banner small

Piperaceae

Peperomia Ruiz & Pavon

Last edited by Pieter B. Pelser, 6 September 2017

Photos
  1. Peperomia agusanensis C.DC. in Elmer, LPB 6 (1914) 2294; --Merr., EPFP 2 (1923) 18; --Quisumb., PJS 43 (1930) 210; --Type: Elmer 13635 (PNH, holo, lost?; BISH, BM, CAS, DS, E, G, G-DC, GH, L, LY, MICH, MO, NY, P, U, US, W, Z, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta (Hilong-hilong), Aug-1912. Endemic to the Philippines. MINDANAO: Bukidnon, Agusan del Norte. Mossy forests, c. 1800m.
  2. Peperomia blanda (Jacq.) Kunth s.l., Nov. Gen. Sp. 1 (1816) 67; --Piper blandum Jacq., Collectanea 3 (1791) 210. --Peperomia recurvata var. philippinensis Miq., Lond. Journ. Bot. 4 (1845) 423; --Type: Cuming 1920 (K, holo), Philippines. Pantropical.
  3. Peperomia canlaonensis C.DC., Prodr. 5 (1910) Bot. 408; --Merr., EPFP 2 (1923) 18; --Quisumb., PJS 43 (1930) 200; --Type: Merrill Philip. Pl. 251 (G, holo; FR, G, M, PRC, U, US, Z, iso), Negros: Negros Occidental prov., Mt Kanlaon, Apr-1910. Endemic to the Philippines. NEGROS: Negros Occidental (Mt Kanlaon). Epiphytic in the mossy forests, c. 1800m.
  4. Peperomia copelandii Quisumb., PJS 43 (1930) 201; --Type: Copeland 178 (PNH), Luzon: Benguet prov., Pauai, May-1913. Endemic to the Philippines. LUZON: Benguet (Pauai), c. 2400m.
  5. Peperomia elmeri C.DC. in Elmer, LPB 3 (1910) 761; --Merr., EPFP 2 (1923) 19; --Quisumb., PJS 43 (1930) 216; --Type: Elmer 10493 (PNH, holo, lost?; BM, E, F, G, G-DC, GH, HBG, K, L, LY, MO, NY, U, US, W, Z, iso), Mindanao: Davao, Mt Apo, Todaya, ‘hanging from the under side of a thick inclined humus-covered forest trunk in moist forests at 4000ft’, May-1909. Endemic to the Philippines. MINDANAO: Davao (Mt Apo). Lower montane forest, c. 1200m, on tree trunks.
  6. Peperomia exigua (Blume) Miq., Syst. Pip. (1843) 101; --Merr., EPFP 2 (1923) 19; --Quisumb., PJS 43 (1930) 219; --Piper exiguum Blume, Verh. Bat. Genoots. 11 (1826) 232; --Type: from Java. --Peperomia hymenophylla Miq., Syst. Piperac. (1843) 78; --Type: Meyen s.n. (B, holo, lost; U, iso), Philippines. India, Malesia, tropical America. MINDANAO: Davao (Mt Apo, Todaya). Dense woods along Baracatan Creek, 500m, creeping and forming dense patches on wet ledges and cliffs.
  7. Peperomia laevifolia (Blume) Miq., Syst. Pip. (1843) 101; --Merr., EPFP 2 (1923) 19; --Quisumb., PJS 43 (1930) 207; --Piper laevifolium Blume, Cat. Gew. Buitenz. (1823) 33; --Type: from Java. --Peperomia apoana C.DC., PJS 5 c (1910) Bot. 411; --Merr., EPFP 2 (1923) 18; --Type: Copeland 1002 (G-DC, holo), Mindanao: Davao, Mt Apo. Christmas Isl, Indonesia, New Guinea, Philippines. PANAY: Antique (near Culasi), NEGROS: Negros Oriental (Cuernos Mtns), MINDANAO: Zamboanga (Sax River mtns), Bukidnon (Mt Lipa), Davao (Mt Apo). Montane forests to 2200m.
  8. Peperomia lagunaensis C.DC., PJS 5 c (1910) Bot. 407; --Merr., EPFP 2 (1923) 19; --Quisumb., PJS 43 (1930) 206; --Type: Merrill 5130 (G-DC, syn; K, NY, US, isosyn); BS 2446, 6079 (G-DC, syn); BS 6210 (G-DC, syn; US, isosyn), Luzon: Laguna prov., Mt Makiling, 17-Mar-1906. --Peperomia puberulifolia C.DC., PJS 5 c (1910) Bot. 407; --Merr., EPFP 2 (1923) 19; --Type: BS 6047 Foxworthy (PNH, holo, lost?), Luzon: Laguna prov., Mt Banahaw. Endemic to the Philippines. LUZON: Bataan (Mt Mariveles), Laguna (Mt Makiling; Mt Banahaw). Montane forests to 1700m, on mossy ground, boulders and tree trunks.
  9. Peperomia lanaoensis C.DC., PJS 5 c (1910) Bot. 410; --Merr., EPFP 2 (1923) 19; --Quisumb., PJS 43 (1930) 226; --Type: Clemens 625 (G-DC, holo; F, iso), Mindanao: Lanao del Norte prov., Maria Cristina Falls. Endemic to the Philippines. LUZON: Rizal (Mt Irid), SAMAR: Western Samar (Loquilocon), MINDANAO: Zamboanga, Lanao del Norte, Bukidnon (Tangkulan). Low and medium elevation, lithophytic on boulders, ledges of ravines and along streams.
  10. Peperomia latibracteata Quisumb., PJS 43 (1930) 224; --Type: BS 20126 R.C. McGregor (PNH, holo, lost?; G-DC, K, US, iso), Luzon: Nueva Vizcaya prov., Campote, 15-Jan-1913. Endemic to the Philippines. LUZON: Ifugao (Mt Polis), Nueva Vizcaya (Campote). Above 1900m in Ifugao.
  11. Peperomia marivelesana C.DC. in Elmer, LPB 3 (1910) 762; --Merr., EPFP 2 (1923) 19; --Quisumb., PJS 43 (1930) 220; --Type: Merrill 3721 (PNH, holo, lost?), Luzon: Bataan prov., Mt Mariveles; Elmer 6820 (G, K, NY, syn), Luzon: Bataan prov., Mt Mariveles, Nov-1904. Endemic to the Philippines. LUZON: Ilocos Norte (Mt Nagapatan), Nueva Vizcaya (Caraballo Mtns), Bataan (Mt Mariveles), Laguna (Mt Makiling), MINDORO: Mindoro Oriental (Mt Halcon), PALAWAN (Mt Victoria). Mossy rocks and tree trunks, 450-1200m.
  12. Peperomia merrillii C.DC., PJS 5 c (1910) Bot. 411; --Merr., EPFP 2 (1923) 19; --Quisumb., PJS 43 (1930) 212; --Peperomia bilineata var. merrillii (C.DC.) Quisumb., ined.; --Type: Merrill 4180 (G-DC, syn; K, NY, US, isosyn), Luzon: Cavite prov., Maragondon, Jul-1905; BS 1071 (G-DC, syn; NY, P, US, isosyn); For. Bur. 3359 (G-DC, syn; BR, K, NY, P, US, isosyn); Loher 4585 (K, US, isosyn). Endemic to the Philippines. LUZON: Rizal (Montalban; Mt Canumay; Mt Susong Dalaga; San Andales (possibly San Andres), Cavite (Maragondon), Laguna (Mt Makiling), Sorsogon (Mt Bulusan). Lithophytic at medium elevation to 500m.
  13. Peperomia mindoroensis C.DC., PJS 5 c (1910) Bot. 413; --Merr., EPFP 2 (1923) 19; --Quisumb., PJS 43 (1930) 215; --Type: Merrill 6107 & 6184 (PNH, syn, lost?), Mindoro: Mindoro Oriental prov., Mt Halcon. Endemic to the Philippines. MINDORO: Mindoro Oriental (Mt Halcon). Mossy trees, c. 1500m.
  14. Peperomia negrosensis C.DC. in Elmer, LPB 3 (1910) 760; --Merr., EPFP 2 (1923) 20; --Quisumb., PJS 43 (1930) 213; --Type: Elmer 9425 (G, holo; BISH, BM, E, L, NY, U, US, W, Z, iso), Negros: Negros Oriental prov., Cuernos Mtns, ‘tufted or climbing upon moss-covered rocks in well-shaded ravines’, Mar-1908. Endemic to the Philippines. NEGROS: Negros Oriental (Cuernos Mtns). Mossy rocks in shaded ravines. Photos
  15. Peperomia pallidibacca C.DC. in Elmer, LPB 3 (1910) 763; --Merr., EPFP 2 (1923) 20; --Quisumb., PJS 43 (1930) 199; --Type: Elmer 9344 (G, holo; BM, E, L, NY, US, iso), Luzon: Benguet prov., Baguio, Mar-1907. --Peperomia ramosii C.DC., Candollea 1 (1923) 315, nom. nud.; Candollea 2 (1925) 188; --Type: BS 12113 Ramos (G-DC, holo; BM, G-DC, K, L, US, iso), Luzon: Rizal prov., Tanay, San Isidro, Jun-1910. --Peperomia panaiana C.DC., Candollea 1 (1923) 330; --Type: McGregor 8461 (G-DC, holo; K, iso), Luzon. Endemic to the Philippines. LUZON: Mountain Province (Pinggat), Benguet (Mt Tonglon = Mt Sto Tomas; Baguio), Rizal (Montalban, Mt Lumutan; San Isidro). In the mossy forests, up to 2200m.
  16. Peperomia pellucidopunctulata C.DC. in Elmer, LPB 3 (1910) 760; --Merr., EPFP 2 (1923) 20; --Quisumb., PJS 43 (1930) 209; --Type: Elmer 8436 (G-DC, syn), Luzon: Benguet prov., Baguio; Elmer 9970 (G-DC, syn). Endemic to the Philippines. LUZON: Benguet (Baguio), Albay (Mt Mayon). On rocks and tree trunks at medium and high elevation.
  17. Peperomia recurvata (Blume) Miq., Syst. Pip. (1843) 141. Indonesia, Philippines. Photos
  18. Peperomia rubrivenosa C.DC., PJS 5 c (1910) Bot. 409; --Merr., EPFP 2 (1923) 21; --Quisumb., PJS 43 (1930) 225; --Type: R.S. Williams 1083 (PNH, holo, lost?; G-DC, drawing; NY, iso), Luzon: Benguet prov., Baguio. --Peperomia macgregorii C.DC., PJS 5 c (1910) Bot. 412; --Merr., EPFP 2 (1923) 19; --Type: BS 8380 (G-DC). --Peperomia rivulorum C.DC. in Elmer, LPB 3 (1910) 762; --Merr., EPFP 2 (1923) 21; --Quisumb., PJS 43 (1930) 228; --Type: Elmer 11147 (PNH, holo?, lost?; BISH, BM, E, G, GH, K, L, LY, MO, NY, US, W, Z, iso), Mindanao: Davao, Mt Apo, Todaya, ‘creeping and forming patches in thin earth of large, more or less moss-covered boulders lying in streamlets of dense woods near the Sibulan River, 1750ft’, Jul-1909. Endemic to the Philippines. LUZON: Benguet, MINDANAO: Davao (Catalonan, Mt Apo: Todaya). On trees and mossy boulders. Mossy forest, 2100m.
