fern Fl. Philipp. 2 (1960) 248] at the turn of the 1900s. Photos

Rumohra sensu Copel. (1960) non Raddi = Arachniodes

Teratophyllum Mettenius

1. Teratophyllum aculeatum (Blume) Mett. ex Kuhn var. aculeatum.
2. Teratophyllum arthropteroides (Christ) Holttum.
3. Teratophyllum articulatum (J Sm. ex Fée) Mett. ex Kuhn; --Arthrobotrya articulata J Sm.
4. Teratophyllum koordersii Holttum.
5. Teratophyllum leptocarpum (Fée) Holttum.
6. Teratophyllum luzonicum Holttum.

Equisetaceae

Equisetum L.

 
1. Equisetum ramosissimum Desf. ssp. debile (Roxb. ex Vauch.) Hauke, Amer. Fern J. 52 (1962) 33. Photos.  

Gleicheniaceae

Edited by Pieter B. Pelser, 10 June 2012, following Christenhusz et al. 2011

Dicranopteris Bernhardi

1. Dicranopteris curranii Copel., PJS 81 (1952) 4; --Copel., FFP 1 (1958) 46; Sumatra, Peninsular Malaysia, W Java, Lesser Sunda Isls (Flores), Borneo, Sulawesi, Philippines. LUZON (Laguna), NEGROS, MINDANAO. Photos  
2. Dicranopteris linearis (Burm.) Underw., Bull. Torrey Bot. Club 34 (1907) 250; --Copel., FFP 1 (1958) 45. Photos  
   • var. altissima Holttum. 
   • var. latiloba Holttum.
   • var. rigida (Blume) Holttum. Dicranopteris crassifolia (C.Presl.) Ching, Sunyatsenia 5 (1940) 275; --Copel., FFP 1 (1958) 46.
   • var. subspeciosa Holttum.

Diplopterygium (Diels) Nakai

Photos

1. Diplopterygium elmeri (Copel.) Nakai, Bull. Natl. Sci. Mus. 29 (1950) 49; --Gleichenia elmeri Copel. in Elmer, LPB 3 (1910) 799; --Hicriopteris elmeri (Copel.) Copel., FFP 1 (1958) 43.
2. Diplopterygium laevissimum (H.Christ) Nakai, Bull. Natl. Sci. Mus. 29 (1950) 52; --Gleichenia laevissima Christ, Bull. Ac. Inst. Geogr. Bot. 3, 11 (1902) 268; --Hicriopteris laevissima (Christ) Ching, Sunyatesenia 5 (1940) 280; --Copel., FFP 1 (1958) 44.
3. Diplopterygium longissimum (Blume) Nakai, Bull. Natl. Sci. Mus. 29 (1950) 53; --Gleichenia longissima Blume, Enum. Pl. Jav. (1828) 250. --Hicriopteris glauca (Thunb.) Ching, Sunyatsenia 5 (1940) 279; --Copel., FFP 1 (1958) 43 S China to Indochina through Malesia to Melanesia (Fiji, Tahiti). In Malesia not collected in the Lesser Sunda Isls E of Flores, Sulawesi, the Moluccas and New Guinea. Throughout the Philippines, usually at 800-1500m alt. in exposed places. Photos 

Gleichenia Smith

1. Gleichenia dicarpa R Br., Prodr. Fl. Nov. Holl. (1810) 161; --Copel., FFP 1 (1958) 38. Australia, New Caledonia, Malesia: New Guinea and Philippines. MINDANAO (Agusan del Norte, Mt Urdaneta; Davao, Mt Apo). Photos  
2. Gleichenia hirta Blume, Enum. Pl. Jav. (1828) 250; --Copel., FFP 1 (1958) 40. Sticherus hirtus (Blume) Ching, Sunyatsenia 5 (1940) 203 (err. ‘hirsutus’) Moluccas and the Philippines. LUZON, MINDORO, MINDANAO. Photos  
3. Gleichenia microphylla R Br., Prodromus Florae Novae Hollandiae (1810).
4. Gleichenia paleacea (Copel.) Holttum, Reinwardtia 4 (1957) 265; --Hicriopteris paleacea Copel., PJS 81 (1952) 3; FFP 1 (1958) 44. 
5. Gleichenia peltophora Copel., PJS 40 (1929) 292, t. 1; --Copel., FFP 1 (1958) 39. New Guinea, S C Sulawesi, Borneo and Philippines. MINDANAO: South Cotabato (Mt Matutum), alt. 1600m.
6. Gleichenia sordida Copel. in Elmer, LPB 3 (1910) 798; --Hicriopteris sordida (Copel.) Copel., FFP 1 (1958) 44.
7. Gleichenia vulcanica Blume, Enum. Pl. Jav. (1828) 251; Sumatra, Peninsular Malaysia, W java, Borneo, Philippines, Sulawesi and New Guinea. MINDORO: Mindoro Oriental (Mt Halcon), PALAWAN (Mt Pulgar and Mt Mantalingahan, Co 1773), MINDANAO. Photos

Sticherus C.Presl.

Photos

1. Sticherus hispidus (Mett. ex Kuhn) Copel., Gen. Fil. (1947) 27; FFP 1 (1958) 41; --Gleichenia hispida Mett. ex Kuhn, Verh. K.K. Zool.-Bot. Ges. Wien 25 (1875) 600. --Sticherus koordersii (Christ) Copel., FFP 1 (1958) 40. --Sticherus pinnatus Copel. Sumatra, Java, Bali, N Sulawesi, Ternate and Philippines. LUZON: Benguet, Albay (Mt Mayon), NEGROS.
2. Sticherus loheri (Christ) Copel., FFP 1 (1958) 42; --Gleichenia loheri Christ, Bull. Herb. Boiss. 2, 6 (1906) 1009. --Sticherus perpaleaceus Copel., PJS 81 (1952) 3; FFP 1 (1958) 42. 
   • var. loheri LUZON: Benguet (Mt Pulog), Rizal, Quezon (Mt Banahaw).
   • var. major Holttum, Reinwardtia 4 (1959) 272; NEGROS: Negros Occidental (Mt Kanlaon). 
3. Sticherus truncatus (Willd.) Nakai, Bull. Natl. Sci. Mus. 29 (1950) 20; --Gleichenia truncata (Willd.) Sprengel, Syst. Veg. ed. 16, 4 (1827) 25. Photos
• var. truncata. --Sticherus laevigatus (Willd.) C.Presl., Tent. (1836) 52; --Copel., FFP 1 (1958) 41; Throughout Malesia, apparently most common in the west. 

Of doubtful occurrence in the Philippines

1. Sticherus milnei (Baker) Ching, Sunyatsenia 5 (1940) 284; --Gleichenia milnei Baker, Syn. Fil. (1874) 449; Sulawesi, Moluccas, New Guinea, Admiralty Isls, Solomons, New Hebrides. ?LUZON

Hymenophyllaceae

Edited by Pieter B. Pelser, 10 June 2012, following Ebihara et al. 2006.

Abrodictyum C.Presl.

Callistopteris C.Presl.

Cephalomanes C.Presl.

1. Cephalomanes atrovirens C.Presl., Hymen. (1843) 17, pl. 5; --Copel., FFP 1 (1958) 77; --Trichomanes atrovirens (C.Presl.) Kunze, Bot. Zeit. 6 (1847) 371; --Copel., PJS 51 (1933) 251, pl. 52, Fig. 3; pl. 55, Fig. 2. Photos.  
2. Cephalomanes crassum (Copel.) MG Price; --Trichomanes crassum Copel.
3. Cephalomanes javanicum (Blume) Bosch var. asplenioides (C.Presl.) K Iwats. --Cephalomanes oblongifolium C.Presl., Epim. Bot. (1849) 19, pl. 10; --Copel., FFP 1 (1923) 78. --Trichomanes asplenioides C.Presl. --Trichomanes laciniatum Roxb. Photos

Crepidomanes (C.Presl.) C.Presl.

1. Crepidomanes bipunctatum (Poir.) Copel., PJS 67 (1938) 59; FFP 1 (1958) 72; --Trichomanes bipunctatum Poir. in Lam., Encycl. 8 (1808) 69; --Copel., PJS 51 (1933) 177, pl. 18, Fig. 3, 1-4.
2. Crepidomanes brevipes (C.Presl.) Copel., PJS 67 (1938) 60; FFP 1 (1958) 71; --Trichomanes brevipes (C.Presl.) Baker, Syn. Fil. (1867) 84; --Copel., PJS 51 (1933) 182, pl. 20.; --Didymoglossum brevipes C.Presl.
3. Crepidomanes christii (Copel.) Copel., PJS 67 (1938) 60; FFP 1 (1958) 73; --Trichomanes christii Copel., PJS 1 (1906) Suppl. 251; PJS 51 (1933) 185, pl. 21.
4. Crepidomanes grande (Copel.) Ebihara & K. Iwats., Blumea 51 (2006) 239; --Cephalomanes grande (Copel.) K Iwats.; --Trichomanes grande Copel., PJS 6 c (1911) 70; PJS 51 (1933) 224, pl. 40, Fig. 1-4; --Nesopteris grandis (Copel.) Copel., PJS 67 (1938) 66; FFP 1 (1958) 76.
5. Crepidomanes humile (Forst.) Bosch; --Trichomanes humile Forst., Prodr. (1786) 12; --Copel., PJS 51 (1933) 164, pl. 12; --Crepidopteris humilis (Forst.) Copel., PJS 67 (1938) 58; FFP 1 (1958) 70. --Trichomanes gracillimum Copel., PJS 51 (1933) 168, pl. 13; --Crepidopteris gracillima (Copel.) Copel., PJS 67 (1938) 58; FFP 1 (1958) 70.
6. Crepidomanes minutum (Blume) K Iwats.; --Trichomanes minutum Blume, Enum. Pl. Jav. (1828) 223; --Trichomanes minutum Blume forma alagensis (C Chr.) Copel; --Gonocormus minutus (Blume) Bosch, Hymen. Javan. (1861) 7, pl. 3; --Copel., FFP 1 (1958) 68. --Trichomanes alagensis Christ, PJS 3 c (1908) Bot. 270; --Copel., PJS 51 (1933) 152, pl. 6, Fig. 4-7; --Gonocormus alagensis (Christ) Copel., PJS 67 (1938) 57. Photos
7. Crepidomanes rupicolum (Raciborski) Copel., PJS 67 (1938) 59; FFP 1 (1958) 73; --Trichomanes rupicolum Raciborski, Pterid. Buitenz. (1898) 24. 
8. Crepidomanes sp. 1. --Trichomanes latifrons Bosch; --Vandenboschia latifrons (Bosch) Copel.  
9. Crepidomanes thysanostomum (Makino) Ebihara & K.Iwats., Blumea 51 (2006) 240; --Cephalomanes thysanostomum (Makino) K Iwats.; --Trichomanes thysanostomum Makino, Bot. mag. Tokyo 12 (1898) 193; --Nesopteris thysanostoma (Makino) Copel., PJS 67 (1938) 66. 

Crepidopteris Copeland = Crepidomanes

Didymoglossum Desv.

1. Didymoglossum mindorense (H.Christ) K.Iwats., Blumea 51 (2006) 236; -- Trichomanes mindorense Christ, PJS 3 c (1908) Bot. 270; --Copel., PJS 51 (1933) 209, pl. 34,  Fig. 1, 2; --Microgonium mindorense (Christ) Copel., PJS 67 (1938) 61; PJS 51 (1933) 164, pl. 12. --Trichomanes craspedoneurum Copel., PJS 7 c (1912) Bot. 58; PJS 51 (1933) 208, pl. 33, Fig. 5-7; --Microgonium craspedoneurum (Copel.) Copel., PJS 67 (1938) 61; FFP 1 (1958) 74.
2. Didymoglossum motleyi (Bosch) Ebihara & K. Iwats., Blumea 51 (2006) 236; --Trichomanes motleyi Bosch, Ned. Kruid. Arch. 5 (1861) 145; --Copel., PJS 51 (1933) 201, pl. xx, Fig. 1-4; --Microgonium motleyi Bosch, Hymen. Javan. (1861) 5, pl. 1; --Copel., FFP 1 (1958) 75. --Trichomanes beccarianum Cesati, Atti Accad. Napoli, Part 8 7 (1876) 8, pl. 1, Fig. 2; --Copel., PJS 51 (1933) 200, pl. 29; --Microgonium beccarianum (Cesati) Copel., PJS 67 (1938) 63; FFP 1 (1958) 74. Photos