  19. Peperomia tetraphylla (G.Forst.) Hook. & Arn., The Botany of Captain Beechey’s Voyage 2 (1832) 97. --Peperomia reflexa (L.f.) A.Dietr., Sp. Pl. ed. 6 1 (1831) 180, nom. illeg., auct. non Kunth. --Peperomia reflexa (L.f.) A.Dietr. var. capensis (Miq.) C.DC., Prodr. 16, 1 (1869) 451; --Quisumb., PJS 43 (1930) 196; --Peperomia reflexa (L.f.) A.Dietr. forma capensis Miq., Syst. Pip. (1843) 169; --Merr., EPFP 2 (1923) 20. --Peperomia reflexa (L.f.) A.Dietr. forma subsessilifolia C.DC., Prod. 16 (1869) 452; --Merr., EPFP 2 (1923) 20; --Type: Elmer 8576 (BISH, iso), Luzon: Benguet prov., Baguio, Mar-1907. --Peperomia reflexa (L.f.) A.Dietr. var. (or forma?) calcicola C.DC., --PJS 5 c (1910) Bot. 410; --Merr., EPFP 2 (1923) 20; --Type: Elmer 6077 (PNH, holo?, lost?; US, iso), Luzon: Benguet prov., Baguio, Apr-1904. --Peperomia reflexa (L.f.) A.Dietr. var. parvilimba C.DC., --Quisumb., PJS 43 (1930) 198; --Type: Merrill 3918 (PNH, syn?, lost?), Luzon: Pampanga prov., Mt Arayat, on rocks near the summit, c. 1000m. Pantropical. LUZON: Mountain Province (Mt Data), Benguet (Pauai, Mt Pulag, Mt Lusong, Mt Sto Tomas, Baguio), Pampanga (Mt Arayat), MINDANAO: Bukidnon (Mt Candoon), Davao (Mt Apo).
  20. Peperomia tomentosa (Vahl) A.Dietr., Sp. Pl. 1 (1831) 172; --Piper tomentosum Vahl, Enum. Pl. 1 (1804) 351. --Peperomia tomentosa var. carnosa C.DC., Prodr. 161 (1869) 455; --Merr., EPFP 2 (1923) 21; --Quisumb., PJS 43 (1930) 222. Indonesia, New Guinea, Seychelles, ?Philippines.
  21. Peperomia ventenatii Miq., Syst. Piperac. (1843) 125; --Type: from Indonesia. Indonesia, Philippines.

Non-native:

  1. Peperomia argyreia (Hook.f.) E.Morren, Ann. Hort. Belge Étrangère 17 (1867) 2. An exotic species of American origin, cultivated for its variegated leaves. Photos
  2. Peperomia pellucida (L.) Humb., Bonpl. & Kunth, Nov. Gen. Sp. Pl. 1 (1815) 64; --Merr., Fl. Manila (1912) 170; EPFP 2 (1923) 20; --Quisumb., PJS 43 (1930) 218; --Piper pellucidum L., Sp. Pl. 1 (1753) 30. Native to the Americas. Photos

Piper L.

Last edited by Pieter B. Pelser, 7 September 2017

Photos
  1. Piper abbreviatum Opiz, in C.Presl, Rel. Haenk. 1 (1828) 157; --Merr., EPFP 2 (1923) 2; --Quisumb., PJS 43 (1930) 58; --Gardner, Blumea 51 (2007) 579; --Type: Haenke s.n. (PR, holo), Luzon. --Chavica populifolia Miq., Syst. Pip. (1843) 248; --Type: Cuming 834 (K, holo), Luzon: Sorsogon prov.. --Piper mearnsii C.DC., PJS 5 c (1910) Bot. 447; --Merr., EPFP 2 (1923) 12; --Type: BS 2987 Mearns (PNH, holo), Luzon: Aurora prov., Casiguran. --Piper miquelianum Fern.-Vill., Nov. App. (1880) 176; --Merr., EPFP 2 (1923) 17. --Piper parvispicum C.DC. in Perkins, Fragm. Fl. Philip. (1905) 156; --Type: Warburg 14750 (B, holo), Mindanao: Davao del Sur prov., Mt Dagatpan. --Piper rhombophyllum C.DC., Prod. 16 (1869) 352; --Merr., EPFP 2 (1923) 14. --Piper rubripunctulatum C.DC. in Perkins, Fragm. Fl. Philip. (1905) 158; --Type: Warburg 14746 (B, holo), Mindanao: Davao del Sur prov., Sta Cruz. Philippines to Solomon Isls and Australia. LUZON: Cagayan (Enrile, Peñablanca), Aurora (Casiguran), Laguna (San Antonio, Calauan, Mt Makiling), Quezon (Mt Binuang, Kabibihan), Camarines Norte (Paracale), Camarines Sur (Sagnay, Mt Isarog), Albay, Sorsogon (Mt Bulusan), POLILLO. 
  2. Piper agusanense C.DC. in Elmer, LPB 6 (1914) 2291; --Merr., EPFP 2 (1923) 2; --Quisumb., PJS 43 (1930) 50; --Type: Elmer 13319 (BISH, NY, US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta = Mt Hilong-hilong, Cabadbaran. LUZON: Laguna (Mt Makiling), CAMIGUIN, MINDANAO: Agusan del Norte (Mt Hilong-hilong), Surigao del Norte (Placer). In forest.
  3. Piper angustipeltatum Merr., PJS 17 (1920) 244; EPFP 1 (1922) 3; --Quisumb., PJS 43 (1930) 98; --Type: BS 28846 Ramos & Edaño (PNH, holo, lost?; US, iso), Luzon: Quezon prov., Mt Binuang, damp forests along rivers, 19-May-1917. LUZON: Quezon (Mt Binuang). Low and medium elevation forests along rivers.
  4. Piper aristolochiphyllum Quisumb., PJS 43 (1930) 54; --Type: BS 45695 Ramos & Edaño (PNH, holo, lost?; NY, iso), Luzon: Aurora prov., Mt Alzapan, on forested slopes, c. 1500m, 6-Jun-1925. LUZON: Aurora (Mt Alzapan).
  5. Piper asterostigmum Quisumb., PJS 43 (1930) 91; --Type: C.A. Wenzel 3121 (UC, holo; NY, iso), Mindanao: Surigao del Norte prov., Placer, in forests, c. 150m, 5-Sep-1927. MINDANAO: Surigao del Norte (Placer). Lowland forest.
  6. Piper aurilimbum C.DC. in Elmer, LPB 3 (1910) 768; --Merr., EPFP 1 (1922) 3; --Quisumb., PJS 43 (1930) 55; --Type: Elmer 8866 (MO, NY, US, iso), Luzon: Benguet prov., Baguio, Mar-1907. LUZON: La Union (Disdis), Kalinga (Mt Masingit), Ifugao (Mt Polis), Benguet (Baguio), Rizal. Medium elevation forests.
  7. Piper baccatum Blume, Verh. Bat. Genoots. 11 (1826) 172; --Merr., EPFP 1 (1922) 4; --Quisumb., PJS 43 (1930) 173; --Gardner, Blumea 51 (2007) 579; --Type: from Java. --Piper sarcopodum C.DC., PJS 11 c (1916) Bot. 207; --Merr., EPFP 2 (1923) 4; --Quisumb., PJS 43 (1930) 175; --Type: BS 16536 Ramos (K, NY, US, iso), Luzon: Laguna prov., San Antonio, forests near streams, Sep/Oct-1912. Borneo, India, Java, Singapore, Sri Lanka, Thailand, Philippines. LUZON: Laguna (Mt Makiling; San Antonio), Sorsogon (Mt Bulusan, Irosin), MINDANAO: Lanao del Sur (Camp Keithley), Agusan del Norte (Mt Hilong-hilong). Low and medium elevation forests. Photos
  8. Piper baguionum C.DC. in Elmer, LPB 3 (1910) 775; --Merr., EPFP 1 (1922) 4; --Quisumb., PJS 43 (1930) 96; --Type: Elmer 8784 (BISH, MO, NY, US, iso), Luzon: Benguet prov., Baguio, Mar-1907. --Piper polycladum C.DC., PJS 5 c (1910) Bot. 438; --Merr., EPFP 2 (1923) 14; --Type: R.S. Williams 1134 (PNH, holo, lost?; NY 2 sheets, US, iso), Luzon: Benguet prov., Baguio, 15-Jun-1904. LUZON: Benguet (Baguio).Montane forests and thickets, 1200-1500m.
  9. Piper begoniaefolium (Blume) Quisumb., PJS 43 (1930) 189; --Zippelia begoniaefolia Blume in Roem. & Schultes, Syst. Veg. 7 (1830) 1614, 1615; --Merr., EPFP 2 (1923) 18; --Gardner, Blumea 51 (2007) 584; --Type: from Java. Indochina, Sumatra, Peninsular Malaysia, Java, Borneo, SW Philippines. JOLO. Damp forest, c. 700m.