Gonocormus Bosch = Crepidomanes

Hymenophyllum Smith

1. Hymenophyllum acanthoides (Bosch) Rosenst., Bull. Jard. Bot. Buitenz. 2, 2 (1911) 25; --Copel., PJS 64 (1937) 45, pl. 17; --Meringium acanthoides (Bosch) Copel., PJS 67 (1938) 42; FFP 1 (1958) 63.
2. Hymenophyllum acutum (C. Presl) Ebihara & K. Iwats., Blumea 51 (2006) 232; --Crepidomanes acutum (C. Presl) K Iwats.; --Trichomanes acutum C.Presl., Epim. Bot. (1849) 258. 
3. Hymenophyllum angulosum Christ, PJS 3 c (1908) Bot. 269; --Copel., PJS 64 (1937) 33, pl.  13; --Mecodium angulosum (Christ) Copel., PJS 64 (1937) 109, pl. 50; FFP 1 (1958) 49. 
4. Hymenophyllum badium Hook. & Grev., Icon. Fil. (1828) pl. 76; --Copel., PJS 64 (1937) 144, pl. 76; --Mecodium badium (Hook. & Grev.) Copel., PJS 67 (1938) 23; FFP 1 (1958) 53. --Mecodium crispatum (Wall.) Copel., PJS 67 (1938) 23; FFP 1 (1958) 54; --Hymenophyllum crispatum Wall. in Hook. & Grev., Icon. Fil. (1828) pl. 72. Photos
5. Hymenophyllum bartlettii (Copel.) C Morton; --Meringium bartlettii Copel., PJS 73 (1940) 464; FFP 1 (1958) 62. 
6. Hymenophyllum bicolanum Copel., PJS 64 (1937) 31, pl. 10; --Meringium bicolanum (Copel.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 60.
7. Hymenophyllum blandum Raciborski, Pterid. Buitenz. (1898) 20; --Copel., PJS 64 (1937) 50,  pl. 18; --Meringium blandum (Raciborski) Copel., PJS 67 (1938) 43; FFP 1 (1958) 63.
8. Hymenophyllum bontocense Copel., PJS 64 (1937) 33, pl. 12; --Meringium bontocense (Copel.) Copel., PJS 67 (1938) 41; FFP 1 (1958) 60.
9. Hymenophyllum campanulatum H.Christ, PJS 2 c (1907) Bot.155; --Copel., PJS 64 (1937) 32,  pl. 11; --Meringium campanulatum (Christ) Copel., PJS 67 (1938) 40; FFP 1 (1958) 60.
10. Hymenophyllum crispatum Wall. in Hook. & Grev., Icon. Fil. (1828) pl. 72; --Copel., PJS 64 (1937) 148, pl. 77; --Mecodium crispatum (Wall.) Copel., PJS 67 (1938) 23.
11. Hymenophyllum denticulatum Sw., Schrad. J. (1801) 100; --Copel., PJS 64 (1937) 41, pl. 15; --Meringium denticulatum (Sw.) Copel., PJS 67 (1938) 42; FFP 1 (1958) 62.
12. Hymenophyllum digitatum (Sw.) Fosberg; --Crepidomanes digitatum (Sw.) K Iwats.; --Trichomanes digitatum Sw., Syn. Fil. (1806) 370; --Copel., PJS 51 (1933) 159, pl. 7, Fig. 3, 4; --Microtrichomanes digitatum (Sw.) Copel., PJS 67 (1938) 36; FFP 1 (1958) 55.
13. Hymenophyllum edanoi (Copel.) C Morton; --Mecodium edanoi Copel., PJS 81 (1952) 2; FFP 1 (1958) 50.
14. Hymenophyllum emarginatum Sw., Schrad. J. 2 of 1800 (1801) 101; --Copel., PJS 64 (1937)  118, pl. 58; --Mecodium emarginatum (Sw.) Copel., PJS 67 (1938) 21; FFP 1 (1958) 52. Photos  
15. Hymenophyllum fimbriatum J Sm., J. Bot. 3 (1841) 418, nomen; --Hook., Sp. Fil. (1844) 102,  pl. 36C; --Copel., PJS 64 (1937) 122, pl. 60; --Mecodium fimbriatum (J Sm.) Copel., PJS 67 (1938) 21; FFP 1 (1958) 52.
16. Hymenophyllum holochilum (Bosch) C Chr., Index Fil. (1905) 226, 362; --Copel., PJS 64  (1937) 34, pl. 14; --Meringium holochilum (Bosch) Copel., PJS 67 (1938) 41; FFP 1 (1958) 62. --Hymenophyllum merrillii Christ, PJS 2 c (1907) Bot. 154; --Copel., PJS 64 (1937) 33, pl. 13; --Meringium merrillii (Christ) Copel., PJS 67 (1938) 41; FFP 1 (1958) 61. Photos
17. Hymenophyllum imbricatum Blume, Enum. Pl. Jav (1828) 220; --Copel., PJS 64 (1937) 137,  pl. 70, 71; --Mecodium imbricatum (Blume) Copel., PJS 67 (1938) 22; FFP 1 (1958) 53.
18. Hymenophyllum johorense Holttum, Gard. Bull. Sing. 4 (1929) 408, with fig.; --Copel., PJS 64  (1937) 51, pl. 19; --Meringium johorense (Holttum) Copel., FFP 1 (1958) 64. --Hymenophyllum reductum Copel., PJS 64 (1937) 53, pl. 20; --Meringium reductum (Copel.) Copel., PJS 67 (1938) 43; FFP 1 (1958) 64.
19. Hymenophyllum klabatense H.Christ, Verh. Nat. Ges. Basel 11 (1894) 4; --Copel., PJS 64 (1937) 30; --Meringium klabatense (Christ) Copel., PJS 67 (1938) 40; FFP 1 (1958) 59.
20. Hymenophyllum nitiduloides Copel., PJS 64 (1937) 112, pl. 52; --Mecodium nitiduloides (Copel.) Copel., PJS 64 (1937) 112, pl. 52; FFP 1 (1958) 50. 
21. Hymenophyllum pachydermicum Cesati, Atti Accad. Napoli 8 (1876) 8; --Meringium pachydermicum (Cesati) Copel., PJS 67 (1938) 41; FFP 1 (1958) 57.
22. Hymenophyllum pallidum (Blume) Ebihara & K. Iwats.; --Crepidomanes pallidum (Blume) K Iwats.; --Pleuromanes pallidum (Blume) C. Presl, Epim. Bot. (1849) 258; --Copel., FFP 1 (1958) 67; --Trichomanes pallidum Blume, Enum. Pl. Jav. (1828) 226; --Copel., PJS 51 (1933) 141. Photos
23. Hymenophyllum paniculiflorum C.Presl, Hymen. (1843) 147; --Copel., PJS 64 (1937) 110,  pl. 51; --Mecodium paniculiflorum (C.Presl.) Copel., PJS 67 (1938) 19; FFP 1 (1958) 50.
24. Hymenophyllum pilosissimum C.Chr., Gard. Bull. 7 (1934) 213; --Sphaerocionium pilosissimum (C Chr.) Copel., PJS 67 (1938) 33; FFP 1 (1958) 54. Photos
25. Hymenophyllum polyanthos (Sw.) Sw., Schrad. J. ii of 1800 (1801) 102; --Copel., PJS 64  (1937) 97, pl. 46, 47; --Mecodium polyanthos (Sw.) Copel., PJS 67 (1938) 19; FFP 1 (1958) 49.
26. Hymenophyllum productum Kunze, Bot. Zeit. 6 (1848) 305; --Copel., PJS 64 (1937) 118, pl.  54; --Mecodium productum (Kunze) Copel., PJS 67 (1938) 20; FFP 1 (1958) 51.
27. Hymenophyllum ramosii Copel., PJS 64 (1937) 34, pl. 9, Fig. 4-6; --Meringium ramosii (Copel.) Copel., PJS 67 (1938) 31; FFP 1 (1958) 61.
28. Hymenophyllum reinwardtii Bosch., Pl. Jungh. 1 (1856) 567; --Copel., PJS 64 (1937) 115, pl.  55; --Mecodium reinwardtii (Bosch.) Copel., PJS 67 (1938) 20; FFP 1 (1958) 51.
29. Hymenophyllum serrulatum (C.Presl.) C Chr. --Meringium meyenianum C.Presl., Hymen. (1843) 116, pl. 8 B; --Copel., FFP 1 (1958) 58; --Hymenophyllum meyenianum (C.Presl.) Copel., PJS 64 (1937) 25, pl. 8; FFP 1 (1958) 58. --Hymenophyllum edentulum (Bosch) C Chr., Index Fil. (1905) 360; --Meringium edentulum (Bosch) Copel., PJS 67 (1938) 41; FFP 1 (1958) 58. --Hymenophyllum bakeri Copel., Sarawak Mus. J. 2 (1917) 309; --Copel., PJS 64 (1937) 29; --Meringium bakeri (Copel.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 59. --Hymenophyllum pulchrum Copel., PJS 64 (1937) 22; --Meringium pulchrum (Copel.) Copel., PJS 67 (1938) 42; FFP 1 (1958) 57.
30. Hymenophyllum thuidium Harrington, J. Linn. Soc. Bot. 16 (1877) 25; --Copel., PJS 64 (1937) 116, pl. 56; --Mecodium thuidium (Harrington) Copel., PJS 67 (1938) 20; FFP 1 (1958) 52.
31. Hymenophyllum vittatum Copel., PJS 64 (1937) 31, pl. 9, Fig. 1-3; FFP 1 (1958) 59; --Meringium vittatum (Copel.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 59.
32. Hymenophyllum zamboanganum ined; --Microtrichomanes zamboanganum Copel., PJS 81 (1952) 2; FFP 1 (1958) 56; --Trichomanes zamboanganum (Copel.) C Morton.

Sphaerocionium C.Presl. = Hymenophyllum

Vandenboschia Copel.

Photos

1. Vandenboschia auriculata (Blume) Copel.; --Crepidomanes auriculatum (Blume) K Iwats.; --Trichomanes auriculatum Blume. Photos
2. Vandenboschia johnstonensis (Bailey) Copel.; --Crepidomanes johnstonense (Bailey) K Iwats.; --Trichomanes johnstonense Bailey.
3. Vandenboschia maxima (Blume) Copel.; --Crepidomanes maximum (Blume) K Iwats.; --Trichomanes maximum Blume.

Hypodematiaceae 

Edited by Julie F. Barcelona, 20 January 2013

Didymochlaena Desvaux

1. Didymochlaena truncatula (Sw.) J Sm. Photos

Hypodematium Kunze

1. Hypodematium crenatum (Forssk.) Kuhn. Photos

Leucostegia C.Presl.

1. Leucostegia immersa (Wall.) C.Presl., Tent. (1836) 95, t. 4, Fig. 11. Photos
2. Leucostegia pallida Mett. Photos

Lindsaeaceae

Isoloma J Smith = Lindsaea

Lindsaea Dryander

1. Lindsaea adiantoides J Sm. --Lindsaea tropidorhachis Alderw. Photos
2. Lindsaea apoensis Copel. --Lindsaea havicei Copel. Photos
3. Lindsaea bouillodii Christ, Not. Syst. 1 (1909) 59;  Thailand and N Vietnam to Malesia: Peninsular Malaysia, Sumatra, W Java, Borneo, Banka and Natuna Is. New to the Philippines: PALAWAN: Mt. Mantalingahan, alt. 1200m, Co 1793 (A, PNH, PUH). 
4. Lindsaea capillacea Christ
5. Lindsaea cultrata (Willd.) Sw., Syn. Fil. (1806) 119; Lindsaea decomposita sensu Copel. (1960) non Willd., pro parte.
6. Lindsaea divergens Hook. & Grev. Isoloma divergens (Hook. & Grev.) J Sm.
7. Lindsaea doryphora Kramer Lindsaea parasitica sensu Copel. (1960) non Wall. Photos  
8. Lindsaea ensifolia Sw. ssp. ensifolia Photos
9. Lindsaea fissa Copel. Lindsaea hymenophylloides sensu Copel. (1960), non Blume. Photos  
10. Lindsaea gueriniana (Gaudich.) Desv., Mem. Soc. Linn. Paris 6 (1827) 312; --Isoloma guerinianum (Gaudich.) Fée. --Isoloma fuligineum (Copel.) Copel. Photos
11. Lindsaea hamiguitanensis D.N.Karger & V.B.Amoroso, Phytotaxa 56: 18. --Type: Karger 444 (holo PNH, iso UC, Z). Endemic to MINDANAO: Mt. Hamiguitan. Photos
12. Lindsaea heterophylla Dryander
13. Lindsaea javanensis Blume
14. Lindsaea lobata Poir. in Lam., Encycl. Suppl. 3 (1813) 448; 
15. Lindsaea longifolia Copel. 
16. Lindsaea lucida Blume, Enum. Pl. Jav. (1828) 216; Lindsaea securifolia C.Presl. ex Goldm. Lindsaea gracilis Blume in Copel. (1960), excl. syn. ssp. lucida. Photos  
17. Lindsaea merrillii Copel. ssp. merrillii
18. Lindsaea oblanceolata Alderw., Bull. Jard. Bot. Buitenz. 2, 23 (1916) 15. Photos
19. Lindsaea obtusa J Sm. ex Hook., Sp. Fil. 1 (1846) 224; Lindsaea decomposita sensu Copel. (1960) non Willd., excl. syn. Lindsaea davallioides sensu Copel. (1960) non Blume. Photos  
20. Lindsaea orbiculata (Lam.) Mett. ex Kuhn Photos
    • var. orbiculata
    • var. conmixta (Tagawa) Kramer
21. Lindsaea ovata J Sm. Isoloma ovatum (J Sm.) C.Presl.
22. Lindsaea pellaeformis Christ Luzon: Isabela prov., Divilacan, Aubarede peninsula, 50 m alt., L. Co 3580, distributed as L. gueriniana (A, PNH, PUH, CAHUP, L, US, K, MICH). In here sympatrically occurring with L. gueriniana, which according to Kramer (1971), might just be an extreme form of that variable species. New to the Philippines. Previously known only from Sulawesi. 
23. Lindsaea philippinensis Kramer. --Isoloma angustum (Copel.) Tard., comb. superfl.; non L. angusta Copel. (1952). According to Kramer (1971), doubtfully distinct from L. jamesonioides Baker of Sabah. Photos  
24. Lindsaea polyctena Kramer. Photos
25. Lindsaea pulchella (J Sm.) Mett. ex Kuhn. Photos
    
    • var. pulchella. --Lindsaea adiantifolia (Hook.) Copel. --Lindsaea adiantoides sensu Copel. (1960), pro parte.
    • var. blanda (Mett. ex Kuhn) Kramer. --Lindsaea cyathicola Copel.
26. Lindasea ramosii Copel. Very doubtfully distinct from L. longifolia.
27. Lindsaea repens (Bory) Thwaites
    
    • var. sessilis (Copel.) Kramer, Blumea 15 (1968) 568. --Lindsaea longa Copel. --Lindsaea cultripinna Copel.
    • var. pectinata (Blume) Mett. ex Kuhn in Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 277; --Lindsaea pectinata Blume. --Lindsaea macraeana (Hook. & Arn.) Copel., at least in part. --? Lindsaea roxasii Copel. --Lindsaea angusta Copel. Aberrant form from Mindanao called L. angusta Copel. may be distinguished as forma angusta (Copel.) Kramer, Blumea 15 (1968) 568. Photos
28. Lindsaea rigida J Sm. ex Hook., Sp. Fil. 1 (1846) 217, t. 63A. --Lindsaea monosora Copel.
29. Lindsaea tetragona Kramer var. tetragona. --Lindsaea tenuifolia sensu Copel. (1960) non Blume, excl. syn.

Doubtful species

1. Lindsaea decrescens Copel. “As stated by Copeland, this is intermediate between L. repens (‘L. macraena, L. longa’) and L. fissa. The isotype in MICH from Copeland’s herbarium is rather scrappy. I doubt whether it represents a distinct species” –Kramer, Fl. Males. Ser. ii 1 (1971) 253.
2. Lindsaea longa Copel. Probably only an usually deeply incised form of L. repens (Bory) Thwaites var. sessilis (Copel.) Kramer.

Sphenomeris Maxon: see Odontosoria for the Philippine species

Odontosoria (Pr.) Fée

Photos

1. Odontosoria biflora (Kaulf.) C.Chr., Index Filic. (1905) 207; --Sphenomeris biflora (Kaulf.) Tagawa. --Sphenomeris chusana sensu Copel. (1960), pro parte 
2. Odontosoria chinensis (L.) J.Sm., Bot. Voy. Herald [Seemann] 10 (1857) 430. 1857; --Sphenomeris chinensis (L.) Maxon. --Sphenomeris chusana (L.) Copel. ssp. chinensis Tropical to subtropical Asia and Oceania. Throughout the Philippines. Terrestrial or epilithic in exposed sites, from low altitudes to 2100m. Photos
   • ssp. divaricata (Christ) Kramer
3. Odontosoria retusa (Cav.) J.Sm., Bot. Voy. Herald [Seemann] 10 (1857) 430. 1857; --Sphenomeris retusa (Cav.) Maxon. Photos  

Osmolindsaea (K.U.Kramer) Lehtonen & Christenh.

Edited by Julie F. Barcelona, 2 May 2013

Photos

1. Osmolindsaea odorata (Roxb.) Lehtonen & Christenh., Bot. J. Linn. Soc. 163 (2010): 335 (-336); --Lindsaea odorata Roxb., Calc. J. Nat. Hist. 4 (1844) 511; --Lindsaea cultrata sensu Copel. (1960), pro parte Lindsaea cultrata (Willd.) Sw. var. varia Copel. --?Lindsaea cultrata var. minor Hook. --Lindsaea loheriana Christ. Photos

Tapeinidium (C.Presl.) C.Chr.

1. Tapeinidium acuminatum Kramer. Photos  
2. Tapeinidium calomelanos Kramer
3. Tapeinidium gracile (Blume) Alderw.
4. Tapeinidium luzonicum (Hook.) Kramer Photos
   • var. luzonicum
   • var. leptophyllum Kramer
5. Tapeinidium pinnatum (Cav.) C Chr. Photos  
6. Tapeinidium biserratum (Blume) Alderw. Probably a natural hybrid. According to Kramer (Fl. Males. Ser. ii 1 (1971): “The type of this ‘species’ is intermediate between T. luzonicum and T. pinnatum; there are 25-30 intermediates… Their status is not clear. They occur throughout the common range of the two species”.

Lomariopsidaceae 

Cyclopeltis J Smith

1. Cyclopeltis crenata (Fee) C Chr. --Cyclopeltis zamboangana Copel. Photos
2. Cyclopeltis presliana (J.Sm.) Berkeley. --Cyclopeltis cumingiana (Fee) C Morton.

Lomariopsis Fée

1. Lomariopsis kingii (Copel.) Holttum.
2. Lomariopsis lineata (C.Presl.) Holttum. --Lomariopsis smithii Fée. --Lomariopsis papyracea Copel. Photos
3. Lomariopsis subtrifoliata (Copel.) Holttum.

Doubtfully occurring in the Philippines

1. Lomariopsis spectabilis (Kunze) Mett. Philippine specimens formerly so-named to this species all appear to have narrower fertile pinnae and hence believed by Holttum to be L. kingii. (Fl. Males. ser. 2, 1 (1978) 262. Photos

Lygodiaceae

Lygodium Swartz

1. Lygodium auriculatum (Willd.) Alston, Reinwardtia 5 (1959) 16. --Lygodium semihastatum (Cav.) Desv., Prodr. (1827) 203; --Copel., FFP 1 (1958) 34. Indochina, Micronesia, Malesia: E Borneo, Philippines. LUZON, POLILLO, MINDORO, SAMAR, MINDANAO. Terrestrial, usually in thickets and disturbed forests. Photos  
2. Lygodium circinnatum (Burm.) Sw., Syn. Fil. (1806) 153; --Copel., FFP 1 (1958) 33. --Lygodium basilanicum Christ, PJS 2 c (1907) Bot. 179; --Copel., FFP 1 (1958) 34. Sri Lanka and India to S China, across Malesia to the Solomons. Throughout the Philippines. Terrestrial, common in brush and open sites and disturbed lowland forests. Photos  
3. Lygodium flexuosum (L.) Sw., Schrader’s J. (1801) 106; --Copel., FFP 1 (1958) 35. Sri Lanka, India, Himalayas to S China, Hongkong, Taiwan, Ryukyus, across Malesia, N Queensland, Melanesia. Terrestrial, in brush and open forests at low altitudes. Photos  
4. Lygodium japonicum (Thunb.) Sw., Schrader’s J. (1801) 106; --Copel., FFP 1 (1958) 36. --Lygodium mearnsii Copel., PJS 3 c (1908) Bot. 37; --Copel., FFP 1 (1958) 35. Sri Lanka, Himalayas (Kashmir eastwards) to E China (Zejiang), Korea, Japan and southwards to Thailand and S China; in Malesia: Banka, C & E Java, Sulawesi, Philippines, Moluccas, Lesser Sunda Isls and New Guinea. Naturalized in Florida and Texas. Terrestrial in open forests and throroughly cleared sites at low to medium altitudes. Photos  
5. Lygodium longifolium (Willd.) Sw. in Schrader’s J. Bot. 1801, pt. 2 (1803) 305; S India, Hainan. In Malesia: Sumatra, Peninsular Malaysia, Riouw and Lingga Isls., Borneo, Philippines. LUZON (doubtful), PALAWAN.
6. Lygodium merrillii Copel., PJS 2 c (1907) Bot. 146, t. 4; FFP 1 (1958) 37. --Lygodium matthewii Copel., PJS 3 c (1908) Bot. 36; --Copel., FFP 1 (1958) 37. SW China (Guizhou), N Vietnam, Sumatra, Borneo (Sarawak), Philippines. LUZON: Isabela; S Luzon, MINDORO, LEYTE. Photos
7. Lygodium microphyllum (Cav.) R Br., Prodr. Fl. Nov. Holl. (1810) 162. --Lygodium scandens (L.) Sw., Schrader’s J. (1801) 106; --Copel., FFP 1 (1958) 36. Tropical Africa, SE Asia, N to Bangladesh and Hongkong and the Ryukyus, across Malesia to the Solomons, New Caledonia, N and E Australia S to New South Wales. LUZON to MINDANAO (along the Pacific side), MINDORO, CULION, BASILAN. Terrestrial in moe or less open sites. Photos
8. Lygodium salicifolium Sw.
9. Lygodium versteegii Christ, Nova Guinea 8 (1909) 161; --Copel., FFP 1 (1958) 37. New Guinea and Philippines. LUZON: Laguna (Mt Makiling), Quezon (Guinayangan), Camarines (Bicol National Park), MINDANAO: Agusan del Norte (Mt Hilong-hilong). Photos  

Marattiaceae

Angiopteris Hoffmann

1. Angiopteris evecta Sw., Syn. Fil. (1806) 395; --Copel., FFP 1 (1958) 22, in obs., pro syn. A. palmiformis. --Angiopteris palmiformis (Cav.) C Chr., Dansk Bot. Arki 3, 9 (1937) 30, t. 3, Fig. 8, 9; --Copel., FFP 1 (1958) 22. --Angiopteris elmeriana Copel. in Elmer, LPB 5 (1913) 1679; --Copel., FFP 1 (1958) 23. --Angiopteris uncinata Christ, PJS 2 c (1907) Bot. 186, non Vriese (1853); --Copel., FFP 1 (1958) 24. --Angiopteris caudata Vriese, Monog. (1853) 20; --Copel., FFP 1 (1958) 24. --Angiopteris cartilagidens Christ, Bull. Boiss. 6 (1898) 207, t. 4, Fig. 8; --Copel., FFP 1 (1958) 24. --Angiopteris pruinosa Kunze, Bot. Zeit. 4 (1846) 417; --Copel., FFP 1 (1958) 25. --Angiopteris angustifolia C Presl var. prunifera Hieron., Hedwigia 61 (1919) 257; --Copel., FFP 1 (1958) 25. Photos  

Christensenia Maxon

1. Christensenia aesculifolia (Blume) Maxon, Proc. Biol. Soc. Wash. 18 (1905) 240. Photos
   • ssp. aesculifolia; --Copel., FFP 1 (1958) 27; --Aspidium aesculifolium Blume, Enum. Pl. Javae 2 (1898) 143; --Kaulfussia aesculifolia (Blume) Blume, Enum. Pl. Javae 2 (1898) 260; --Type: Blume? (L51148), Java. India (Assam) to Malesia.  
2. Christensenia lobbiana (de Vriese) Rolleri, Amer. Fern Jour. 83(1) (1993) 17; --Kaulfussia lobbiana de Vriese & Harting, Monogr. Marattiac. (1853) 14; --Type: "Habitat in insulam Javam ". --Christensenia cumingiana Christ, Philipp. J. Sci. 2 (1907) 186; --Type: Copeland, s.n. (holo, MICH (HC1110)?, iso, PNH62411?).  Philippines, Borneo, Thailand, Sumatra and Java. Photos. 
   There may very well be one morphologically variable species. I have seen both 3-5-foliate fronds in one plant in Panay. Larger fronds tend to be more dissected than smaller ones.

Macroglossum Copeland [sometimes included in Angiopteris]

1. Macroglossum alidae Copel.

Marattia Swartz p.p. = Ptisana

Ptisana Murdock

Edited by Julie F. Barcelona, 13 January 2013

Photos 

1. Ptisana fraxinea (Sm.) Murdock, Taxon 57(3) (2008) 746; --Marattia fraxinea Sm., Pl. Ic. ined. 2. (1790) t. 48. Photos
2. Ptisana pellucida (C.Presl) Murdock, Taxon 57(3) (2008) 747; --Marattia pellucida C Presl, Suppl. Tent. (1845) 10; --Copel., FFP 1 (1958) 26. --Marattia ternatea Christ, PJS 2 c (1907) Bot. 185; --Copel., FFP 1 (1958) 26. C LUZON southwards to MINDANAO: Mt. Matutum. Photos  
3. Ptisana sylvatica (Blume) Murdock, Taxon 57(3) (2008) 747; --Marattia sylvatica Blume, Enum. (1828) 256; --Copel., FFP 1 (1958) 27. LUZON: Cordillera Central. Common in mossy montane forests to c. 2400m. Photos

Marsileaceae

Marsilea L.

1. Marsilea crenata C.Presl. Photos  

Matoniaceae

Matonia R Brown

1. Matonia foxworthyi Copel.

Nephrolepidaceae

Edited by Julie F. Barcelona, 16 January 2013

Nephrolepis Schott

1. Nephrolepis acutangula C.Presl. --Nephrolepis pubescens Copel.
2. Nephrolepis auriculata (L.) Trimen.
3. Nephrolepis biserrata (Sw.) Schott.
4. Nephrolepis clementis Christ. Probably synonymous with Nephrolepis radicans.
5. Nephrolepis cordifolia (L.) Presl Nephrolepis glabra Copel. Photos
6. Nephrolepis exaltata (L.) Schott. Photos
7. Nephrolepis falcata (Cav.) C Chr. Photos
8. Nephrolepis hirsutula (G Forst.) C.Presl.
9. Nephrolepis multiflora (Roxb.) Jarrett ex C Morton.
10. Nephrolepis radicans (Burm.) Kuhn.

Oleandraceae

Oleandra Cavanilles

1. Oleandra benguetensis Copel.
2. Oleandra cumingii J Sm. --Oleandra scandens Copel.
3. Oleandra herrei Copel. Photos
4. Oleandra maquilingensis Copel. --Oleandra colubrina sensu Copel., Fern Fl. Philipp. 1 (1958) 181, pro parte.
5. Oleandra nitida (Copel.) Copel.
6. Oleandra pistillaris (Sw.) C Chr. --Oleandra neriiformis Cav. --Oleandra colubrina (Blanco) Copel. Photos
7. Oleandra sibbaldii Grev.

Ophioglossaceae

Botrychium Swartz

1. Botrychium daucifolium Wall. ex Hook. & Grev., Icon. Fil. (1829) t. 161; --Copel., FFP 1 (1958) 20. Photos  
2. Botrychium lanuginosum Wall. ex Hook. & Grev., Icon. Fil. (1828) t. 79; --Copel., FFP 1 (1958) 19.

Helminthostachys Kaulfuss

1. Helminthostachys zeylanica (L.) Hook., Genera (1840) t. 47; --Copel., FFP 1 (1958) 20; -- Holttum, Rev. Fl. Mal. 2 (1954) 42, Fig. 2. Sri Lanka and India to Hainan, Taiwan, across Malesia to New Caledonia. Throughout the Philippines. Terrestrial in open brushy places near sea level on damp ground. Photos  

Ophioglossum L.