  10. Piper betle L., Sp. Pl. (1753) 28; --Merr., Fl. Manila (1912) 170; EPFP 1 (1922) 3; --Quisumb., PJS 43 (1930) 85; --Gardner, Blumea 51 (2007) 579. --Piper anisodorum Blanco, Fl. Filip. ed. 2 (1845) 16; Fl. Filip. ed. 3, 1 (1877) 31; --Piper anisum-olens Blanco, Fl. Filip. (1837) 23; --Type: non extant; 'Illustrative specimen': Merrill Sp. Blancoanae 788 (US), Luzon: Rizal prov., Antipolo, Jan-1915. --Piper bathycarpum C.DC. in Perkins, Fragm. Fl. Philip. (1905) 153; EPFP 2 (1923) 4, as ‘bathicarpum’; --Type: Warburg 14844, Jolo. --Piper puberulinodum C.DC. in Elmer, LPB 3 (1910) 773; --Merr., EPFP 2 (1923) 14; --Type: Elmer 11972 (BISH, MO, US), Mindanao: Davao del Sur prov., Mt Apo, Todaya, Oct-1909. --Piper carnistylum C.DC. in Elmer, LPB 3 (1910) 774; --Merr., EPFP 2 (1923) 6; --Type: Elmer 8904, Luzon: Quezon prov., Lucban. --Piper betle var. fenixii (C.DC.) Quisumb., PJS 43 (1930) 89; --Piper fenixii C.DC., PJS 5 c (1910) Bot. 425; --Merr., EPFP 2 (1923) 8; --Type: BS 3652 E. Fenix (US, iso), Batanes Isls, May-Jun-1907. --Piper betle var. macgregorii (C.DC.) Quisumb., PJS 43 (1930) 89; --Piper macgregorii C.DC., PJS 11 c (1916) Bot. 215; --Merr., EPFP 2 (1923) 11; --Type: BS 18491 R.C. McGregor, Biliran: Biliran prov., in forest, c. 300m. --Piper betle var. densum (Blume) C.DC., Prod. 16 (1869) 360. --Piper malarayatensis C.DC., PJS 5 c (1910) Bot. 442; --Merr., EPFP 2 (1923) 11. --Piper philippinense C.DC., Prodr. 16, 1 (1869) 353; --Fern.-Vill., Novis. App. (1880) 175; --S.Vidal, Phan. Cuming. Philip. (1885) 138; Rev. Pl. Vasc. Filip. (1886) 219; --Type: Cuming 485, Luzon: Laguna prov., Calauan. Widely cultivated in Asia and through Malesia to the Solomon Isls and Fiji. BATAN: Basco, LUZON: Ilocos Norte (Burgos), Ilocos Sur (Cervantes), Pangasinan (San Quintin), Isabela (Malunu), Nueva Vizcaya (Dupax), Aurora (Casiguran; Umiray), Tarlac (Bamban), Bataan (Mt Mariveles, Lamao River; Dinalupihan), Bulacan (Norzagaray; Angat; Galas), Rizal (Antipolo; Morong), Cavite (Mendez; Alfonso), Laguna (Los Baños; Calauan; Mt San Cristobal; San Antonio; Paete), Quezon (Mt Binuang; Lucban; Guinayangan), Camarines Sur (Sagnay; Camaligan), ALABAT, MINDORO: Mindoro Oriental (Pinamalayan), BOHOL: Bohol (Sevilla), BILIRAN, LEYTE: Leyte (Dagami; Tigbao), TAWI-TAWI, JOLO, MINDANAO: Agusan del Norte (Mt Hilong-hilong), Surigao del Norte (Surigao; Placer), Davao del Sur (Mt Apo, Todaya), Davao Oriental (Mati). Occurring wild but also extensively cultivated throughout the Archipelago. Gardens, old clearings and secondary forests, at low and medium elevation up to c. 1400m. Betel pepper, Ikmo. Photos
  11. Piper breviamentum C.DC., PJS 5 c (1910) Bot. 434; --Merr., EPFP 2 (1923) 4; --Quisumb., PJS 43 (1930) 62; --Type: R.S. Williams 2104 (PNH, holo, lost?; NY 3 sheets, US, iso), Mindanao: Zamboanga, Sax River, 4-Feb-1905. --Piper copelandii C.DC., PJS 5 c (1910) Bot. 447; --Merr., EPFP 2 (1923) 15; --Type: Copeland 1298 (PNH, holo, lost?; US, iso), Mindanao: Davao del Sur prov., Mt Apo, Todaya, Apr-1904. --Piper sarcostylum C.DC., PJS 11 c (1916) Bot. 216; --Merr., EPFP 2 (1923) 15; --Type: Merrill 7305 (US, iso), Mindanao: Agusan del Norte prov., Agusan River, Oct-1910.
  12. Piper brevicuspe (Miq.) Merr., EPFP 2 (1923) 5; --Quisumb., PJS 43 (1930) 26; --Rhyncholepis brevicuspis Miq., Syst. Pip. (1843) 283; --Type: Cuming 1843, Bohol: Bohol prov. --Piper rhyncholepis C.DC., Prodr. 16, 1 (1869) 344, as ‘rhyncholepsis’; PJS 5 c (1910) Bot. 209; Candollea 1 (1923) 212; --Fern.-Vill., Novis. App. (1880) 176; --S.Vidal, Phan. Cuming. Philip. (1885) 138; Rev. Pl. Vasc. Filip. (1886) 220. --Rhyncholepis cumingiana Miq., Syst. Pip. (1843) 282 (non Piper cumingiana Miq.); --Type: Cuming 1697, Samar. BOHOL (Valencia; Kalingohan), LEYTE: Leyte (Tingib; Dagami, Panda; Jaro, Buenavista), SAMAR: Yaborog; Northern Samar (Paranas), Western Samar (Loquilocon), MINDANAO: Agusan del Norte (Mt Urdaneta = Mt Hilonghilong), Surigao del Norte (Surigao; Placer), Davao Oriental (Mt Mayo). Low and medium elevation damp forests.
  13. Piper brevistigmum C.DC. in Elmer, LPB 6 (1914) 2292; --Merr., EPFP 1 (1922) 5; --Quisumb., PJS 43 (1930) 148; --Type: Elmer 13684 (BISH, NY US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta = Mt Hilong-hilong, in forest, c. 1000m, Sep-1912. Mindanao: Agusan del Norte (Mt Urdaneta = Mt Hilong-hilong), in forest, c. 1000m.
  14. Piper cacuminum C.DC. in Elmer, LPB 3 (1910) 765; --Merr., EPFP 1 (1922) 5; --Quisumb., PJS 43 (1930) 68; --Type: Elmer 6890 (PNH, holo, lost?), Luzon: Bataan prov., Mt Mariveles, in the mossy forest near summit. LUZON: Zambales (Mt Tapulao), Bataan (Mt Mariveles). Montane mossy forests, 1100-1500m.
  15. Piper caninum Blume, Verh. Bat. Genoots. 11 (1826) 214; --Merr., EPFP 1 (1922) 5; --Quisumb., PJS 43 (1930) 120; --Gardner, Blumea 51 (2007) 580; --Type: from Java. --Piper caninum var. basilanum (C.DC.) Quisumb., PJS 43 (1930) 124; --Piper basilanum C.DC., PJS 5 c (1910) Bot. 457; --Merr., EPFP 2 (1923) 4; --Type: Hallier s.n. (PNH, holo, lost?), Basilan, Jan-1904. --Piper caninum var. glabribracteum C.DC., PJS 5 c (1910) Bot. 459; --Merr., EPFP 2 (1923) 5; --Quisumb., PJS 43 (1930) 121; --Type: Clemens 176 (PNH, holo, lost?), Mindanao: Lanao del Sur prov., Lanao Lake, Camp Keithley. --Piper caninum var. hallieri (C.DC.) Quisumb., PJS 43 (1930) 123; --Piper hallieri C.DC., PJS 5 c (1910) Bot. 458; --Merr., EPFP 2 (1923) 9; --Type: Hallier s.n. (PNH, holo, lost?), Basilan, Jan-1904. --Piper caninum var. lanaoense C.DC., PJS 5 c (1910) Bot. 459; --Merr., EPFP 2 (1923) 5; --Quisumb., PJS 43 (1930) 122; --Type: Clemens s.n. (PNH, holo, lost?), Mindanao: Lanao del Sur prov., Lake Lanao, Camp Keithley, Sep-1906. --Piper caninum var. latibracteum C.DC. in Elmer, LPB 3 (1910) 787; --Merr., EPFP 2 (1922) 6; --Quisumb., PJS 43 (1930) 126; --Type: Elmer 7627 (BISH, MO, syn); --Elmer 7990 (NY, syn), Luzon: Quezon prov., Lucban, May-1907; --Elmer 8844 (MO, NY, syn), Luzon: Benguet prov., Baguio, Mar-1907. --Piper caninum var. oblongifolium Quisumb., PJS 43 (1930) 122; --Type: BS 35070 Ramos & Pascasio (PNH, holo, lost?), Bucas Grande, in open forests at low elevation, 8-Jun-1919. --Piper acutibaccum C.DC., PJS 5 c (1910) Bot. 459; --Merr., EPFP 2 (1923) 2; --Quisumb., PJS 43 (1930) 146; --Type: BS 10031 Ramos (PNH, holo, lost), Luzon: Laguna prov., San Antonio. --Piper apoanum C.DC. in Elmer, LPB 3 (1910) 785; --Merr., EPFP 1 (1922) 3; --Quisumb., PJS 43 (1930) 131; --Type: Elmer 11174 (MO, NY, US, iso), Mindanao: Davao del Sur prov., Mt Apo, Todaya, in bamboo thickets near Sibulan River, c. 800m, Jul-1909. --Piper arborisedens C.DC., PJS 11 c (1916) Bot. 223; --Merr., EPFP 2 (1923) 3; --Quisumb., PJS 43 (1930) 145; --Type: BS 24934 Ramos (NY, US, iso), Luzon: Laguna prov., San Antonio, in forest, 20-Aug-1910. --Piper cabadbaranum C.DC. in Elmer, LPB 6 (1914) 2292; --Merr., EPFP 1 (1922) 5; --Quisumb., PJS 43 (1930) 143; --Type: Elmer 14136 (NY, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta = Mt Hilong-hilong, wind-swept steep forested slopes of Cawilanan Peak, 4000ft, Oct-1912. --Piper dagatpanum C.DC. in Perkins, Fragm. Fl. Philip. (1905) 154; --Merr., EPFP 2 (1923) 7; --Quisumb., PJS 43 (1930) 132; --Type: Warburg 14739 (B, holo), Mindanao: Davao del Sur prov., Mt Dagatpan, Forests, c. 1300m. --Piper haenkeanum Opiz in C.Presl, Rel. Haenk. 