1. Ophioglossum costatum R.Br.
2. Ophioglossum gramineum Willd. var. gramineum .
3. Ophioglossum intermedium Hook., Icon. Pl. 10 (1854) t. 995; --Copel., FFP 1 (1958) 18. Photos
4. Ophioglossum nudicaule L.f., Suppl. (1781) 443; --Copel., FFP 1 (1958) 18.
   • var. nudicaule
   • var. macrorrhizum (Kunze) Clausen
5. Ophioglossum pendulum L., Sp. Pl. ed. 2 (1763) 1518; --Copel., FFP 1 (1958) 19; --Holttum, Rev. Fl. Mal. 2 (1954) 40. Photos.  
   • forma pendulum Old World tropics from Madagascar through tropical Asia to Polynesia. Throughout the Philippines. Epiphytic in everwet rainforests.
   • forma pendulum X forma nutans Alderw.
   • forma pendulum X forma falcatum (C Presl) Wief. 
6. Ophioglossum ramosii Copel., PJS 56 (1935) 97, pl. 1, Fig. 1, 2; FFP 1 (1958) 18. Photos
7. Ophioglossum reticulatum L., Sp. Pl. (1753) 1063; --Copel., FFP 1 (1958) 17; forma reticulatum. --Ophioglossum petiolatum Hook., Exot. Fl. (1823) pl. 56; --Copel., FFP 1 (1958) 17. --Ophioglossum nudicaule L. f. var. grandifolium Clausen, Mem. Torr. Bot. Club 19 (1938) 151; --Copel., FFP 1 (1958) 18. Pantropic. Probably throughout the Philippines. Terrestrial in humid mostly shaded sites. Only present during the wet season, 0-1600m. Photos  
   • forma dilatatum (Miq.) Wief.
   • forma complicatum (Miq.) Wief. 

Osmundaceae

Osmunda L.

1. Osmunda banksiifolia (C Presl) Kuhn, Ann. Lugd.-Bat. 4 (1869) 299; --Copel., FFP 1 (1958) 30. Photos
2. Osmunda bromeliifolia (C Presl) Copel., PJS 2 c (1907) Bot. 147; --Copel., FFP 1 (1958) 30.
3. Osmunda herbacea Copel., PJS 40 (1929) 291; FFP 1 (1958) 31.

Plagiogyriaceae

Plagiogyria (Kunze) Mett.

 
1. Plagiogyria adnata (Blume) Bedd.
2. Plagiogyria egenolfioides (Baker) Copel. var. latipinna (Copel.) XC Zhang & Noot. --Plagiogyria tuberculata Copel. Photos
3. Plagiogyria euphlebia (Kunze) Mett. --Plagiogyria christii Copel. Photos  
4. Plagiogyria falcata Copel.
5. Plagiogyria glauca (Blume) Mett. --Plagiogyria nana Copel.; --Plagiogyria glauca var. nana (Copel.) C Chr. Photos
6. Plagiogyria pycnophylla (Kunze) Mett. Photos
7. Plagiogyria stenoptera (Hance) Diels.

Polypodiaceae

including Drynariaceae, Grammitidaceae, Loxogrammaceae, Platyceriaceae

Edited by Barbara Parris, 30 September 2012

Acrosorus Copeland

1. Acrosorus friderici-et-pauli (Christ) Copel. --Acrosorus exaltatus Copel. --Acrosorus merrillii Copel. --Acrosorus symmetricus Copel. --Acrosorus triangularis (Scort. ex Bedd.) Copel.
2. Acrosorus nudicarpus PM Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp. 32 (1980) 47,  t. 2; --Xiphopteris nudicarpa (PM Zamora & Co) Parris in Parris et al., Pl. Mt Kinabalu 1 (1992) 77. --Grammitis palasaba MG Price, Contr. Univ. Mich. Herb. 16 (1987) 196.
3. Acrosorus streptophyllus (Baker) Copel. Recorded by Price (1972); some material so identified belongs to A. friderici-et-pauli. Occurrence in the Philippines needs confirmation.

Aglaomorpha Schott

1. Aglaomorpha acuminata (Willd.) Hovenkamp; --Photinopteris acuminata (Willd.) C Morton. --Photinopteris speciosa C.Presl. Photos
2. Aglaomorpha cornucopia (Copel.) Roos; --Thayeria cornucopia Copel. Photos
3. Aglaomorpha heraclea (Kunze) Copel.; --Drynariopsis heraclea (Kunze) Ching. Photos
4. Aglaomorpha meyeniana (Hook.) Schott. Photos
5. Aglaomorpha pilosa (Hook. & Bauer) Copel. Photos
6. Aglaomorpha splendens (Hook. & Bauer) Copel. Photos

 Anapausia C.Presl.  = Leptochilus

Arthromeris (T Moore) J Sm.

1. Arthromeris proteus (Copel.) Tagawa.

Belvisia Mirbel

1. Belvisia glauca (Copel.) Copel.; --Hymenolepis platyrhynchos Kunze var. glauca Copel.; --Hymenolepis glauca (Copel.) C.Chr. Photos
2. Belvisia mucronata (Fée) Copel. var. mucronata
3. Belvisia platyrhynchos (Kunze) Copel. Photos
4. Belvisia spicata (L.f.) Copel. --Belvisia revoluta (Blume) Copel.
5. Belvisia validinervis (Kunze) Copel. var. validinervis Photos

Calymmodon C.Presl.

Edited by Barbara Parris, 30 September 2012

1. Calymmodon clavifer (Hook.) T Moore.
2. Calymmodon cucullatus (Nees & Blume) C.Presl. Photos
3. Calymmodon gracilis (Fée) Copel. Photos
4. Calymmodon muscoides (Copel.) Copel. Photos
5. Calymmodon ordinatus Copel.

Christopteris Copeland

1. Christopteris sagitta (Christ) Copel. --Christopteris copelandii Christ. Photos

Chrysogrammitis Parris

1. Chrysogrammitis glandulosa (J Sm.) Parris. --Ctenopteris merrittii (Copel.) Tagawa.
2. Chrysogrammitis musgraviana (Baker) Parris. --Xiphopteris apoensis (Copel.) Copel.

Ctenopteris Blume = Chrysogrammitis, Ctenopterella, Dasygrammitis,  Prosaptia, Themelium, Tomophyllum

  Christiopteris Copeland = Christopteris

 Colysis Presl = Leptochilus

  Crypsinus Presl = Selliguea

Ctenopterella Parris

1. Ctenopterella blechnoides (Grev.) Parris, Gard. Bull. Sing. 58 (2007) 235; --Ctenopteris blechnoides (Grev.) Wagner & Grether. Photos
2. Ctenopterella denticulata (Blume) Parris. --Ctenopteris solida Kunze in Zoll., Syst. Verz. (1854) 37.
3. Ctenopterella pediculata (Baker) Parris, Gard. Bull. Sing. 58 (2007) 237; --Ctenopteris pediculata (Baker) Copel.

Dasygrammitis Parris

1. Dasygrammitis fuscata (Blume) Parris, Gard. Bull. Sing. 58 (2007) 239; --Ctenopteris fuscata (Blume) Kunze.
2. Dasygrammitis malaccana (Baker) Parris. --Ctenopteris matutumensis Copel., Fern Fl. Philip. 3 (1960) 528, anglice, non rite publice. --Ctenopteris pachycaulis (Copel.) Copel.
3. Dasygrammitis mollicoma (Nees & Blume) Parris, Gard. Bull. Sing. 58 (2007) 239; --Ctenopteris mollicoma (Nees & Blume) Kunze. Photos

 Dendroglossa C.Presl. = Leptochilus

 Diblemma J Smith = Microsorum samarense

 Drymoglossum C.Presl. = Pyrrosia

Drynaria (Bory) J Smith

1. Drynaria descensa Copel. Photos
2. Drynaria quercifolia (L.) J Sm. India and S China, across Malesia to tropical Australia and Fiji. Throughout the Philippines. A very common epiphyte or lithophyte in seasonal areas, in open forests, about settlements, etc., usually at low altitudes. Photos
3. Drynaria rigidula (Sw.) Bedd. Photos
4. Drynaria sparsisora (Desv.) T Moore. Photos

Drynariopsis (Copeland) Ching = Aglaomorpha

Goniophlebium (Blume) C.Presl.

1. Goniophlebium benguetense (Copel.) Copel.; --Schellolepis benguetensis (Copel.) Pic.-Serm.
2. Goniophlenium coadunatum Barcelona & MG Price.
3. Goniophlebium mengtzeense (Christ) Rödl-Linder; --Goniophlebium argutum auctt. philipp. non J Sm.
4. Goniophlebium percussum (Cav.) Wagner & Grether; --Schellolepis percussa Pic.-Serm.
5. Goniophlebium persicifolium (Desv.) Bedd.; --Schellolepis persicifolia (Desv.) Pic.-Serm. Photos
6. Goniophlebium pseudoconnatum (Copel.) Copel. Photos
7. Goniophlebium subauriculatum (Blume) C.Presl. --Goniophlebium tomentellum (C Chr.) Copel.; --Schellolepis tomentella (C Chr.) Pic.-Serm. Goniophlebium integrum Copel. Photos
8. Goniophlebium terrestre Copel.; --Schellolepis terrestris (Copel.) MG Price

 Grammitis auctt. Males., non Swartz = Oreogrammitis

Lecanopteris Reinwardt

Edited by Julie F. Barcelona, 15 May 2012

1. Lecanopteris deparioides (Cesati) Baker, J. Bot. 19 (1881) 366; --NPfl. 328. --Lecanopteris curtisii Baker, J. Bot. (1881) Icon. Plant, pl. 1607; --H Gay, Gard. Bull. Sing. 45 (1993) 301. --Lecanopteris pumila Copel., Polyp. Philipp. (1905) 134; Fern Fl. Philip. 3 (1960) 499, non Blume (1829.)-- Lecanopteris philippinensis Alderw., Bull. Jard. Bot. Buitenz. Ser. 2, no. 1 (1911) 8. Sumatra, Peninsular Malaysia, Philippines to the Moluccas. MINDANAO: Lanao del Sur, Davao (Mt Apo), Davao Oriental (Mt Hamiguitan). Altitude 800-1200m. Photos
2. Lecanopteris lomarioides (Mett.) Copel. LUZON: Nueva Ecija.
3. Lecanopteris luzonensis Hennipman, Blumea 32. --Lecanopteris carnosa auctt. philipp. non (Reinw.) Blume  LUZON: Pangasinan, Pampanga, Bataan (Mt Mariveles), Laguna, Quezon, Camarines Norte. Photos
4. Lecanopteris sinuosa (Wall. ex Hook.) Copel.

Lemmaphyllum C.Presl.

1. Lemmaphyllum accedens (Blume) Donk; --Weatherbya accedens (Blume) Copel. Photos
2. Lemmaphyllum carnosum (Hook.) C.Presl. var. carnosum. --Lemmaphyllum spathulatum (C.Presl.) C.Presl. Photos

Lepisorus (J Smith) Ching

1. Lepisorus longifolius (Blume) Holttum; --Pleopeltis longifolia (Blume) Bedd.; --Paragramma longifolia (Bl) T Moore. Photos
2. Lepisorus thunbergianus (Kaulf.) Ching; --Pleopeltis thunbergiana Kaulf.

Leptochilus Kaulf.

1. Leptochilus axillaris (Cav.) Kaulf.
2. Leptochilus decurrens Blume; --Anapausia decurrens (Blume) C.Presl.
3. Leptochilus ellipticus (Thunb.) Noot.; --Colysis elliptica (Thunb.) Ching. --Colysis pothifolia (D Don) C.Presl. Photos
4. Leptochilus hemionitideus (Wall.) C.Presl. (Colysis) For some reason, this is not accounted for in the FM revision, although quite a common species in the Philippines. Photos
5. Leptochilus macrophyllus (Blume) Noot. Photos
   
   • var. macrophyllus; --Colysis macrophylla (Blume) C.Presl. --Colysis bolsteri (Copel.) Alderw.
   • var. fluviatilis (Lauterb.) Noot.
   • var. pedunculatus (Hook. & Grev.) Noot. --Colysis membranacea (Blume) C.Presl.
6. Leptochilus minor Fée Dendoglossa minor (Fée) Copel.; --Colysis minor (Fée) MG Price. Photos

Loxogramme (Blume) C.Presl.

1. Loxogramme antrophyoides (Baker) C Chr.
2. Loxogramme avenia (Blume) C.Presl. --Loxogramme malayana Copel. Photos
3. Loxogramme chinensis Ching. Possible new record for the Philippines.
4. Loxogramme conferta Copel.
5. Loxogramme dimorpha Copel.
6. Loxogramme forbesii Copel.
7. Loxogramme parallela Copel.
8. Loxogramme scolopendrioides (Gaudich.) C Morton. --Loxogramme scolopendrina (Bory) C.Presl. Photos
9. Loxogramme subselliguea (Baker) Alston. --Loxogramme major Copel.

Micropolypodium Hayata

1. Micropolypodium pulogense (Copel.) AR Sm.; --Xiphopteris pulogensis (Copel.) Copel.

Microsorum Link

1. Microsorum commutatum (Blume) Copel.; --Phymatosorus commutatus (Blume) Pic.-Serm. --Microsorum phanerophlebium (Copel.) Copel. Photos
2. Microsorum congregatifolium (Alderw.) Holttum. --Microsorum mindanaense (Christ) Copel. Photos
3. Microsorum ensatum (Thunb.) H Ito. --Neocheiropteris ensata (Thunb.) Ching.
4. Microsorum heterocarpum (Blume) Ching. Photos
5. Microsorum heterolobum (C Chr.) Copel. Probably a hybrid.
6. Microsorum insigne (Blume) Copel. --Microsorum decurrens (Brackenr.) Copel. --Microsorum palmatum auct. non (Blume) Fee. Photos
7. Microsorum longissimum Fée. --Microsorum sablanianum (Christ) Copel. Photos
8. Microsorum membranifolium (R Br.) Ching. --Microsorum plukenetii (C.Presl.) MG Price. --Microsorum alternifolium auct.: Copel., Gen. Fil. (1947), pro parte. --Microsorum longipes Copel., nom. nud. --Microsorum nigrescens (Blume) Copel.; --Phymatosorus nigrescens (Blume) Pic.-Serm. --? Microsorum schneideri (Christ) Copel. Photos
9. Microsorum monstrosum (Copel.) Copel. Photos
10. Microsorum pentaphyllum (Baker) Copel. --Microsorum curranii (Copel.) Copel. Photos
11. Microsorum pteropus (Blume) Copel.
12. Microsorum punctatum (L.) Copel. --Microsorum musifolium (Blume) Copel. --Microsorum validum (Copel.) Ching. Photos
13. Microsorum rubidum (J Sm.) Copel. Photos
14. Microsorum samarense (J Sm.) Bosman; --Diblemma samarense J Sm. --Microsorum tenuilore (J Sm. ex Mett.) Copel. Photos
15. Microsorum sarawakense (Baker) Ching. --Microsorum rizalense Copel. Photos
16. Microsorum scolopendria (Burm. f.) Copel.; --Phymatosorus scolopendria (Burm. f.) Pic.-Serm. Tropical Africa to Polynesia and Australia. Throughout the Philippines. Epiphytic or lithophytic in exposed sites and disturbed forests at low elevations, even on coastal bluffs subjected to salt spray. Common. Photos
17. Microsorum zippelii (Blume) Ching. Photos

 Neocheiropteris Christ = Microsorum

Oreogrammitis Copeland

1. Oreogrammitis adspersa (Blume) Parris, Gard. Bull. Sing. 58 (2007) 255; --Grammitis adspersa (Blume) Blume.
2. Oreogrammitis caespitosa (Blume) Parris, Gard. Bull. Sing. 58 (2007) 257; --Grammitis caespitosa Blume.
3. Oreogrammitis congener (Blume) Parris, Gard. Bull. Sing. 58 (2007) 258; --Grammitis congener Blume.
4. Oreogrammitis dolichosora (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 258; --Grammitis dolichosora (Copel.) Copel.
5. Oreogrammitis fasciata (Blume) Parris, Gard. Bull. Sing. 58 (2007) 259; --Grammitis fasciata Blume.
6. Oreogrammitis fenicis (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 259; --Grammitis fenicis Copel.
7. Oreogrammitis knutsfordiana (Baker) Parris, Gard. Bull. Sing. 58 (2007) 262; --Grammitis knutsfordiana (Baker) Copel.
8. Oreogrammitis locellata (Baker) Parris, Gard. Bull. Sing. 58 (2007) 262; --Grammitis locellata (Baker) Copel. --Grammitis stenocrypta Copel.
9. Oreogrammitis marivelesensis (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 263; --Grammitis marivelesensis Copel.
10. Oreogrammitis merrillii (Baker) Parris, Gard. Bull. Sing. 58 (2007) 263; --Grammitis merrillii (Copel.) Copel.
11. Oreogrammitis oncobasis (MG Price) Parris, Gard. Bull. Sing. 58 (2007) 264; --Grammitis oncobasis MG Price.
12. Oreogrammitis padangensis (Baker) Parris, Gard. Bull. Sing. 58 (2007) 264; --Grammitis padangensis (Baker) Copel.
13. Oreogrammitis pilosiuscula (Blume) Parris.
14. Oreogrammitis reinwardtii (Blume) Parris, Gard. Bull. Sing. 58 (2007) 266; --Grammitis reinwardtii Blume.
15. Oreogrammitis scabristipes (Baker) Parris, Gard. Bull. Sing. 58 (2007) 267; --Grammitis scabristipes (Baker) Copel. --Grammitis bulbotricha (Copel.) Copel. --Grammitis limapes Copel.
16. Oreogrammitis sumatrana (Baker) Parris, Gard. Bull. Sing. 58 (2007) 269; --Grammitis sumatrana (Baker) Copel.
17. Oreogrammitis torricelliana (Brause) Parris, Gard. Bull. Sing. 58 (2007) 270; --Grammitis torricelliana (Brause) Parris. --Grammitis microtricha Copel.
18. Oreogrammitis aff. insularis.
19. Oreogrammitis aff. knutsfordiana.

Photinopteris J Smith = Aglaomorpha

 Phymatodes C.Presl. = Microsorum

 Phymatopsis J Smith = Selliguea

 Phymatopteris Pichi-Sermolli = Selliguea

 Phymatosorus Pichi-Sermolli = Microsorum

Platycerium Desvaux

1. Platycerium coronarium (König ex Müller) Desv. Tenasserim, Thailand and Malesia. In the Philippines reported from LUZON: Cagayan to MINDANAO: Davao. High canopy epiphyte in lowland primary dipterocarp forests. Also cultivated as an ornamental.
2. Platycerium grande (Fée) Kunze MINDANAO: Zamboanga, Lanao, Davao (Mt Apo). Also reported by Copeland (1960) from S LUZON (Mt Cristobal), possibly in error. Majestic crown epiphyte in forest, lowland to 1000m. Also cultivated as an ornamental. Photos

 Pleopeltis Humboldt & Bonpland = Lepisorus

Podosorus Holttum

1. Podosorus angustatus Holttum.

Prosaptia C.Presl.

1. Prosaptia alata (Blume) Christ. --Prosaptia pinnatifida C.Presl. --Prosaptia toppingii Copel.
2. Prosaptia ancestralis Copel.
3. Prosaptia barathrophylla (Baker) MG Price. Philippine material with the following two names are referable to this species: Prosaptia khasyana (Hook.) C Chr. & Tardieu, Notul. Syst. (Paris) 8 (1939) 180; --Ctenopteris khasyana (Hook.) Holttum. 
4. Prosaptia celebica (Blume) Tagawa & K Iwats., Acta Phytotax. Geobot. 24 (1969) 61; --Ctenopteris celebica (Blume) Copel. --Ctenopteris pseudoarticulata (Copel.) Tagawa. Photos
5. Prosaptia contigua (Forst.) C.Presl. Photos
6. Prosaptia cryptocarpa Copel. Photos
7. Prosaptia davalliacea (F Muell. & Baker) Copel. --Prosaptia linearis Copel. Photos
8. Prosaptia micropora (Copel.) Parris. --Ctenopteris microcarpa Copel. Fern Fl. Philip. 3 (1960) 531, anglice, non rite publice.
9.  Prosaptia multicaudata (Copel.) Parris, Kew Bull. 50 (1995) 166; --Ctenopteris multicaudata (Copel.) Copel.
10. Prosaptia negrosensis (Copel.) MG Price, Contr. Univ. Michigan Herb. 17 (1990) 276; --Ctenopteris negrosensis (Copel.) Copel. --Ctenopteris spongiosa (Copel.) Copel.
11. Prosaptia nutans (Blume) Mett.; --Ctenopteris nutans (Blume) J Sm.
12. Prosaptia obliquata (Blume) Mett. not C Chr. & Tardieu, Notul. Syst. (Paris) 8 (1939) 180; --Ctenopteris obliquata (Blume) Copel. Photos
13. Prosaptia pectinata T Moore. --Prosaptia leysii (Baker) Ching.
14. Prosaptia urceolaris (Hayata) Copel.
15. Prosaptia venulosa (Blume) MG Price, Contr. Univ. Michigan Herb. 16 (1987) 197; --Ctenopteris venulosa (Blume) Kunze. Photos
16. Prosaptia aff. cavisora

Pteropsis Desvaux = Pyrrosia

Pyrrosia Mirbel

1. Pyrrosia lanceolata (L.) Farw. --Pyrrosia adnascens (Sw.) Ching. --Pyrrosia varia (Kaulf.) Farwell. India and S China, across Malesia to Polynesia. Throughout the Philippines. In exposed sites, open forests, etc., Epiphytic or lithophytic, usually at low altitude. Photos
2. Pyrrosia longifolia (Burm.f.) C Morton. --Pyrrosia acrostichoides (G Forst.) Ching. Photos
3. Pyrrosia nummularifolia (Sw.) Ching. Photos
4. Pyrrosia piloselloides (L.) MG Price; --Drymoglossum piloselloides (L.) Presl; --Pteropsis piloselloides (L.) Desv. --Pteropsis heterophyllum auct. non (L.) Trimen India to New Guinea. Throughout the Philippines. Epiphytic on tree trunks and branches or epilithic on outcrops and boulders along rivers and streams at low altitudes. Common. Photos
5. Pyrrosia porosa (C.Presl.) Hovenkamp var. porosa. --Pyrrosia mollis auct. non (Kunze) Ching. Photos
6. Pyrrosia rasamalae (Raciborski) Shing. --Pyrrosia lanuginosa (Giesenh.) Copel. This species has often been named P. floccigera, properly a synonym of P.  albicans, a species not known to occur in the Philippines. Photos
7. Pyrrosia samarensis (C.Presl.) Ching. --Pyrrosia philippinensis Copel., nom. superfl. Photos
8. Pyrrosia sphaerotricha (Mett.) Ching. Photos
9. Pyrrosia splendens (C.Presl.) Ching. Photos

Radiogrammitis Parris

1. Radiogrammitis alepidota (MG Price) Parris, Gard. Bull. Sing. 58 (2007) 241; --Grammitis alepidota MG Price, Philip. Agriculturist 57 (1973) 34, Fig. 1;  Taiwan (BJ Wang 17447-B, HAST) and Philippines. LUZON (Ilocos Norte, Mt Burnay; Cagayan, Mt Cetaceo; Quezon, Mt Banahaw, type locality), PALAWAN (Mt Mantalingahan). Photos
2. Radiogrammitis jagoriana (Mett. ex Kuhn) Parris, Gard. Bull. Sing. 58 (2007) 242; --Grammitis jagoriana (Mett.) Copel.
3. Radiogrammitis setigera (Blume) Parris. --Grammitis intromissa (Christ) Parris. --Grammitis fasciculata Blume, nom. illegit. pro Polypodium setigerum Blume. Photos
4. Radiogrammitis subpinnatifida (Blume) Parris; --Xiphopteris subpinnatifida (Blume) Copel.

 Schellolepis Pic.-Serm. = Goniophlebium

Scleroglossum Alderw.

1. Scleroglossum pusillum (Blume) Alderw. --Scleroglossum minus (Fée) C Chr. Photos
2. Scleroglossum sulcatum (Kuhn) Alderw. --Scleroglossum intermedium (Copel.) C Chr. --Scleroglossum pusillum Copel., non (Blume) Alderw.

Selliguea Bory

1. Selliguea albidosquamata (Blume) Parris; --Crypsinus albidosquamatus (Blume) Copel. Photos
2. Selliguea caudiformis (Blume) Carruth. --Selliguea calophlebia (Copel.) Copel.
3. Selliguea elmeri (Copel.) Ching; --Selliguea feei Bory var. elmeri (Copel.) Tagawa. Photos
4. Selliguea enervis (Cav.) Ching; --Crypsinus enervis (Cav.) Copel. Photos
5. Selliguea feei Bory; --Selliguea feei Bory var. vulcanica (Blume) Tagawa. Photos
6. Selliguea glauca (J Sm. ex T Moore) Hovenkamp; --Crypsinus glaucus (J Sm. ex T Moore) Copel. Photos
7. Selliguea heterocarpa (Blume) Blume.
8. Selliguea lagunensis (Christ) Hovenkamp; --Crypsinus lagunensis (Christ) Copel.; --Phymatopteris lagunensis (Christ) Pic.-Serm. Photos
9. Selliguea lateritia (Baker) Hovenkamp.
10. Selliguea oodes (Kunze) Hovenkamp; --Crypsinus oodes (Kunze) Copel. Photos
11. Selliguea pyrolifolia (Goldm.) Hovenkamp; --Crypsinus pyrolifolius (Goldm.) Copel. Photos
12. Selliguea stenophylla (Blume) Parris; --Crypsinus stenophyllus (Blume) Copel.
13. Selliguea taeniata (Sw.) Parris; --Crypsinus taeniatus (Sw.) Copel., including var. palmatus C Chr.; --Phymatopteris taeniata (Sw.) Pic.-Serm. --Crypsinus ramosii Copel. Photos
14. Selliguea triloba (Houtt.) MG Price; --Crypsinus trilobus (Houtt.) Copel.; --Phymatopteris triloba (Houtt.) Pic.-Serm. --Selliguea matutumensis Copel.
15. Selliguea whitfordii (Copel.) Hovenkamp; --Crypsinus whitfordii (Copel.) Copel. --Crypsinus subdimorphus Copel. Photos

Doubtful occurrence in the Philippines:

1. Selliguea hastata (Thunb.) Copel.; --Crypsinus hastatus (Thunb.) Copel.;  --Phymatopteris hastata (Thunb.) Pic.-Serm. India, China, Japan. In the Philippines known only from LUZON: Benguet. Epiphytic or lithophytic, alt. c. 2400m. According to Hovenkamp et.al. [Fl. Males. ser. 2, 3 (1998) 227], simple fronded plants from the Philippines have been identified as Phymatopteris hastata by Zamora & Co [Guide to Philipp. Flora & Fauna 2 (1986) 29], and that other than through this reference, P. hastata is not known from the Philippines.  This is incorrect as Copeland (Fern Fl. Philipp. 2 (1960) 505) reported this species to be common in Benguet at 2400 m.

Themelium (T Moore) Parris

1. Themelium cf. blechnifrons (Hayata) Parris. --Ctenopteris curtisii (Baker) Copel. Photos
2. Themelium halconense (Baker) Parris, Kew Bull. 59 (2004) 225; --Ctenopteris halconensis (Copel.) Copel.
3. Themelium loherianum (Christ) Parris, Kew Bull. 59 (2004) 225; --Grammitis loheriana (Christ) Copel.
4. Themelium taxodioides (Baker) Parris, Kew Bull. 52 (1997) 739; --Ctenopteris taxodioides (Baker) Copel.
5. Themelium tenuisectum (Blume) Parris, Kew Bull. 52 (1997) 740; --Ctenopteris tenuisecta (Blume) J Sm. Photos
6. Themelium yoderi (Copel.) Parris, Kew Bull. 52 (1997) 739; --Ctenopteris yoderi (Copel.) Copel.
7. Themelium aff. tenuisectum sp. 1.
8. Themelium aff. tenuisectum sp. 2.