1 (1828) 159; --Merr., EPFP 2 (1923) 9; --Quisumb., PJS 43 (1930) 138; --Chew, Blumea 20 (1972) 147; --Type: Haenke s.n. --Piper malalaganum C.DC., PJS 5 c (1910) Bot. 461; --Merr., EPFP 2 (1923) 11; --Quisumb., PJS 43 (1930) 137; --Type: Copeland 696 (PNH, holo, lost?), Mindanao: Davao del Sur prov., Malalag. --Piper marivelesanum C.DC. in Perkins, Fragm. Fl. Philip. (1905) 155; --Merr., EPFP 2 (1923) 11; --Type: Warburg 13640, Luzon: Bataan prov., Mt Mariveles. --Piper merrittii C.DC., PJS 5 c (1910) Bot. 460; --Merr., EPFP 2 (1923) 12; --Type: FB 6138 Merritt (PNH, holo, lost?), Mindoro: Mindoro Oriental prov., Balete. --Piper negrosense C.DC. in Elmer, LPB 3 (1910) 786; --Merr., EPFP 2 (1923) 17; --Type: Elmer 9482 (MO, NY, US, iso), Negros: Negros Oriental prov., Cuernos Mtns, ‘climber upon low trees and over large boulders in the hemp region at 3000ft’, Mar-1908. --Piper obovatibracteum C.DC. in Elmer, LPB 3 (1910) 784; --Merr., EPFP 2 (1923) 13; --Type: Elmer 7927 (BISH, L, NY, US, iso), Luzon: Quezon prov., Lucban, in woods at 800m, May-1907. --Piper parcipilum C.DC., PJS 5 c (1910) Bot. 445; --Type: M.S. Clemens s.n., Mindanao: Lanao del Sur prov., Lake Lanao, Camp Keithley. --Piper perpunctatum C.DC., PJS 11c (1916) Bot. 219; --Merr., EPFP 2 (1923) 13; --Type: Merrill 7246 (US, iso), Palawan: Palawan prov., Malampaya, Sep-1910. --Piper sablanum (C.DC.) Quisumb., PJS 43 (1930) 135; --Piper caninum Blume var. sablanum C.DC., PJS 5 c (1910) Bot. 459; --Merr., EPFP 2 (1923) 6; --Type: Elmer 6150 (PNH, holo, lost?; NY, US, iso), Luzon: Benguet prov., Sablan, Apr-1904. --Piper tenuipedunculum C.DC., PJS 5 c (1910) Bot. 460; --Merr., EPFP 2 (1923) 16; --Quisumb., PJS 43 (1930) 136; --Type: R.S. Williams 2343 (PNH, holo, lost?; NY 2 sheets, iso), Mindanao: Zamboanga prov., San Ramon, in forests, c. 400ft, 3-Feb-1905. --Piper tenuirameum C.DC. in Perkins, Fragm. Fl. Philip. (1905) 159; --Merr., EPFP 2 (1923) 16; --Type:   Warburg 13317, Luzon: Rizal prov., Binangonan. --Piper villilimbum C.DC. in Elmer, LPB 3 (1910) 788; --Merr., EPFP 2 (1923) 17; --Type: Elmer 7382 (L, MO, US, syn), Luzon: Quezon prov., Lucban, May-1907; --Elmer 7624 (L, MO, NY, syn), Luzon: Quezon prov., Lucban, in woods at 2500ft, May-1907. --Piper viminale Opiz in C.Presl, Rel. Haenk. 1 (1828) 150; --Merr., EPFP 2 (1923) 17; --Quisumb., PJS 43 (1930) 126. W Malesia to the Solomon Isls and Australia. LUZON: Pangasinan (Mt San Isidro Labrador), Abra (Mt Posuey), Benguet (Baguio; Sablan), Cagayan (Peñablanca), Isabela (San Mariano), Aurora (Casiguran), Nueva Ecija (Mt Umingan = Mt Mingan), Zambales (Subic), Pampanga (Mt Pinatubo), Bataan (Dinalupihan; Limay; Mt Mariveles), Bulacan (Angat), Rizal (Binangonan; Bosoboso), Laguna (near Famy; Siniloan-Infanta Road; San Antonio; Paete), Quezon (Mt Tulaog; Dolores; Lucban; Lucban-Mauban Road; Tagkawayan), Camarines Norte (Niog, Sagnay; Paracale), Camarines Sur (Maagnas; Mt Isarog), Sorsogon (Mt Bulusan; Mt Labao), POLILLO (Mt Malulud; Barrio Saloang), CATANDUANES (Mt Mariguidon; Sto Domingo River), MINDORO: Mindoro Occidental (Mt Calavite; Paluan), Mindoro Oriental (Balete; Baco River; Mt Halcon; Naujan; Pinamalayan; Bongabong River; Camantigue), BUSUANGA, PALAWAN (Malampaya; Lake Manguao), SIBUYAN (Mt Giting-giting, Magallanes), PANAY: Antique (Culasi), Capiz (Jamindan; Galecia; Mt Kinablangan), Aklan (Libacao), NEGROS: Negros Oriental (Cuernos Mtns), BOHOL (Kalingohan; Bilar), LEYTE: Leyte (Dagami; Tacloban, Tigbao; Jaro, Buenavista), SAMAR (Cauayan Valley; Mt Canislagan; Camaniwan, Catubig River), CAMIGUIN (Mambajao; Mt Hibok-hibok; Mt Mahilog), JOLO (Bud-wak), BASILAN, MINDANAO: Zamboanga (San Ramon), Zamboanga del Sur (Malangas), Lanao del Sur (Camp Keithley), Bukidnon (Tangkulan), Davao, Davao del Sur (Mt Apo; Mt Dagatpan; Malalag), Agusan del Norte (Mt Hilong-hilong), Agusan del Sur (Talicogon), Surigao del Norte (Surigao; Placer), SIARGAO (Dapa), BUCAS GRANDE. Lowland and medium elevation forests and thickets, also along streambanks, ascending to 1300m. Conservation status: Other Threatened Species (DAO 2017-11). Photos
  16. Piper celtidiforme Opiz in C.Presl, Rel. Haenk. 1 (1828) 152; --Merr., EPFP 2 (1922) 6; --Quisumb., PJS 43 (1930) 177; --Gardner, Blumea 51 (2007) 580; --Type: Haenke s.n. (PR, holo), Luzon. --Piper celtidiforme forma luzonense (C.DC.) Quisumb., PJS 43 (1930) 180; --Type: Jagor 722 (B, holo), Luzon. --Piper celtidiforme forma tubuanense Quisumb., PJS 43 (1930) 182; --Type: BS 36704 Ramos & Edaño (PNH, holo, lost?), Mindanao: Zamboanga del Sur prov., Mt Tabuan. --Piper celtidiforme forma usterii (C.DC.) Quisumb., PJS 43 (1930) 181; --Piper usterii C.DC., Beitr. Kenntn. Philipp. (1905) 125; --Type: Usteri s.n. (PNH, holo; lost?), Guimaras. --Piper celtidiforme var. vaginans Quisumb., PJS 43 (1930) 182; --Type: BS 19822 R.C. McGregor (PNH, holo, lost?), Luzon: Ifugao prov., Mt Polis. --Piper catubigense Merr., PJS 17 (1920) 244; EPFP 2 (1922) 6; --Quisumb., PJS 43 (1930) 183; --Type: BS 24278 Ramos (PNH, holo, lost?; NY, US, iso), Samar: Catubig River, on trees in damp forests at low elevation, 13-Mar-1916. --Piper corylistachyon (Miq.) C.DC., Prodr. 16, 1 (1869) 346; PJS 5 c (1910) Bot. 438; PJS 11 c (1916) Bot. 218; LPB 3 (1910) 776; Candollea 1 (1923) 188; --Fern.-Vill., Novis. App. (1880) 175; --S.Vidal, Phan. Cuming. Philip. (1885) 138; Rev. Pl. Vasc. Filip. (1886) 219; --Merr., Sp. Blancoanae (1918) 118; --Chavica corylistachya Miq., Syst. Pip. (1843) 281; --Type: Cuming 1044 (K, holo; L, iso), Philippines. --Piper corylistachyon formae d and d2 C.DC., PJS 5 c (1910) Bot. 439; --Type: Gregory 110, Luzon: Quezon prov., Atimonan. --Piper longilimbum C.DC., PJS 11 c (1916) Bot. 221; --Merr., EPFP 2 (1923) 11; --Type: BS 17602 Ramos (US, iso), Samar: Mt Canislagan, Mar/Apr-1914. --Piper penninerve C.DC. in Perkins, Fragm. Fl. Philip. (1905) 157; --Merr., EPFP 2 (1923) 13; --Quisumb., PJS 43 (1930) 185; --Type: Warburg 14744 (B, holo), Mindanao: Davao del Sur prov., Mt Dagatpan. --Piper villirache C.DC., PJS 11 c (1916) Bot. 214; --Merr., EPFP 2 (1923) 17; --Quisumb., PJS 43 (1930) 187; --Type: BS 15773 E. Fenix (US, iso), Mindanao: Bukidnon prov., Sumilao, Aug-1912. --Piper warburgii C.DC. in Perkins, Fragm. Fl. Philip. (1905) 159; Candollea 1 (1923) 285; --Type: Warburg 13116 (B, holo), Luzon: Quezon prov., Sampaloc. Philippines, New Guinea, Solomon Isls. LUZON: Ilocos Norte (Bangui to Claveria), Ilocos Sur, Pangasinan (Umingan), Apayao (Abulug River), Kalinga (Mt Masingit), Ifugao (Mt Polis), Aurora (Casiguran), Nueva Ecija (Carranglan), Zambales (Anuling), Bataan (Mt Mariveles), Pampanga (Mt Abu; Mt Arayat), Rizal (Sumag; Mt Angilog; Antipolo; Calauis River; Montalban; Binangonan; Bosoboso; Tanay, San Isidro; Mt Irid), Laguna (Los Baños; Mt Makiling; Sta Cruz; Liliw, Pangil, Paete; San Antonio), Quezon (Infanta; Mt Binuang; Lucban; Atimonan; Sampaloc; Tagkawayan), POLILLO (Burdeos), Camarines Sur (Niog; Bagacay), Albay (Carcraray; Mt Mayon), Sorsogon (Mt Lalao; Mt Bulusan), CATANDUANES (Calolbong), MINDORO: Mindoro Occidental (Paluan), Mindoro Oriental (Lake Naujan; Cauayan; Pinamalayan), MARINDUQUE, PANAY: Antique (Culasi; Tibiao), Capiz (Mt Madia-as; Mt Bomlon; Jamindan; Agraman, Locero), Iloilo (Suague River; Tigom River), GUIMARAS, NEGROS, CEBU (Catmon), BOHOL (Bilar), BILIRAN, LEYTE: Leyte (Tigbao; Jaro, Buenavista; Dagami), Southern Leyte (Malitbog), SAMAR: (Lanang; Catubig River), Western Samar (Catbalogan; Loquilocon), BASILAN, MINDANAO: Zamboanga del Sur (Malangas; Mt Tubuan; Port Banga), Davao del Sur (Mt Dagatpan), Davao Oriental (Mt Mayo; Baganga), Bukidnon (Sumilao), Agusan del Norte (Agusan River, Waloe), Surigao del Norte (Surigao; Placer), SIARGAO. Lowland to medium elevation forests and thickets generally, ascending to 1500m.
  17. Piper cordatilimbum Quisumb., PJS 43 (1930) 141; --Type: BS 42699 Ramos (PNH, holo, lost?), Bohol: Bohol prov., Bilar, Aug/Oct-1923. LUZON: Laguna (San Antonio), Quezon (Lucban), CATANDUANES (Mt Mariguidon), BOHOL (Bilar), LEYTE: Leyte (Jaro, Buenavista; Dagami), MINDANAO: Davao del Sur (Mt Apo, Todaya), Davao Oriental (Mt Mayo). Low and medium elevation forests, ascending to 1200m.
  18. Piper costulatum C.DC. in Elmer, LPB 3 (1910) 764; --Merr., EPFP 2 (1922) 7; --Quisumb., PJS 43 (1930) 66; --Type: Elmer 6805 (NY, iso), Luzon: Bataan prov., Mt Mariveles, Lamao River, Nov-1904. --Piper podandrum C.DC., PJS 5 c (1910) Bot. 436; --Merr., EPFP 2 (1923) 13; --Type: FB 8141 Curran & Merritt, Luzon: Zambales prov. LUZON: Cagayan (Abulug River), Isabela (Mt Moises), Nueva Vizcaya (Noso to Imugan; Caraballo Mtn), Aurora (Mt Dingalan), Nueva Ecija (Mt Umingan = Mt Mingan), Zambales (Mt Tapulao), Bataan (Mt Mariveles, Lamao River), Rizal (Mt Lumutan; Paningtingan), Quezon (Mt Binuang), CATANDUANES (Sto Domingo River). Low and medium elevation forests, ascending to 1500m.
  19. Piper curtifolium C.DC., PJS 5 c (1910) Bot. 421; --Merr., EPFP 2 (1923) 7; --Quisumb., PJS 43 (1930) 76; --Type: BS 7107 Ramos (PNH, holo, lost?), Luzon: Abra prov., Mt Paraga. --Piper mindorense C.DC., PJS 5 c (1910) Bot. 423; --Merr., EPFP 2 (1923) 12; --Quisumb., PJS 43 (1930) 79; --Type: FB 4474 Merritt (PNH, holo, lost?), Mindoro: Mindoro Oriental prov., Mt Halcon. LUZON: Ilocos Norte (Mt Palimlim), Abra (Mt Paraga), Mountain Province (Mt Pukis), Benguet (Mt Pulogloco), Rizal (Mt Lumutan), Quezon (Mt Binuang), MINDORO: Mindoro Oriental (Mt Halcon). Montane forests to 2250m.
  20. Piper davaoense C.DC. in Perkins, Fragm. Fl. Philip. (1905) 154; --Merr., EPFP 2 (1923) 7; --Quisumb., PJS 43 (1930) 160; --Type: Warburg 14740 (B, holo), Mindanao: Davao del Sur prov., Mt Dagatpan. --Piper laxirameum C.DC. in Elmer, LPB 3 (1910) 779; --Merr., EPFP 2 (1923) 10; --Type: Elmer 10503 (BISH, MO, NY, US, iso), Mindanao: Davao del Sur prov., Mt Apo, Todaya. ‘In dense humid woods of fertile steep ravines at 4000ft..’, May-1909. MINDANAO: Davao del Sur (Mt Dagatpan; Mt Apo, Todaya; Malita). Forests, 600-1300m.
  21. Piper decumanum L. in Stickman, Herb. Amb. (1754) 19; --Merr., EPFP 2 (1923) 8; --Quisumb., PJS 43 (1930) 42; --Gardner, Blumea 51 (2007) 581; --Type: from Moluccas. Philippines, Sulawesi, Moluccas, New Guinea. LEYTE (Mt Cabalian), MINDANAO: Lanao del Sur (Camp Keithley), Bukidnon (Sumilao), Agusan del Norte (Mt Urdaneta = Mt Hilong-hilong). Medium elevation forests.
  22. Piper delicatum C.DC. in Elmer, LPB 3 (1910) 778; --Merr., EPFP 2 (1922) 8; --Quisumb., PJS 43 (1930) 74; --Type: Elmer 8583 (NY, US, iso), Luzon: Benguet prov., Baguio, ‘in woods at 7000ft of (Mt) Santo Tomas’, Mar-1907. --Piper delicatum var. glabrum C.DC., PJS 5 c (1910) Bot. 444; --Merr., EPFP 2 (1923) 8; --Type: R.S. Williams 1216 (PNH, holo, lost?), Luzon: Benguet prov., Mt Sto Tomas. --Piper polisanum C.DC., PJS 11c (1916) Bot. 209; --Merr., EPFP 2 (1923) 14; --Type: BS 19617 & 19816 R.C. McGregor (syn), Luzon: Ifugao prov., Mt Polis. LUZON: Ilocos Norte (Mt Palimlim), Ifugao (Mt Polis), Mountain Province (Mt Data; Bauco; Malamey; Mt Masapilid; Mt Caua), Benguet (Pauai; Baguio; Mt Sto Tomas). Mossy forests to about 2400m.
  23. Piper densibaccum C.DC., PJS 5c (1910) Bot. 454; --Merr., EPFP 2 (1923) 5; --Quisumb., PJS 43 (1930) 131; --Type: M.S. Clemens s.n. (PNH, holo, lost?), Mindanao: Lanao del Sur prov., Lake Lanao, Camp Keithley, Mar-1906. MINDANAO: Lanao del Sur (Camp Keithley).
  24. Piper dipterocarpinum C.DC., PJS 5 c (1910) Bot. 455; --Merr., EPFP 2 (1923) 8; --Quisumb., PJS 43 (1930) 133; --Type: FB 9146 Whitford & Hutchinson (PNH, holo, lost?; NY, iso), Mindanao: Zamboanga del Sur prov., near Port Banga, in dipterocarp forest, c. 20 m, Jan-1908. --Piper malindangense C.DC., PJS 5 c (1910) Bot. 436; --Merr., EPFP 2 (1923) 11; --Type: FB 4758 Mearns & Hutchinson (PNH, holo, lost?), Mindanao: Misamis Occidental prov., Mt Malindang. BASILAN (Binauangan), MINDANAO: Zamboanga del Sur (Banga), Misamis Occidental (Mt Malindang), Lanao del Sur (Camp Keithley). Lowland forests.
  25. Piper elmeri Merr., PJS 17 (1920) 245; --Quisumb., PJS 43 (1930) 151; --Type: BS 24256 Ramos (PNH, holo, lost?; US, iso), Samar: Catubig River, 10-Feb-1916. LUZON: Laguna (Mt Makiling), Sorsogon (Mt Pocdol), SAMAR (Catubig River). Damp forests, ascending to 450m on Mt Makiling.
  26. Piper ensifolium Quisumb., PJS 43 (1930) 119; --Type: BS 45433 Ramos & Edaño (PNH, holo, lost?; NY, US, iso), Luzon: Aurora prov., Casiguran, in damp forests along stream, low elevation, 29-May-1925. LUZON: Ilocos Norte (Bangui to Claveria), Aurora (Casiguran).
  27. Piper eupodum C.DC., PJS 11 c (1916) Bot. 219; --Merr., EPFP 2 (1923) 8; --Quisumb., PJS 43 (1930) 47; --Type: C.A. Wenzel 1005 (syn), Leyte: Leyte prov., Jaro, Buenavista; BS 15227 Ramos (US, syn), Leyte: Dagami, Aug-1912. LEYTE: Leyte (Dagami; Jaro, Buenavista). Forests, c. 500m.
  28. Piper firmolimbum C.DC., PJS 5 c (1910) Bot. 435; --Merr., EPFP 2 (1923) 9; --Quisumb., PJS 43 (1930) 93; --Type: Clemens s.n. (PNH, holo, lost?), Mindanao: Lanao del Sur prov., Lake Lanao, Camp Keithley. --Piper firmolimbum var. parvilimbum Quisumb., PJS 43 (1930) 95; --Type: Elmer 14517 (PNH, holo, lost?; NY, iso), Luzon: Sorsogon prov., Irosin, Mt Bulusan, Sep-1916. LUZON: Laguna (Mt Makiling), Sorsogon (Mt Bulusan, Irosin), ALABAT, LEYTE: Leyte (Tacloban, Tigbao), MINDANAO: Lanao del Sur (Camp Keithley). Low and medium elevation forests.
  29. Piper fragile Benth. in Hook., London J. Bot. 2 (1843) 234; --Quisumb., PJS 43 (1930) 99; --Gardner, Blumea 51 (2007) 581; --Type: from New Guinea. --Piper fragile var. multinerve C.DC., PJS 5 c (1910) Bot. 421; --Merr., EPFP 2 (1923) 9; --Type: Merrill 3366 (NY, iso), Luzon: Camarines Sur prov., Pasacao, near seashore. Nov-1903. Borneo (Banguey Is off Sabah), Philippines, Moluccas, New Guinea, Solomon Isls, Micronesia. LUZON: Quezon (Mt Pular), Camarines Sur (Pasacao), MINDANAO: Davao Oriental (Mt Mayo; Mati), BUCAS GRANDE. On rocks near the sea, lowland and medium elevation forests and thickets.
  30. Piper fuscinervum Quisumb., PJS 43 (1930) 83; --Type: BS 35380 Martelino & Edaño (PNH, holo, lost?), Panay: Aklan prov., Libacao. PANAY: Capiz (Mt Madia-as; Jamindan), Aklan (Libacao), MINDANAO: Bukidnon (Sumilao; Tangkulan). Low and medium elevation forests.
  31. Piper halconense C.DC., PJS 5 c (1910) Bot. 422; --Merr., EPFP 2 (1923) 9; --Quisumb., PJS 43 (1930) 69; --Type: FB 4393 Merritt (PNH, holo, lost?), Mindoro: Mindoro Oriental prov., Mt Halcon. --Piper cagayanense C.DC., PJS 5 c (1910) Bot. 435; --Merr., EPFP 2 (1923) 5; --Type: BS 7484 Ramos (PNH, holo, lost?), Luzon: Cagayan prov., Pamplona. --Piper atrospicum C.DC., PJS 11 c (1916) Bot. 208; --Merr., EPFP 2 (1923) 3; --Quisumb., PJS 43 (1930) 71; --Type: BS 15359 Ramos (US, iso), Leyte: Leyte prov., Dagami, in mossy forest, Aug-1912. LUZON: Apayao (Tauit; Mt Sulu), Cagayan (Pamplona), Isabela (San Mariano), Zambales (Mt Tapulao; Iba), Quezon (Mt Banahaw; Lucban), Sorsogon (Mt Bulusan, Irosin), MINDORO: Mindoro Oriental (Mt Halcon), LEYTE: Leyte (Dagami), MINDANAO: Agusan del Norte (Mt Urdaneta = Mt Hilong-hilong). Medium elevation forests upwards, ascending to 1500m.