Thylacopteris Kunze ex J Smith

1. Thylacopteris papillosa (Blume) J Sm. Photos

Tomophyllum (E Fournier) Parris

1. Tomophyllum macrum (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 247; --Ctenopteris macra (Copel.) Copel.
2. Tomophyllum millefolium (Blume) Parris, Gard. Bull. Sing. 58 (2007) 247; --Ctenopteris millefolia (Blume) Copel.
3. Tomophyllum minutum (Blume) Parris, Gard. Bull. Sing. 58 (2007) 247; --Ctenopteris minuta (Blume) Holttum Possibly conspecific with C. macra. It is more likely to be a misidentification of Tomophyllum macrum.  Blume’s name has priority.
4. Tomophyllum repandulum (Mett.) Parris.
5. Tomophyllum subsecundodissectum (Zoll.) Parris, Gard. Bull. Sing. 58 (2007) 249; --Ctenopteris subsecundodissecta (Zoll.) Copel. Photos

Psilotaceae

Psilotum L.

1. Psilotum complanatum Sw., Syn. Fil. (1806) 188, 414, t. 4, Fig. 5. Photos
2. Psilotum nudum (L.) Griseb., Prodr. Aethéogam. (1805) 112. Photos.  
3. Psilotum triquetrum Sw., Schrad. J. Bot. 1800 (2) 1801.

Tmesipteris Bernhardi

1. Tmesipteris lanceolata Dang, Le Botaniste 2me Serie (1891). Tmesipteris tannensis auctt. non (Spreng.) Bernh. Photos.  

Pteridaceae 

Including Acrostichaceae, Adiantaceae, Antrophyaceae, Ceratopteridaceae, Cheilanthaceae, Hemionitidaceae, Parkeriaceae, Sinopteridaceae, Taenitidaceae, Vittariaceae

Acrostichum L.

1. Acrostichum aureum L. Photos
2. Acrostichum speciosum Willd. Photos

Adiantum L.

1. Adiantum capillus-veneris L. Photos
2. Adiantum caudatum L. 
3. Adiantum cupreum Copel.
4. Adiantum edgeworthii Hook. 
5. Adiantum flabellulatum L.
6. Adiantum hispidulum Sw. Photos  
7. Adiantum malesianum Ghatak. Photos  
8. Adiantum mindanaoense Copel. 
9. Adiantum philippense L. Photos  
10. Adiantum scabripes Copel. Photos  
11. Adiantum setulosum J Sm. --Adiaantum diaphanum sensu Copel. non Blume, Enum. Pl. Jav. (1828) 215. Photos
12. Adiantum soboliferum Wall.
13. Adiantum stenochlamys Baker. --Adiantum opacum Copel., PJS 1 (1906) Suppl. 255, t. 3. Photos  

Aleuritopteris Fée

1. Aleuritopteris farinosa (Forssk.) Fée Cheilanthes agetae (Saiki) Kuo; --Cheilanthes farinosa (Forssk.) Kaulf.
2. Aleuritopteris albofusca (Baker) Pic.-Serm. Cheilanthes anceps Blanf. --Cheilanthes albofusca Baker. Photos
3. Aleuritopteris rufa (D Don) Copel. Cheilanthes dubia Hope; --Cheilanthes rufa D Don

Antrophyum Kaulfuss

1. Antrophyum immersum (Bory ex Willd.) Mett. --Antrophyum nanum Fée. Photos
2. Antrophyum latifolium Blume. Photos  
3. Antrophyum ledermannii Hieron.
4. Antrophyum plantagineum (Cav.) Kaulf. Photos
5. Antrophyum reticulatum (Forst.) Kaulf. --Antrophyum alatum Brackenr. --Antrophyum callifolium Blume. --Antrophyum clementis Christ. Photos
6. Antrophyum semicostatum Blume.
7. Antrophyum sessilifolium (Cav.) Sprengel. Photos
8. Antrophyum williamsii Benedict.

Ceratopteris Brongniart

1. Ceratopteris thalictroides (L.) Brongn. --Ceratopteris siliquosa (L.) Copel. Photos  

Cheilanthes Swartz

1. Cheilanthes belangeri (Bory) C Chr.
2. Cheilanthes javensis (Willd.) T Moore. --Cheilanthes densa (J Sm.) Fée. --Cheilanthes dilimanensis PM Zamora
3. Cheilanthes mysurensis Wall. ex Hook.
4. Cheilanthes tenuifolia (NL Burm.) Sw., Syn. Fil. (1806) 129, 332. Photos

Coniogramme Fée

1. Coniogramme fraxinea (D Don) Diels in Engl. & Prantl, Nat. Pflanzenfam. 1 (1899) 262; 
2. Coniogramme macrophylla (Blume) Hieron., Hedwigia 57 (1916) 291. Photos
3. Coniogramme serrulata Fée -- Mém. Foug., 5. Gen. Filic. 167 t. 14 B. f. 2. (1850). Photos
4. Coniogramme subcordata Copel.

Copelandiopteris Stone = Pteris

Doryopteris J Sm.

1. Doryopteris concolor (Langsdorff & Fischer) Kuhn.; --Cheilanthes concolor (Langsdorff & Fischer) R & A Tryon. --Doryopteris cuspidata Copel. Photos
2. Doryopteris ludens (Wall.) J Sm. Photos  

Gymnopteris Bernhardi [= Hemionitis]

1. Gymnopteris vestita (Wall.) Underw.

Haplopteris C.Presl

Edited by Julie F. Barcelona, 16 January 2013

Photos

1. Haplopteris alternans (Copel.) Barcelona, ined.; --Vittaria alternans Copel. --Vittaria hecistophylla Copel. Photos
2. Haplopteris anguste-elongata (Hayata) E.H. Crane; --Vittaria anguste-elongata Hayata. --Vittaria stenophylla Copel.
3. Haplopteris crispomarginata (Christ) Barcelona, ined.; --Vittaria crispomarginata Christ. --Vittaria merrillii Christ.
4. Haplopteris elongata (Sw.) E.H. Crane; --Vittaria elongata Sw. --Vittaria intermedia Blume. --Vittaria philippinensis Christ
5. Haplopteris ensiformis (Sw.) E.H. Crane; --Vittaria ensiformis Sw. --Vittaria incurvata Cav. --Vittaria lineata auct. non Smith. --Vittaria pachystemma Christ. Photos
6. Haplopteris scolopendrina (Bory) E.H. Crane; --Vittaria scolopendrina (Bory) Thwaites. --Vittaria subcoriacea Christ. Photos
7. Haplopteris taeniophylla (Copel.) E.H. Crane; --Vittaria taeniophylla Copel.
8. Haplopteris zosterifolia (Willd.) E.H. Crane; --Vittaria zosterifolia Willd. --Vittaria copelandii Alderw.

Monogramma Comm. ex Schkuhr.

1. Monogramma dareicarpa Hook. --Monogramma capillaris Copel., PJS 6c (1911) Bot. 147. Photos

Onychium Kaulfuss

1. Onychium japonicum (Thunb.) Kunze. Photos  
2. Onychium siliculosum (Desv.) C Chr.

Parahemionitis Panigrahi

Edited by Julie F. Barcelona & Pieter B. Pelser, 25 May 2012

1. Parahemionitis cordata (Roxb. ex Hook. & Grev.) Fraser-Jenk., New Sp. Syndr. Indian Pteridol. (1997) 187. --Parahemionitis arifolia (Burm. f.) Panigr.; --Hemionitis arifolia (Burm. f.) T Moore. Photos  

Pityrogramma Link

Pteris L.

1. Pteris armata C.Presl. Photos
2. Pteris biaurita L. --Pteris pacifica Hieron. --Pteris cumingii Hieron. Photos  
3. Pteris blumeana J Agardh. Photos  
4. Pteris brevis Copel. 
5. Pteris calocarpa (Copel.) MG Price; --Schizostegeopsis calocarpa Copel.; --Copelandiopteris calocarpa (Copel.) Stone.
6. Pteris cretica L. --Pteris stenophylla sensu Copel., FFP 1 (1958) 129, non Hook. & Grev. Photos  
7. Pteris dataensis Copel.
8. Pteris distans J Sm. ex Hook. Possibly a hybrid or a bizarre freak.
9. Pteris edanoi Copel., PJS 81 (1952) 8, 'edanyoi'.
10. Pteris elmeri Christ.
11. Pteris endoneura MG Price; --Copelandiopteris endoneura (MG Price) Stone.
12. Pteris ensiformis Burm. Photos
    • var. ensiformis. --Pteris mutilata L.
    • var. victoriae Hort.
13. Pteris fauriei Hieron., Hedwigia 55 (1914) 345; --Hatusima, Mem. Fac. Kagoshima Univ. 5 (1966) 20. --Pteris rigidula Copel., FFP 1 (1958) 137.
14. Pteris glaucovirens Goldm. 
15. Pteris heteromorpha Fée. Photos  
16. Pteris loheri Copel.
17. Pteris longipinnula Wall. ex J Agardh. Photos
18. Pteris luzonensis Hieron.
19. Pteris macgregorii Copel. 
20. Pteris melanocaulon Fée
21. Pteris melanorhachis Copel. 
22. Pteris mertensioides Willd. --Pteris gardneri (Fée) Hook. --Pteris excelsa Gaudich. Photos  
23. Pteris micracantha Copel.
24. Pteris mucronulata Copel. 
25. Pteris opaca (C.Presl.) J Sm. ex Fée. --Pteris taenitis Copel. Photos
26. Pteris oppositipinnata Fée. --Pteris asperula J Sm. --Pteris caesia Copel. Photos  
27. Pteris pachysora (Copel.) MG Price; --Schizostegeopsis pachysora Copel.; --Copelandiopteris pachysora (Copel.) Stone.
28. Pteris pedicellata Copel. 
29. Pteris pellucida C.Presl.
30. Pteris philippinensis Fée.; --Pteris longipes D Don var. philippinensis (Fée) Christ. --Pteris longipes D Don, Prod. Fl. Nepal. (1825) 15; --Copel., FFP 1 (1958) 133. Photos  
31. Pteris purpureorhachis Copel. 
32. Pteris ramosii Copel. Possibly a variant of P. philippinensis.
33. Pteris rangiferina C.Presl.
34. Pteris ryukyuensis Tagawa, Acta Phytotax. Geobot. 4 (1935) 204; --Hatusima, Mem. Fac. Kagoshima Univ. 5 (1966) 20. --? Pteris cretica sensu Copel., FFP 1 (1958) 129, quoad pl. ex Batanes Islands, non L. Photos  
35. Pteris schlechteri Brause. Photos  
36. Pteris semipinnata L. Photos  
37. Pteris spinescens C.Presl. --Pteris setuloso-costulata Hayata.
38. Pteris squamipes Copel. 
39. Pteris tripartita Sw. --Pteris microdictyon (Fée) Hook. Photos  
40. Pteris vittata L. Photos  
41. Pteris wallichiana J Agardh Confirmed to be in Luzon: Ilocos Norte prov., Mt. Burnay (Iwatsuki & Price, 1975).
42. Pteris whitfordii Copel.

Schizoloma Gaudich. = Taenitis

Syngramma J Smith

1. Syngramma alismifolia (C.Presl.) J Sm., Lond. J. Bot. 4 (1845) 168, t. 7-8, Fig. B. Photos  
2. Syngramma vittiformis J Sm.
3. Syngramma wallichii (Hook.) Bedd.

Taenitis Willdenow

1. Taenitis blechnoides (Willd.) Sw., Syn. Fil. 24 (1806) 220. Photos  
2. Taenitis luzonica (Alderw.) Holttum; --Syngramma luzonica Alderw.
3. Taenitis cordata (Gaudich.) Holttum; --Schizoloma cordatum Gaudich.

Vaginularia Fée

Edited by Julie F. Barcelona, 24 January 2013

Photos 

1. Vaginularia paradoxa (Fée) Mett.; --Monogramma paradoxa (Fée) Mett. Photos  
2. Vaginularia trichoidea (J Sm.) Fée; --Monogramma trichoidea J Sm. ex Hook. Photos

Vittaria Smith = Haplopteris p.p.

Saccolomataceae

Edited by Julie F. Barcelona, 16 January 2013

Orthiopteris Copel.

Photos

1. Orthiopteris campylura (Kunze) Copel.; --Saccoloma campylurum Mett. Photos  

Saccoloma Kaulf. = Orthiopteris

Salviniaceae (including Azollaceae)

Azolla Lam.

1. Azolla pinnata R.Br. var. imbricata (Roxb. ex Griff.) Bonap. --Azolla imbricata (Roxb. ex Griff.) Nakai
2. Azolla rubra R.Br., Prodr. Fl. Nov. Holland. 167 (1810) Bak. Hdb. 137. Photos

Salvinia Seguier

1. Salvinia sp.

Schizaeaceae

Schizaea Smith

1. Schizaea dichotoma (L.) Sm., Mém. Acad. Turin 5 (1793) 422; --Copel., FFP 1 (1958) 32. Photos  
2. Schizaea digitata (L.) Sw., Syn. Fil. (1806) 150, 380, t. 4, Fig. 1; --Copel., FFP 1 (1958) 32; --Actinostachys digitata (L.) Wall. ex Brackenr. Photos  
3. Schizaea inopinata Selling, Svensk. Bot. Tidskr. 40 (1946) 274, Fig. 1-7; --Copel., FFP 1 (1958) 32; --Actinostachys inopinata (Selling) Reed. Photos
4. Schizaea malaccana Baker

Tectariaceae

Aenigmopteris Holttum

1. Aenigmopteris dubia (Copel.) Holttum; --Ctenitis dubia (Copel.) Copel. LUZON: Quezon (Mt Banahaw de Lucban), NEGROS: Negros Oriental (Cuernos Mtns), LEYTE, CAMIGUIN. Terrestrial, altitude 850-1200m.
2. Aenigmopteris mindanaensis Holttum.