  32. Piper interruptum Opiz in C.Presl, Rel. Haenk. 1 (1828) 157; --Merr., EPFP 2 (1923) 9; --Quisumb., PJS 43 (1930) 154; --Gardner, Blumea 51 (2007) 581; --Type: Haenke s.n. (PR, holo), Luzon. --Piper interruptum var. cumingianum (Miq.) Quisumb., PJS 43 (1930) 158; --Piper cumingianum Miq., Syst. Pip. (1843) 329; --Type: Cuming 1190 (K, holo), Luzon: Albay prov. --Piper interruptum Opiz var. herbaceum C.DC., PJS 5 c (1910) Bot. 448; --Merr., EPFP 2 (1923) 9; --Type: Merrill 3182 (PNH, holo, lost?), Luzon: Bataan prov., Mt Mariveles, Lamao River. --Piper interruptum var. laevirameum (C.DC.) Quisumb., PJS 43 (1930) 158; --Piper laevirameum C.DC., PJS 5 c (1910) Bot. 450; --Merr., EPFP 2 (1923) 10; --Type: Clemens 1255 (PNH, holo, lost?), Mindanao: Lanao del Sur prov., Lake Lanao, Camp Keithley. --Piper interruptum var. loheri (C.DC.) Quisumb., PJS 43 (1930) 157; --Piper loheri C.DC., PJS 5 c (1910) Bot. 450; --Merr., EPFP 2 (1923) 2; --Type: Loher 4552 (US, syn), Luzon: Rizal prov., San Mateo, 12-Jul-90. --Piper interruptum var. multiplinerve C.DC. in Elmer, LPB 3 (1910) 785; --Merr., EPFP 2 (1923) 9; --Quisumb., PJS 43 (1930) 159; --Type: BS 1870 Foxworthy (PNH, holo, lost?), Luzon: Bataan prov., Lamao. --Piper interruptum Opiz var. subarborescens C.DC., PJS 5 c (1910) Bot. 448; --Merr., EPFP 2 (1923) 10; --Type: BS 4585 Ramos (US, iso), Luzon: Rizal prov., Bosoboso, in forest, Aug-1904. --Piper abraense C.DC., PJS 5 c (1910) Bot. 451; --Merr., EPFP 2 (1923) 11; --Type: BS 7195 Ramos (PNH, holo, lost?; NY, US, iso), Luzon: Abra prov., Baret, Jan-1909. --Piper clemensiae C.DC., PJS 5 c (1910) Bot. 449; --Merr., EPFP 2 (1923) 6; --Quisumb., PJS 43 (1930) 166; --Type: Clemens 1200 (PNH, holo, lost?), Mindanao: Lanao del Sur prov., Lake Lanao, Camp Keithley. --Piper elliptibaccum C.DC., PJS 5 c (1910) Bot. 449; --Merr., EPFP 2 (1923) 8; --Quisumb., PJS 43 (1930) 165; --Type: BS 6344 C. Robinson (PNH, holo, lost?), Luzon: Camarines Sur prov., Maagnas. --Piper loheri C.DC. forma multiplinerve C.DC., PJS 5 c (1910) Bot. 450; --Merr., EPFP 2 (1923) 11; --Type: --BS 1870 Foxworthy (syn), Luzon: Bataan prov., Lamao, Dec-1904; --BS 6015 C. Robinson (syn), Luzon: Laguna prov., Liliw, Mar-1904. --Piper pilispicum C.DC., PJS 5 c (1910) Bot. 452; --Merr., EPFP 2 (1923) 13; --Type: BS 5720 Ramos (PNH, holo, lost?; US, iso), Luzon: Benguet prov., Dec-1908. --Piper pulogense C.DC., PJS 5 c (1910) Bot. 453; --Merr., EPFP 2 (1923) 14; --Quisumb., PJS 43 (1930) 162; --Type: FB 16240 Curran, Merritt & Zschokke (PNH, holo, lost?; US, iso), Luzon: Benguet prov., Mt Pulag, 2250m, Jan-1909. --Piper samaranum C.DC., PJS 11 c (1916) Bot. 223; --Merr., EPFP 2 (1923) 15; --Type: BS 17546 Ramos, Samar: Cauayan Valley. ?Taiwan, Philippines and probably elsewhere in W Malesia, New Guinea, Solomon Isls, Vanuatu, Australia. LUZON: Ilocos Norte (Bangui to Claveria; Mt Nagapatan; Mt Piao), La Union (Bauang; San Fernando), Pangasinan (Mt San Isidro Labrador; Villasis, Libtong), Abra (Baret), Mountain Province (Mt Malaya; Bagnen; Gintadan; Bauco; Mt Polis; Mt Caua), Benguet, Cagayan (Abulug to Linao), Isabela (San Mariano), Nueva Vizcaya (Campote), Aurora (Casiguran), Nueva Ecija (Bongabon), Zambales (Candelaria; Botolan, Mt Cabangaan), Bataan (Mt Mariveles, Lamao), Pampanga (Arayat), Bulacan (Angat; Norzagaray; Malolos), Rizal (Mt Lumutan; Mt Susong Dalaga; Mt Angilog; Mt Irid; Mt Tokduan Banoy; Montalban; San Mateo; Antipolo; Pasig; San Isidro; San Andales; Mt Masungit), Manila (Singalong), Laguna (Los Baños; Mt Makiling; Calauan; Liliw; Mt San Cristobal), Quezon, Batangas, Camarines Sur (Maagnas), Sorsogon (Mt Bagacaua), MINDORO: Mindoro Occidental (Paluan), Mindoro Oriental (Mt Halcon), CATANDUANES (Sto Domingo River), PANAY: Antique (Culasi), LEYTE: Leyte (Jaro, Masaganap), SAMAR (Cauayan Valley), MINDANAO: Zamboanga (Sax River mountains), Lanao del Sur (Camp Keithley), Bukidnon (Mt Candoon; Tangkulan), Agusan del Norte (Mt Urdaneta = Mt Hilong-hilong; Liluan Lake), Surigao del Norte (Surigao). Low and medium elevation forests and thickets generally, ascending to 2250m in Benguet. Photos
  33. Piper langlassei C.DC., Ann. Conserv. Jard. Bot. Genève 2 (1898) 273; PJS 5 c (1910) Bot. 433; --Merr., EPFP 2 (1923) 10; --Quisumb., PJS 43 (1930) 91; --Type: Langlassé s.n. (P, iso), Luzon: Laguna prov., base of Mt Banahaw, on calcareous soil, 30-Oct-1894. LUZON: Laguna (Mt Banahaw).
  34. Piper lessertianum (Miq.) C.DC., J. Bot. 4 (1866) 164; --Merr., EPFP 2 (1923) 10; --Quisumb., PJS 43 (1930) 36; --Gardner, Blumea 51 (2007) 581; --Chavica lessertiana Miq., Syst. Pip. (1843) 270; --Type: Cuming 1343 (‘1342’) (BM, holo), Luzon: Cagayan prov. --Piper lessertianum var. oblongibaccum (C.DC.) Quisumb., PJS 43 (1930) 39; --Piper oblongibaccum C.DC. in Elmer, LPB 3 (1910) 777; --Merr., EPFP 2 (1923) 12; --Type: Elmer 9456 (BISH, MO, NY, US, iso), Negros: Negros Oriental prov., Cuernos Mtns, in woods of densely forested trail, 4000ft, Mar-1908. Philippines, Sulawesi, New Guinea. LUZON: Apayao (Guiniri), Kalinga (Mt Masingit), Mountain Province (Pininggat), Ifugao (Mt Polis), Cagayan, Aurora (Casiguran), Rizal (Montalban), Quezon (Mt Binuang; Mt Banahaw), Sorsogon (Mt Bulusan), ALABAT, MINDORO: Mindoro Occidental (Mt Calavite), Mindoro Oriental (Mt Halcon), PANAY: Aklan (Mt Magosolon; Libacao), Capiz (Jamindan; Mt Salibongbong; Mt Kinablangan), NEGROS: Negros Occidental (Mt Kanlaon), Negros Oriental (Cuernos Mtns), SAMAR: Northern Samar (Las Navas, Catubig River), MINDANAO: Zamboanga del Sur (Malangas), Lanao del Sur (Camp Keithley), Bukidnon (Mahilucot River), Davao del Sur (Mt Apo), BUCAS GRANDE. Damp forests, mostly from 400-2200m.
  35. Piper longipedicellatum Quisumb., PJS 43 (1930) 144; --Type: BS 38066 Ramos & Edaño (PNH, holo, lost?; US, iso), Luzon: Mountain Province, Mt Caua, on slopes in the mossy forest, c. 1800m, 3-Mar-1920. LUZON: Mountain Province (Mt Caua), Ifugao (Mt Polis). Mossy forests, c. 1800m.
  36. Piper longistigmum C.DC. in Elmer, LPB 3 (1910) 770; --Merr., EPFP 2 (1923) 11; --Quisumb., PJS 43 (1930) 95; --Type: Elmer 7578 (MO, NY, US, iso), Luzon: Quezon prov., Lucban, May-1907. LUZON: Quezon (Lucban), CATANDUANES (Sto Domingo River). Lowland forests.
  37. Piper longivaginans C.DC., PJS 5c (1910) Bot. 444; --Merr., EPFP 2 (1923) 3; --Quisumb., PJS (1930) 72; --Type: BS 6069 C.B. Robinson (US, iso), Luzon: Laguna prov., Mt Banahaw, Mar-1909. --Piper crassinodum C.DC. in Elmer, LPB 3 (1910) 780; --Merr., EPFP 2 (1922) 7; --Type: Elmer 8042 (BISH, MO, NY, US, syn); --Elmer 7626a (MO, US, syn), Luzon: Quezon prov., Lucban, May-1907. --Piper magallanesanum C.DC., PJS 11 c (1916) Bot. 212; --Merr., EPFP 2 (1923) 11; --Type: Elmer 12313, Sibuyan: Mt Giting-giting. LUZON: Nueva Ecija (Mt Umingan = Mt Mingan), Rizal (Montalban; Balacbac; Paningtingan; Mt Lumutan; Mt Irid; Mt Tokduangbanog), Laguna (Mt San Cristobal; Mt Banahaw), Quezon (Mt Tulaog; Mt Binuang; Lucban), Camarines Sur (Mt Isarog), SIBUYAN (Mt Giting-giting). Medium to high montane forests, ascending to 2500m.
  38. Piper macropiper Pennant, Outlines of the Globe 4 (1800) 242; --Merr., J. Arnold Arb. 29 (1948) 191; --Chew, J. Arnold Arb. 53 (1972) 10; --Gardner, Blumea 51 (2007) 582; --Type: Rumphius, Herb. Amboin. 5 (1747) 46. --Piper arborescens Roxb., Hort. Beng. (1814) 80; --Merr., EPFP 1 (1922) 3; --Quisumb., PJS 43 (1930) 20. --Piper arborescens var. angustilimbum Quisumb., PJS 43 (1930) 22; --Type: Elmer 7384 (PNH, holo, lost?; NY, iso), Luzon: Quezon prov., Lucban, May-1907. --Piper arborescens var. hirtellum (Miq.) Merr., EPFP 1 (1922) 3; --Quisumb., PJS 43 (1930) 23; --Piper miniatum Blume var. hirtellum (Miq.) C.DC. in Elmer, LPB 3 (1910) 776; PJS 5 c (1910) Bot. 422; Candollea 1 (1923) 261; --Chavica miniata Miq. var. hirtella Miq., Fl. Ind. Bat. Suppl. (1860-61) 473. --Piper glandulosum Opiz in C.Presl, Rel. Haenk. 1 (1828) 158; --Type: Haenke s.n. (MO, ?iso), Luzon. --Piper pilipes C.DC., PJS 5 c (1910) Bot. 423; --Merr., EPFP 2 (1923) 16; --Quisumb., PJS 43 (1930) 25; --Type: Clemens s.n., Mindanao: Lanao del Sur prov., Lake Lanao, Camp Keithley. --Piper trichophlebium Quisumb., PJS 43 (1930) 24; --Type: Merrill 9404 (PNH, holo, lost?; NY, US, iso), Palawan: Palawan prov., Binaloan, in forests at low elevation, 19-May-1913. LUZON: Ilocos Norte (Bangui to Claveria, Mt Calvario), Abra (Mt Paraga), Apayao (Ngagan), Aurora (Casiguran, Mamatoc River), Bataan (Mt Mariveles), Rizal (Montalban), Laguna (Mt Makiling; San Antonio), Quezon (hills near Malinao; Mt Binuang), Albay, Sorsogon (Mt Bulusan), POLILLO, MINDORO: Mindoro Oriental (Pinamalayan), PALAWAN (Binaloan), PANAY: Mt Bulilao; Capiz (Mt Madia-as; Jamindan; Agsaman; Mt Salibongbong), LEYTE: Leyte (Jaro, Buenavista; Dagami; Tacloban, Tigbao), SAMAR (Ambalate; Catubig River; Catbalogan; Mt Capotoan; Las Navas), BASILAN, MINDANAO: Zamboanga (Sax River), Zamboanga del Sur (Malangas), Lanao del Sur (Lake Lanao, Camp Keithley), Bukidnon (Sumilao; Mt Candoon; Tangkulan), Davao del Sur (Mt Apo, Todaya), Davao Oriental (Mt Mayo), Agusan del Norte (Mt Urdaneta = Mt Hilong-hilong), Surigao del Norte (Placer). Common in lowland and medium elevation forests, ascending to 1000m. Photos
  39. Piper majusculum Blume, Verh. Bat. Genoots. 11 (1826) 210; --Quisumb., PJS 43 (1930) 45; --Gardner, Blumea 51 (2007) 582; --Type: from Java. --Piper rotundistigmum C.DC., PJS 5c (1910) Bot. 425; --Merr., EPFP 2 (1923) 15; --Type: Clemens s.n. (G-DC, holo), Mindanao: Lanao del Sur prov., Lake Lanao, Camp Keithley. --Piper rotundistigmum var. pilosius C.DC., Candollea 1 (1923) 201, 206, --Type: BS 17434 Ramos, Samar. --Piper leyteanum C.DC., PJS 11 c (1916) Bot. 220; Candollea 1 (1923) 206; --Type: C.A. Wenzel 214, Leyte: Leyte prov., Dagami. India, Malaysia, Myanmar, Sumatra, Java, Borneo, Thailand, Philippines, Ambon, New Guinea, Solomon Isls. LUZON: Albay, Sorsogon (Mt Lalao; Mt Bulusan), LEYTE: Leyte (Dagami; Jaro), SAMAR (Lauaan), MINDANAO: Lanao del Sur (Lake Lanao, Camp Keithley), Davao, Agusan del Norte (Mt Urdaneta = Mt Hilong-hilong), Surigao del Norte (Surigao; Placer). Lowland and medium elevation forests. 