Arthropteris J Smith ex JD Hook.

1. Arthropteris beckleri (Hook.) Mett. --Arthropteris obliterata (R Br.) J Sm.
2. Arthropteris palisotii (Desv.) Alston. --Arthropteris glabra Copel.
3. Arthropteris repens (Brackenr.) C Chr.
4. Arthropteris wollastonii (Ridley) Holttum. --Arthropteris orientalis (J. Gmel.) C Chr. 

Ctenitis (C Christensen) C Christensen

1. Ctenitis aciculata (Baker) Ching.
2. Ctenitis atrorubens Holttum. --Ctenitis vilis sensu Copel. (1960), pro parte, non (Kunze) Ching.
3. Ctenitis boholensis Holttum.
4. Ctenitis bulusanica Holttum.
5. Ctenitis decurrentipinnata (Ching) Ching.
6. Ctenitis erythradenia Holttum. --Ctenitis vilis sensu Copel. (1960), pro parte, non (Kunze) Ching.
7. Ctenitis humilis Holttum. Photos
8. Ctenitis paleolata Copel. LUZON: Benguet (Pauai), altitude c. 2400m. Known only from the type collection.
9. Ctenitis pallens (Brackenr.) MG Price. --Ctenitis rizalensis (Christ) Copel. As C. rizalensis: LUZON: Rizal, Laguna (Mt Makiling), SAMAR, MINDANAO: Zamboanga (San Ramon), Davao (Mt Apo). Terrestrial under moist forest shade,  altitude 400-1200m.
10. Ctenitis propinqua (J Sm. ex C.Presl.) Copel. LUZON: Pampanga (Mt Arayat), Laguna (Mt Makiling), Quezon (Mt Banahaw), Batangas (Mt Makulod).
11. Ctenitis setosa (C.Presl.) Holttum. --Ctenitis vilis sensu Copel. (1960), pro parte, non (Kunze) Ching.
12. Ctenitis silvatica Holttum. --Ctenitis vilis sensu Copel. (1960), pro parte, non (Kunze) Ching.
13. Ctenitis subglandulosa (Hance) Ching. --Ctenitis rhodolepis (Clarke) Ching.
14. Ctenitis subobscura (Christ) Holttum.

Excluded

1. Ctenitis subconnexa (Christ) Holttum var. subconnexa and var. alstonii Holttum. A mislocalization to the Philippines by Holttum, Fl. Males. ser. 2, 2 (1991) 28. The locality Batjan is in Halmahera (of the Moluccas).

Heterogonium C.Presl.

1. Heterogonium aspidioides C.Presl. Photos
2. Heterogonium lobulatum Holttum.
3. Heterogonium pinnatum (Copel.) Holttum; --Stenosemia pinnata Copel. Photos
4. Heterogonium profereoides (Christ) Copel. --Heterogonium aspidioides sensu Holttum, Kalikasan 4 (1975) 224, pro parte
5. Heterogonium sagenioides (Mett.) Holttum; --Ctenitis sagenioides (Mett.) Copel. --Ctenitis obscura (Hook.) Copel.; --Heterogonium obscurum (Hook.) Holttum.
6. Heterogonium wenzelii (Copel.) Holttum; --Ctenitis wenzelii Copel. LEYTE.

Pleocnemia C.Presl.

1. Pleocnemia brongniartii (Bory) Holttum; --Arcypteris brongniartii (Bory) Holttum. --Tectaria irregularis sensu Copel. (1960), pro parte.
2. Pleocnemia conjugata (Blume) C.Presl.
3. Pleocnemia cumingiana C.Presl. --Tectaria leuzeana sensu Copel. (1960), pro parte. Photos
4. Pleocnemia irregularis (C.Presl.) Holttum; --Arcypteris irregularis (C.Presl.) Ching; --Tectaria irregularis sensu Copel. (1960), pro parte. Photos
5. Pleocnemia leuzeana (Gaudich.) C.Presl. Photos
6. Pleocnemia macrodonta (Fee) Holttum; --Arcypteris macrodonta (C.Presl.) Holttum. --Tectaria irregularis sensu Copel. (1960), pro parte.
7. Pleocnemia presliana Holttum.  LUZON: Cagayan, Aurora, Laguna, PALAWAN (Mt Gantung; Imolnod), LEYTE (Palo), MINDANAO: Davao (Mt Apo, Todaya), Surigao del Norte. Terrestrial, usually in partly shaded area, near sea-level to at least 400m. Photos

Psomiocarpa C.Presl.

1. Psomiocarpa apiifolia C.Presl. LUZON: Cagayan, Isabela, Bataan, Rizal, Laguna (Mt Makiling), Quezon, POLILLO, MASBATE, LEYTE, SAMAR, MINDANAO: Zamboanga (San Ramon). Terrestrial in forest, from near sea-level to c. 450m altitudes. Photos

Pteridrys C Christensen & Ching

1. Pteridrys microthecia (Fee) C Chr. & Ching.
2. Pteridrys syrmatica (Willd.) C Chr. & Ching.

Stenosemia C.Presl.  = Tectaria, Heterogonium

Tectaria Cavanilles

1. Tectaria adenophora Copel. SIBUYAN.
2. Tectaria angulata (Willd.) Copel. --Tectaria trifolia (Alderw.) C Chr.
3. Tectaria athyriosora MG Price BOHOL, SAMAR, MINDANAO: Surigao del Norte.
4. Tectaria aurita (Sw.) S. Chandra; --Stenosemia aurita (Sw.) Presl. Photos 
5. Tectaria barberi (Hook.) Copel.
6. Tectaria beccariana (Cesati) C Chr.
7. Tectaria calcarea (C.Presl.) Copel. POLILLO, in wet forest, altitude 50m, BOHOL, LEYTE. Photos
8. Tectaria christii Copel. LUZON: Mountain Province, Benguet, MINDORO: Mindoro Oriental (Mt Halcon). Usually on limestone, altitude c. 1500m.
9. Tectaria crenata Cav. --Tectaria grandifolia sensu Copel. (1960), pro parte. Photos
10. Tectaria decurrens (C.Presl.) Copel. --Tectaria peralata Copel. Photos
11. Tectaria devexa (Kunze ex Mett.) Copel. var. devexa. Photos
12. Tectaria dissecta (G Forst.) Lellinger; --Ctenitis dissecta (G Forst.) Copel. Photos
13. Tectaria dolichosora Copel. LUZON: Cagayan, Quezon (Infanta, Tignoan). Steep slopes near summits.
14. Tectaria griffithii (Baker) C Chr. var. griffithii. --Tectaria malayensis (Christ) Copel. Photos
15. Tectaria hilocarpa (Fée) MG Price. --Hemigramma latifolia (Meyen ex Goldm.) Copel., excl. Hemionitis zollingeri and other latter names based on this. Photos
16. Tectaria laxa (Copel.) MG Price; --Ctenitis laxa Copel. LUZON: Ilocos Norte (Mt Quebrada; Mt Siminublan). Altitude 80-1100m. Photos
17. Tectaria lobbii (Hook.) Copel. var. lobbii.
18. Tectaria manilensis (C.Presl.) Holttum var. manilensis; --Lastreopsis manilensis (C.Presl.) MG Price; --Ctenitis manilensis (C.Presl.) Holttum. --Ctenitis loheri (Christ) Copel.
19. Tectaria melanocaula (Blume) Copel. Tectaria ‘melanocaulis’ in Copel. (1960).
20. Tectaria menyanthidis (C.Presl.) Copel.
21. Tectaria mesodon (Copel.) MG Price; --Ctenitis mesodon (Copel.) Copel.; --Dryopteris mesodon Copel. LUZON: Ilocos Norte, MINDANAO: Agusan del Norte (Mt Hilong-hilong), altitude 120m.
22. Tectaria polymorpha (Wall. ex Hook.) Copel.
23. Tectaria ramosii (Copel.) Holttum; --Ctenitis ramosii Copel. As C. ramosii: BOHOL (Dimiao), SAMAR (Catubig River). Possibly near sea-level to c. 300m.
24. Tectaria repanda (Willd.) Holttum. --Tectaria crenata sensu Copel. (1960), pro parte.
25. Tectaria siifolia (Willd.) Copel.
26. Tectaria stalactica MG Price.
27. Tectaria sulitii Copel. --Tectaria loheri Copel. --Tectaria grandifolia sensu Copel. (1960), pro parte According to Holttum, Fl. Males. ser. 2, 2 (1991) 91, there may be intermediates between this species and T. crenata. As T. loheri: LUZON: Rizal (Montalban; Sumag). Photos
28. Tectaria suluensis Holttum.
29. Tectaria tabonensis MG Price.
30. Tectaria taccifolia (Fée) MG Price; --Hemigramma taccifolia (Fée) Copel. Photos
31. Tectaria trichotoma (Fée) Tagawa. --Tectaria balabacensis (Christ) MG Price; --Ctenitis balabacensis (Christ) Copel. As Ct. balabacensis: LUZON: Quezon (Guinayangan), PALAWAN, BALABAC, SIBUYAN. Petrophytic along rivers and streams in forests, altitude 100-200m in Palawan.
32. Tectaria trifida (Fée) MG Price. --Tectaria irrigua (J Sm. ex Presl) Copel.

1. Tectaria ramosii X Psomiocarpa apiifolia C.Presl.
2. Tectaria ramosii X Tectaria aurita (Sw.) S. Chandra.
3. Tectaria repanda X Tectaria hilocarpa (Fée) MG Price.

Tectaridium Copel.

1. Tectaridium macleanii Copel. --Tectaridium primitivum Copel. Photos

Thelypteridaceae

Ampelopteris Kunze

1. Ampelopteris prolifera (Retz.) Copel. Tropical and subtropical regions of the Old World. In the Philippines recorded from LUZON: Abra, Rizal, Laguna, MINDANAO: Davao del Norte (Maco, Lake Leonard), Agusan del Sur (Agusan Marsh). Terrestrial, often forming mats or tangles in swampy sites, lake margins or river embankments. Low altitudes. Photos  

Amphineuron Holttum

1. Amphineuron attenuatum (Kuntze) Holttum. --Cyclosorus alatellus sensu Copel. (1960), excl. syn. Lastrea erubescens sensu Copel. (1960).
2. Amphineuron immersum (Blume) Holttum.; --Lastrea immersa (Blume) T Moore. --Lastrea verrucosa C.Presl. 
3. Amphineuron opulentum (Kaulf.) Holttum. --Cyclosorus extensus (Blume) Ching.
4. Amphineuron terminans (Hook.) Holttum. --Cyclosorus interruptus sensu Copel. (1960), non (Willd.) H Ito Sri Lanka to Polynesia. Throughout the Philippines in the more seasonal regions. Terrestrial, common in thickets on hillsides and in valleys at low and medium altitudes.

Chingia Holttum

1. Chingia christii (Copel.) Holttum.; --Cyclosorus christii (Copel.) Copel.
2. Chingia clavipilosa Holttum var. clavipilosa.
3. Chingia ferox (Blume) Holttum.; --Cyclosorus ferox (Blume) Ching. Photos  
4. Chingia paucipaleata Holttum.
5. Chingia pricei Holttum.
6. Chingia urens Holttum.

Christella Léville emend. Holttum

1. Christella acuminata (Houtt.) Lév.; --Cyclosorus acuminatus (Houtt.) Ching.
2. Christella arida (D Don) Holttum; --Cyclosorus aridus (D Don) Ching. Photos
3. Christella dentata (Forssk.) Brownsey & Jermy.; --Cyclosorus dentatus sensu Copel., 1960, pro parte. --Cyclosorus mindanaensis (Christ) Copel. Pantropic. Throughout the Philippines. Terrestrial in exposed sites, light forests, along stream embankments, etc. Common at low altitudes.
4. Christella hispidula (Decne.) Holttum. --Cyclosorus falcatulus (Christ) Copel.
5. Christella parasitica (L.) Lév. --Cyclosorus dentatus sensu Copel. 1960, pro parte. --Cyclosorus benguetensis Copel. --Cyclosorus procurrens (Mett.) Copel. SE Asia (confined here?, or more widely distributed?). Throughout the Philippines. Terrestrial, open sites and degraded habitats generally, along stream embankments. Altitude?
6. Christella subdentata Holttum.
7. Christella subpubescens (Blume) Holttum.; --Cyclosorus subpubescens (Blume) Ching. --Cyclosorus latipinna (Hook. & Baker) Tard.

Coryphopteris Holttum

1. Coryphopteris borealis Holttum, Blumea 23(1): 35 (1976). 
2. Coryphopteris dura (Copel.) Holttum, Blumea 23(1): 44 (1976); --Lastrea dura (Copel.) Copel.
3. Coryphopteris pubirhachis (Baker) Holttum var. philippinensis Holttum, Blumea 23(1): 37 (1976). --Lastrea viscosa sensu Copel. (1960) non (Baker) Bedd., pro specim. philipp. Photos
4. Coryphopteris squamipes (Copel.) Holttum, Blumea 23(1): 35 (1976); --Lastrea squamipes (Copel.) Copel.

Excluded

1. Coryphopteris viscosa (J Sm.) Holttum, Blumea 19: 33. (1971).

Cyclogramma Tagawa = Cyclosorus

Cyclosorus Link

1. Cyclosorus auriculatus (J.Sm.) C.M.Kuo, Taiwania 47(2): 171 (2002); --Cyclogramma auriculata (J Sm.) Ching. --Thelypteris simulans Ching.
2. Cyclosorus interruptus (Willd.) H Ito. --Cyclosorus gongylodes auctt., doubtfully based on Aspidium gongylodes Schkuhr. Photos

 Haplodictyum C.Presl. = Pronephrium, Sphaerostephanos

Macrothelypteris Ching

1. Macrothelypteris polypodioides (Hook.) Holttum. --Lastrea leucolepis C.Presl. Photos
2. Macrothelypteris torresiana (Gaudich.) Ching; --Lastrea torresiana (Gaudich.) T Moore. Photos

Mesophlebion Holttum [= Thelypteris?]