  40. Piper medinillifolium Quisumb., PJS 43 (1930) 29; --Type: BS 15364 Ramos (PNH, holo, lost?; US, iso), Leyte: Leyte prov., Burauen, 12-Aug-1912. LEYTE: Leyte (Burauen; Jaro, Buenavista). In forests, c. 500m.
  41. Piper melanocaulon Quisumb., PJS 43 (1930) 49; --Type: BS 35725 Martelino & Edaño (PNH, holo, lost?; US, iso), Panay: Iloilo prov., Mt Bolilao, on the summit, c. 1000m, 29-Jun-1919. PANAY: Iloilo (Mt Bolilao). Note by L.L.Co: Resembles Piper eupodum C.DC., but may be readily distinguished from it by its black branches and leaves, shorter peduncles and spikes, laxly arranged fruits, its glabrous rachis, and its bracts.
  42. Piper merrillii C.DC., PJS 5 c (1910) Bot. 426; --Merr., EPFP 2 (1923) 12; --Quisumb., PJS 43 (1930) 52; --Type: Merrill 1809 (PNH, holo, lost?; US, iso), Mindoro: Mindoro Oriental prov., Baco River, Apr-1903. --Piper dagamiense C.DC., PJS 11 c (1916) Bot. 211; --Merr., EPFP 2 (1923) 7; --Type: BS 15181 Ramos (US, iso), Leyte: Leyte prov., Dagami. LUZON: Isabela (San Mariano), Quezon (Mt Binuang), Laguna (San Antonio), ALABAT, MINDORO: Mindoro Occidental (Mt Calavite), Mindoro Oriental (Baco River; Mt Halcon), PALAWAN (Mt Capoas), PANAY: Antique (Culasi), Aklan (Mt Magosolon; Libacao), Capiz (Jamindan), NEGROS: Negros Occidental (Mt Kanlaon; Mt Silay), BOHOL: Bohol (Valencia), LEYTE: Leyte (Dagami), SAMAR (Cauayan; Catubig River), MINDANAO: Davao Oriental (Mt Galintan). Low and medium elevation forests, ascending to 1250m. Photos
  43. Piper multistigmum C.DC., PJS 11 c (1916) Bot. 222; --Merr., EPFP 2 (1923) 12; --Quisumb., PJS 43 (1930) 163; --Type: BS 19819 R.C. McGregor (NY, US, iso), Luzon: Ifugao prov., Mt Polis, Feb-1913. LUZON: Ifugao-Mountain Province (Mt Polis). Forest, elevation cited as 1000m (possibly nearer 2000m). Note by L.L.Co: Resembling Piper interruptum Opiz, but stigma 5 or 6.
  44. Piper mymecophilum C.DC., PJS 11 c (1916) Bot. 211; --Merr., EPFP 2 (1923) 12; --Quisumb., PJS 43 (1930) 56; --Gardner, Blumea 51 (2007) 583; --Type: BS 17599 Ramos (L, US, iso), Samar: Mt Cauayan, Mar/Apr-1914. LEYTE: Leyte (Tigbao; Tacloban), SAMAR (Mt Cauayan; Camanabaan; Catubig River, Pinipisakan). Damp forests, 50-400m.
  45. Piper nigrum L., Sp. Pl. (1753) 28. Photos
  46. Piper ovatibaccum C.DC. in Elmer, LPB 3 (1910) 782; --Merr., EPFP 2 (1923) 13; --Quisumb., PJS 43 (1930) 80; --Type: Elmer 7888 (BISH, MO, NY, US, syn), Luzon: Quezon prov., Lucban, 700m, May-1907; --Elmer 8106 (syn), Luzon: Quezon prov., Lucban. --Piper robinsonii C.DC., PJS 5c (1910) Bot. 445; --Merr., EPFP 2 (1923) 13; --Type: BS 6065 C.B. Robinson (PNH, holo, lost?), Luzon: Laguna prov., Mt Banahaw. LUZON: Rizal (Mt Irid; Balacbac), Laguna (Sta Maria; Paete; San Antonio; Mt Banahaw), Quezon (Infanta; Mt Binuang; Lucban), Camarines Norte (Mt Bagacay), Sorsogon (Mt Kililibong), MINDORO: Mindoro Oriental (Mt Halcon), CATANDUANES (Mt Mariguidon), PANAY: Capiz (Jamindan), LEYTE: Leyte (Dagami; Jaro, Buenavista), MINDANAO: Lanao del Sur (Lake Lanao, Camp Keithley), Bukidnon (Kabaritan; Mt Dalirig; Mt Candoon; Agusan River near Tangkulan), Agusan del Norte (Mt Urdaneta = Mt Hilong-hilong). Medium elevation forests, ascending to 1075m.
  47. Piper parcirameum C.DC. in Elmer, LPB 3 (1910) 781; --Merr., EPFP 2 (1923) 13; --Quisumb., PJS 43 (1930) 65; --Type: Elmer 10899 (BISH, MO, NY, syn), Mindanao: Davao del Sur prov., Mt Apo, Todaya, dense forest at 4000ft of Mt Calelan, Jun-1909; --Elmer 11274 (BISH, MO, NY, syn), Mindanao: Davao del Sur prov., Mt Apo, Todaya, dense forest, 3000ft, Aug-1909. MINDANAO: Davao del Sur (Mt Apo). In forests, 900-1200m.
  48. Piper parong Quisumb., PJS 43 (1930) 105; --Type: BS 24465 Ramos (PNH, holo, lost?; NY, US, iso), Samar: Northern Samar prov., Palapag, Catubig, 11-Mar-1916. SAMAR: Northern Samar. Lowland thickets and streamside forests.
  49. Piper paucinerve C.DC. in Perkins, Fragm. Fl. Philip. (1905) 156; --Merr., EPFP 2 (1923) 13; --Quisumb., PJS 43 (1930) 147; --Type: Warburg 11929 (B, holo), Luzon: Isabela prov., Malunu. LUZON: Isabela (Malunu).
  50. Piper philippinum Miq., Syst. Pip. (1843) 322; --Merr., EPFP 2 (1923) 13; --Quisumb., PJS 43 (1930) 110; --Gardner, Blumea 51 (2007) 583; --Type: Cuming 912 (de Lessert herbarium, holo), Luzon: Albay prov. (Gardner otherwise cites Cuming 1642 (G or P?)). --Piper albidirameum C.DC. in Perkins, Fragm. Fl. Philip. (1905) 153; --Merr., EPFP 2 (1923) 8; --Quisumb., PJS 43 (1930) 115; --Type: Warburg 14751 (B, holo), Mindanao: Davao del Sur prov., Taumo. --Piper crassilimbum C.DC., PJS 11 c (1916) Bot. 210; --Merr., EPFP 2 (1923) 7; --Type: Merrill 7649 (US, syn), Luzon: Benguet prov., Baguio, thickets in limestone region, 1550m, May-1911. --Piper fuscescentirameum C.DC., PJS 11 c (1916) Bot. 217; --Merr., EPFP 2 (1923) 8; --Type: C.A. Wenzel 1184, Leyte: in forest, c. 500m. --Piper lividum C.DC. in Perkins, Fragm. Fl. Philip. (1905) 155; --Merr., EPFP 2 (1923) 8; --Type: Warburg 11930 (B, holo), Luzon: Isabela prov., Malunu. --Piper lucbanense C.DC. in Elmer, LPB 6 (1914) 2293; --Merr., EPFP 2 (1923) 11; --Type: Elmer 7383, Luzon: Quezon prov., Lucban. --Piper maagnasanum C.DC., PJS 5c (1910) Bot. 429; --Merr., EPFP 2 (1923) 8; --Quisumb., PJS (1930) 117; --Type: BS 6355 C.B. Robinson, Luzon: Camarines Sur prov., Maagnas. --Piper pendulifolium C.DC. in Elmer, LPB 3 (1910) 772; --Merr., EPFP 2 (1923) 8; --Type: Elmer 10942 (BISH, iso), Mindanao: Davao del Sur prov., Mt Apo, Todaya, Jul-1909. --Piper petraeum C.DC., PJS 5 c (1910) Bot. 430; --Merr., EPFP 2 (1923) 13; --Type: Elmer 6430 (NY, US, iso), Luzon: Benguet prov., Twin Peaks, on rocks in shady places, May-1904. --Piper psilocarpum C.DC., PJS 11 c (1916) 215; --Merr., EPFP 2 (1923) 8; --Type: C.A. Wenzel 896, Leyte: Leyte prov., Jaro, Buenavista. Forests, c. 500m. S China, Taiwan, Philippines. LUZON: Ilocos Norte (Burgos), Apayao (Ngagan), Benguet (Baguio; Twin Peaks), Isabela (Malunu), Nueva Ecija (Mt Umingan), Aurora (Casiguran; Mt Dingalan), Rizal (Mt Lumutan; Bosoboso; Sta Inez; Mt Irid), Cavite (Mendez), Laguna (Mt Makiling; Mt San Cristobal; Mt Banahaw; Siniloan; San Antonio), Quezon (Mt Tulaog; Mt Binuang; Infanta-Siniloan Trail; Lucban; hills near Malinao; Malbog; Atimonan), Camarines Norte (Paracale), Camarines Sur (Adiagnao), Albay, Sorsogon (Mt Bulusan; Mt Bagacana), ALABAT (Sangerin), MINDORO: Mindoro Occidental (Paluan), Mindoro Oriental (Naujan; Pinamalayan), MASBATE, BATBATAN (W of Panay: Antique prov.), PANAY: Antique (Culasi), Aklan (Libacao; Mt Timbaban), Capiz (Jamindan), BOHOL: Bohol (Valencia), BILIRAN, LEYTE: Leyte (Tacloban, Tigbao; Cabalian; Jaro, Buenavista), SAMAR (Catubig River), MINDANAO: Zamboanga del Sur (Mt Tubuan), Davao del Sur (Taumo; Mt Apo, Todaya), Bukidnon (Botoan; Impasugong), Agusan del Norte (Mt Urdaneta), Surigao del Norte (Surigao; Placer), Davao Oriental (Mati), BUCAS GRANDE. Low and medium elevation forests and thickets, ascending to 1550m.