1. Mesophlebion crassifolium (Blume) Holttum, Blumea 22: 232 (1975); --Lastrea crassifolia (Blume) Moore.
2. Mesophlebion rufescens Holttum, Blumea 22: 230 (1975).

Metathelypteris Ching

1. Metathelypteris dayi (Bedd.) Holttum. --Thelypteris singalanensis sensu Holttum, non (Baker) Ching.
2. Metathelypteris gracilescens (Blume) Ching; --Lastrea gracilescens (Blume) Moore. --Lastrea calva (Copel.) Copel. Photos
3. Metathelypteris uraiensis (Rosenst.) Ching.

Nannothelypteris Holttum [= Cyclosorus?]

1. Nannothelypteris aoristisora (Harrington) Holttum; --Cyclosorus aoristisorus (Harrington) Copel.
2. Nannothelypteris camarinensis Holttum.
3. Nannothelypteris inaequilobata Holttum. --Lastrea nervosa sensu Copel. (1960), pro parte. Photos
4. Nannothelypteris nervosa (Fée) Holttum; --Lastrea nervosa (Fée) Copel., pro parte.
5. Nannothelypteris philippina (C.Presl.) Holttum. --Lastrea confusa (Copel.) Copel.

Parathelypteris (H Ito) Ching emend. Holttum

1. Parathelypteris beddomei (Baker) Ching. Photos
   • var. beddomei; --Lastrea beddomei (Baker) Bedd.
   • var. eugracilis (Copel.) Holttum; --Lastrea eugracilis Copel. 
2. Parathelypteris glanduligera (Kunze) Ching
3. Parathelypteris grammitoides (Christ) Holttum; --Lastrea grammitoides (Christ) Copel.

Phegopteris Fée

1. Phegopteris decursive-pinnata Fée, Mém. Foug., 5. Gen. Filic. 242 (1850) t. 20 A. f. 1. 1850-52. Known only from one collection on Mt. Iraya, Batan I., Batanes (Barcelona 990).

Plesioneuron (Holttum) Holttum

1. Plesioneuron savaiense (Baker) Holttum. --Lastrea quadriaurita (Christ) Copel. Photos

Pneumatopteris Nakai

1. Pneumatopteris costata (Brackenr.) Holttum. --Lastrea cavitensis Copel.
2. Pneumatopteris glabra (Copel.) Holttum; --Cyclosorus glaber Copel. --Cyclosorus viridis Copel. Photos
3. Pneumatopteris laevis (Mett.) Holttum. --Cyclosorus nitidulus sensu Copel. (1960), pro parte. --Cyclosorus microlonchus sensu Copel. (1960), pro parte. Photos
4. Pneumatopteris ligulata (C.Presl.) Holttum; --Lastrea ligulata C.Presl. --Lastrea foxii (Christ) Copel. Photos 
5. Pneumatopteris lithophila Holttum.
6. Pneumatopteris michaelis Holttum.
7. Pneumatopteris microloncha (Christ) Holttum; --Cyclosorus microlonchus (Christ) Copel.
8. Pneumatopteris nitidula (C.Presl.) Holttum; --Cyclosorus nitidulus (C.Presl.) Copel. (1960), pro parte. Photos
9. Pneumatopteris truncata (Poir.) Holttum; --Cyclosorus truncatus (Poir.) Farw. Photos

Pronephrium C.Presl.

1. Pronephrium amphitrichum Holttum.
2. Pronephrium asperum (C.Presl.) Holttum. --Cyclosorus philippinarum (Fée) Copel.
3. Pronephrium bakeri (Harrington) Holttum; --Haplodictyum bakeri (Harrington) Ching; --Cyclosorus bakeri (Harrington) Copel.
4. Pronephrium balabacensis PM Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp. 32  (1980) 45, t. 2. Possibly synonymous with Pronephrium merrilliii. The exindusiate condition of the sori mentioned in the original description may actually be due to small, fugaceous indusia.
5. Pronephrium bulusanicum (Holttum) Holttum; --Haplodictyum bulusanicum Holttum.
6. Pronephrium clemensiae (Copel.) Holttum. Photos
   • var. clemensiae; --Cyclosorus clemensiae (Copel.) Ching.
   • var. degenerum (Christ) Holttum; --Cyclosorus degener (Christ) Copel. 
7. Pronephrium cuspidatum (Blume) Holttum; --Cyclosorus cuspidatus (Blume) Copel.
8. Pronephrium granulosum (C.Presl.) Holttum; --Cyclosorus granulosus (C.Presl.) Copel. --Cyclosorus subdimorphus Copel.
9. Pronephrium gymnopteridifrons (Hayata) Holttum. --Cyclosorus pustulosus Copel.
10. Pronephrium heterophyllum (C.Presl.) Holttum; --Haplodictyum heterophyllum C.Presl.
11. Pronephrium hosei (Baker) Holttum; --Cyclosorus hosei (Baker) Copel.
12. Pronephrium lineatum (Blume) Presl.
13. Pronephrium melanophlebium (Copel.) Holttum forma melanophlebium; --Cyclosorus melanophlebius (Copel.) Copel.
    forma proliferum Co, ined.
14. Pronephrium menisciicarpon (Blume) Holttum. --Cyclosorus latifolius (C.Presl.) Copel.
15. Pronephrium merrillii (Christ) Holttum; --Cyclosorus merrillii (Christ) Copel.
16. Pronephrium nitidum Holttum. --Cyclosorus urophyllus sensu Copel. (1960), excl. loc. Malacca
17. Pronephrium ramosii (Christ) Holttum var. ramosii. --Cyclosorus cuspidatus sensu Copel. (1960), excl. syn.; --Meniscium cuspidatum Blume. Photos
18. Pronephrium rhombeum (Christ) Holttum; --Cyclosorus rhombeus (Christ) Copel. --Cyclosorus lanceolus (Christ) Copel.
19. Pronephrium rubidum (J Sm. ex Hook.) Holttum; --Cyclosorus rubidus (Hook.) Copel.
20. Pronephrium samarense (Copel.) Holttum; --Cyclosorus samarensis Copel.
21. Pronephrium solsonicum Holttum.
22. Pronephrium trachyphyllum Holttum.
23. Pronephrium triphyllum (Sw.) Holttum; --Cyclosorus triphyllus (Sw.) Tard.
24. Pronephrium X xiphioides (Christ) Holttum; --Cyclosorus xiphioides (Christ) Copel. --Cyclosorus edanyoi Copel. --Cyclosorus rhombeus sensu Copel. (1960), pro parte. Photos

Pseudocyclosorus Ching [= Cyclosorus?]

1. Pseudocyclosorus tylodes (Kunze) Moore; --Lastrea tylodes (Kunze) Moore. ‘xylodes’ in Copeland’s Fern Flora.

Pseudophegopteris Ching

1. Pseudophegopteris aurita (Hook.) Ching.
2. Pseudophegopteris paludosa (Blume) Ching. --Lastrea pyrrhorhachis sensu Copel., Fern Fl. Philipp. (1960) 330, pro parte. Photos

Sphaerostephanos J Sm.

1. Sphaerostephanos alcasidii Holttum.
2. Sphaerostephanos acrostichoides (Desv.) Holttum. --Sphaerostephanos smithianus (C.Presl.) Holttum. --Cyclosorus boholensis Copel. --Cyclosorus productus sensu Copel. (1960), pro parte.
3. Sphaerostephanos angustifolius (C.Presl.) Holttum; --Cyclosorus angustifolius (C.Presl.) Copel.
4. Sphaerostephanos appendiculatus (Blume) Holttum
5. Sphaerostephanos cartilagidens PM Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp.  32 (1980) 43, t. 1.
6. Sphaerostephanos convergens Holttum.
7. Sphaerostephanos dichrotrichoides (Alderw.) Holttum; --Lastrea dichrotrichoides (Alderw.) Copel.
8. Sphaerostephanos diminutus (Copel.) MG Price; --Cyclosorus diminutus (Copel.) Copel.; --Pronephrium diminutum (Copel.) Holttum. --Sphaerostephanos lucbanii Holttum
9. Sphaerostephanos diversilobus (C.Presl.) Holttum; --Cyclosorus diversilobus (C.Presl.) Ching.
10. Sphaerostephanos ellipticus (Rosenst.) Holttum var. ellipticus. --Cyclosorus ellipticus (Rosenst.) Copel. var. glabrior Holttum
11. Sphaerostephanos erectus (Copel.) Holttum; --Cyclosorus erectus Copel.
12. Sphaerostephanos foxworthyi Holttum
13. Sphaerostephanos grandescens Holttum
14. Sphaerostephanos hernaezii Holttum
15. Sphaerostephanos heterocarpus (Blume) Holttum. Eight variants of this polymorphic species were enumerated (but no formal names given) by Holttum in the his FM revision. Two are definitely occurring in the Philippines, while another 2 resembles those ascribed to Borneo and New Guinea forms. Thus: -Mindanao form -Luzon form [as Cyclosorus bordenii (Christ) Ching in ‘Fern Flora’]. From Palawan is one collection resembling the Borneo form, while another one from San Ramon, Zamboanga is nearer to the New Guinea form. Photos
16. Sphaerostephanos hirsutus (Kunze ex Mett.) Holttum var. hirsutus. --Cyclosorus adenophorus (C Chr.) Ching.
17. Sphaerostephanos irayensis (Copel.) Holttum; --Plesioneuron irayense (Copel.) Holttum; --Cyclosorus irayensis Copel. --Sphaerostephanos fenixii Holttum.
18. Sphaerostephanos lastreoides (C.Presl.) Holttum. --Cyclosorus duplosetosus Copel. Photos
19. Sphaerostephanos latebrosus (Kunze ex Mett.) Holttum. --Sphaerostephanos todayensis (Christ) Holttum; --Cyclosorus todayensis (Christ) Copel. --Cyclosorus sagittifolioides Copel. --Cyclosorus halconensis Copel. --Cyclosorus polypterus Copel.
20. Sphaerostephanos lobatus (Copel.) Holttum; --Cyclosorus lobatus Copel. --Cyclosorus bordenii sensu Copel. (1960), pro parte.
21. Sphaerostephanos loherianus (Christ) Holttum; --Cyclosorus loherianus (Christ) Copel.
22. Sphaerostephanos magnus (Copel.) Holttum; --Cyclosorus magnus Copel.
23. Sphaerostephanos major (Copel.) Holttum; --Haplodictyum majus Copel. --Cyclosorus dimorphus Copel.
24. Sphaerostephanos norrisii (Rosenst.) Holttum. --Sphaerostephanos elmerorum (Copel.) Holttum; --Cyclosorus elmerorum Copel. --Cyclosorus ellipticus sensu Copel. (1960), pro parte.
25. Sphaerostephanos perglanduliferus (Alderw.) Holttum.
26. Sphaerostephanos polisianus Holttum.
27. Sphaerostephanos polycarpus (Blume) Copel. --Sphaerostephanos asplenioides J Sm.
28. Sphaerostephanos porphyricola (Copel.) Holttum; --Cyclosorus pophyricola (Copel.) Ching.
29. Sphaerostephanos productus (Kaulf.) Holttum; --Cyclosorus productus (Kaulf.) Ching. --Cyclosorus serratus Copel. --Cyclosorus weberi Copel. --Cyclosorus rigidus Copel. --Cyclosorus leucadenius Copel. --Sphaerostephanos punctatus Holttum. --Cyclosorus megaphyllus sensu Copel. (1960). --Cyclosorus nitidulus sensu Copel. (1960), pro parte.
30. Sphaerostephanos santomasii Holttum.
31. Sphaerostephanos sessilipinna (Copel.) Holttum; --Cyclosorus sessilipinna (Copel.) Ching. --Cyclosorus austrophilippinus (Copel.) Copel.
32. Sphaerostephanos simplicifolius (J Sm. ex Hook.) Holttum; --Cyclosorus simplicifolius (J Sm.) Copel.
33. Sphaerostephanos spenceri (Christ) Holttum; --Cyclosorus spenceri (Christ) Copel.
34. Sphaerostephanos stenodontus (Copel.) Holttum; --Cyclosorus stenodontus Copel.
35. Sphaerostephanos subcordatus Holttum.
36. Sphaerostephanos tephrophyllus (Copel.) Holttum; --Cyclosorus tephrophyllus (Copel.)
37. Sphaerostephanos unitus (L.) Holttum Photos
    • var. unitus; --Cyclosorus unitus (L.) Ching Sri Lanka, S India, across Malesia. Throughout the Philippines. Terrestrial. Degraded habitats generally, along waterways, etc. at low to medium altitudes. Very common.
    • var. mucronatus (Christ) Holttum.
38. Sphaerostephanos urdanetensis (Copel.) Holttum; --Cyclosorus urdanetensis Copel. --Cyclosorus matutumensis Copel.
39. Sphaerostephanos williamsii (Copel.) Holttum; --Lastrea williamsii Copel.

Status?

1. Sphaerostephanos mindorensis Holltum, Kalikasan 4 (1975) 54. Type: MG Price 690 (K,  holotype; isotype in PNH). Mindoro: Mt Halcon.  MINDORO: Mt Halcon, in forest, 750m. Known from two collections from the type locality. Not accounted for in Holltum’s Flora Malesiana revision.

Stegnogramma Blume [= Cyclosorus?]

1. Stegnogramma calcicola Co, sp. nov., ined. BOHOL.
2. Stegnogramma dissitifolia Holttum.
3. Stegnogramma gymnocarpa (Copel.) K Iwats. ssp. gymnocarpa; --Lastrea gymnocarpa Copel.
   

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Acknowledgements

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Last updated May 6, 2013