  51. Piper quinqueangulatum Miq., Syst. Pip. (1854) 85; --Gardner, Blumea 51 (2007) 583; --Type: from Java. --Piper cristatum C.DC. in Elmer, LPB 3 (1910) 770; --Merr., EPFP 2 (1923) 7; --Type: Elmer 10703 (G-DC, lecto; BISH, BM, E, GH, K, MO, NY, US, isolecto), Mindanao: Davao del Sur prov. (Mt Apo, Todaya). Epiphytic in very humid forest along Baruring River, 3500ft, May-1909. --Piper korthalsii Miq., Ann. Mus. Bot. Lugd.-Bat. 1 (1863) 139; --C.DC., Candollea 1 (1923) 71.--Merr., EPFP 2 (1923) 10; --Quisumb., PJS 43 (1930) 169. --Piper korthalsii Miq. var. longibracteum C.DC. in Elmer, LPB 3 (1910) 763; --Type: Clemens 463, Mindanao: Lanao del Sur prov., Lake Lanao, Camp Keithley. Philippines, Java, Malaysia, Sumatra, Thailand. LUZON: Apayao (Abulug River; Tamoe; Guiniri), Mountain Province (Bauco), Ifugao (Mt Polis), Benguet (Baguio; Sablan; San Fernando Trail), Cagayan (trail from Sitio Viola to Mt Nanig), Isabela (San Mariano), Nueva Vizcaya (Campote), Aurora (Casiguran), Rizal (Montalban; Mt Irig; Mt Irid), Laguna (San Antonio), MINDORO: Mindoro Oriental (Mt Halcon; Pinamalayan; Bongabong River), PANAY: Antique (Culasi), Capiz (Jamindan; Mt Agramilig), LEYTE: Leyte (Dagami; Jaro, Buenavista), MINDANAO: Lanao del Sur (Lake Lanao, Camp Keithley), Davao del Sur (Mt Apo, Todaya), Agusan del Norte (Mt Urdaneta = Mt Hilong-hilong), Surigao del Norte (Surigao; Placer). Damp lowland to montane forests, 400-2200m, one record from 2800m. Photos
  52. Piper ramosii C.DC., PJS 5 c (1910) Bot. 426; --Merr., EPFP 2 (1923) 14; --Quisumb., PJS 43 (1930) 82; --Type: BS 1755 Ramos, Luzon: Rizal prov., Bosoboso. --Piper calvifolium C.DC., PJS 11 c (1916) Bot. 217; --Merr., EPFP 2 (1923) 5; --Type: Loher 6794, pro parte (K), Luzon. LUZON: Rizal (Mt Lumutan; Montalban; Antipolo; Bosoboso; San Isidro; San Andales). Low and medium elevation forests. Note by L.L.Co: Nearest to P. ovatibaccum C.DC., differing in its auriculate leaves and reniform anthers.
  53. Piper retrofractum Vahl, Enum. 1 (1804) 314; --Merr., EPFP 2 (1923) 14; --Quisumb., PJS 43 (1930) 106; --Gardner, Blumea 51 (2007) 583; --Type: from East Indies. --Piper palawanum C.DC., PJS 11 c (1916) Bot. 210; --Merr., EPFP 2 (1923) 13; --Type: BS 1553 E. Fenix (US, iso), Palawan: Palawan prov., Babuyan. --Piper parvifolium Blanco, Fl. Filip. (1837) 23; Fl. Filip. ed. 2 (1845) 17; Fl. Filip. ed. 3, 1 (1877) 32; --Type: not extant; 'illustrative specimen': Merrill Sp. Blancoanae 523 (US), Luzon: Bulacan prov., Angat, Sep-1913, leg. M Ramos. --Piper retrofractum Vahl var. latifolium C.DC. in Perkins, Fragm. Fl. Philip. (1905) 157; --Type: Warburg 12125 (B, holo), Luzon: Cagayan prov., Buguey. Indochina, Peninsular Malaysia to the Moluccas. BABUYAN ISLS GROUP (DALUPIRI; CAMIGUIN), LUZON: Ilocos Norte (Burgos), La Union (San Fernando), Pangasinan (Rosales; Umingan), Kalinga (Tabuk), Abra (Caburao), Cagayan (Buguey), Nueva Vizcaya (Dupax), Nueva Ecija (Cabanatuan), Bataan (Mt Mariveles; Duale), Bulacan (Angat), Rizal (Mt Lumutan; Montalban; Masambong; Parañaque; Baras; Pililla; Jalajala; Malapad na Bato), Cavite, Laguna (Los Baños; Galas; Pagsanjan), MINDORO: Mindoro Oriental (Puerto Galera; Baco; Pinamalayan), PALAWAN (Babuyan; Ulugan Bay; Taytay; Puerto Princesa; Iwahig; Alfonso XIII = Quezon); PANAY: Antique (Lipata), Capiz (Mt Timbaban), Iloilo, GUIMARAS. Lowland thickets, 0-850m. Conservation status: Other Threatened Species (DAO 2017-11). Photos
  54. Piper sarmentosum Roxb., Fl. Ind. 1 (1820) 162; --Merr., EPFP 1 (1922) 15; --Quisumb., PJS 43 (1930) 101; --Gardner, Blumea 51 (2007) 584. --Piper allenii C.DC., PJS 5 c (1910) Bot. 441; --Merr., EPFP 2 (1923) 3; --Type: Allen 152 (PNH, holo, lost?), Mindanao: Surigao del Norte prov., Surigao. --Piper siassiense C.DC., PJS 5 c (1910) Bot. 443; --Merr., EPFP 2 (1923) 15; --Type: Merrill 5311 (NY, US, iso), Sulu Archipelago: Siasi Island, in coconut groves, 12-Oct-1906. --Piper zamboangae C.DC., PJS 5 c (1910) Bot. 424; --Merr., EPFP 2 (1923) 17; --Type: Hallier s.n. (PNH, holo, lost?), Mindanao: Zamboanga. India, S China, southward to the Moluccas. PALAWAN (Lake Manguao; Brookes Point), NEGROS: Negros Occidental (Kabankalan), SAMAR: Eastern Samar (Borongan), CAMIGUIN (Mambajao), SIASI MINDANAO: Zamboanga, Surigao del Norte (Surigao). Lowland thickets, coconut groves, etc. Photos
  55. Piper sibulanum C.DC. in Perkins, Fragm. Fl. Philip. (1905) 158; --Quisumb., PJS 43 (1930) 103; --Type: Warburg 14742 (B, holo), Mindanao: Davao del Sur prov. --Piper oophyllum C.DC., PJS 5 c (1910) Bot. 430; --Merr., EPFP 2 (1923) 17; --Type: Copeland 333 (US, iso), Mindanao: Davao, sea-level in coconut groves, Mar-1904. --Piper williamsii C.DC., PJS 5 c (1910) Bot. 441; --Type: R.S. Williams 2750 (PNH, holo, lost?; NY, iso), Mindanao: Davao del Sur prov., Sta Cruz, 13-Apr-1905. MINDANAO: Zamboanga del Sur (Malangas), Bukidnon (Tangkulan), Davao, Davao del Sur (Sta Cruz), Davao Oriental (Mati). Lowland forests and coconut groves.
  56. Piper simile Quisumb., PJS 43 (1930) 34; --Type: BS 33105 (PNH, holo, lost?), Luzon: Ilocos Norte prov., Bangui to Claveria, c. 500m, 11-Aug-1918. LUZON: Ilocos Norte (Bangui to Claveria), Aurora (Mt Alzapan). Medium elevation to high montane forests, 500-1900m.
  57. Piper sorsogonum C.DC., Candollea 1 (1923) 223, nom. nud., 2 (1923) 188; --Quisumb., PJS 43 (1930) 150; --Type: Elmer 14921 (NY, US, iso), Luzon: Sorsogon prov., Mt Bulusan, Irosin, Sep-1916. LUZON: Camarines Norte (Mt Bagacay), Sorsogon (Mt Bulusan), LEYTE: Leyte (Mt Abucayan). Lowland and medium elevation forests.
  58. Piper spathelliferum Quisumb., PJS 43 (1930) 164; --Type: BS 26383 Ramos & Edaño (PNH, holo, lost?), Luzon: Nueva Ecija prov., Mt Umingan, on slope near summit of Mt Umingan, c. 400m, 20-Aug-1916. LUZON: Nueva Ecija (Mt Umingan). Note by L.L.Co: Similar to Piper elliptibaccum C.DC., differing in the presence of glandular brown dots on the lower surface of lamina.
  59. Piper subbulatum K.Schum. & Lauterb., Fl. Schutzgeb. Südsee (1900) 262; --Gardner, Blumea 48 (2003) 64; Blumea 51 (2007) 584; --Type: from New Guinea. --Piper lageniovarium C.DC. in Elmer, LPB 3 (1910) 797; --Merr., EPFP 2 (1923) 10; --Quisumb., PJS 43 (1930) 44; --Type: Elmer 10589 (BISH, MO, NY, US, iso), Mindanao: Davao del Sur prov., Mt Apo, Todaya, May-1909. New Guinea, New Ireland and New Britain, Bougainville, Solomon Isls, Vanuatu, Philippines. MINDANAO: Bukidnon (Kalasungay), Davao del Sur (Mt Apo, Todaya). Medium elevation thickets and forests, streamsides, etc. (0-)1300-1700m.
  60. Piper subprostratum C.DC., PJS 5 c (1910) Bot. 425; --Merr., EPFP 2 (1923) 15; --Quisumb., PJS 43 (1930) 40; --Type: FB 6751 ML Merritt (US, iso), Mindoro: Mindoro Oriental prov., S of Lake Naujan, Mar/Apr-1907. LUZON: Rizal (Mt Lumutan), MINDORO: Mindoro Oriental (S of Lake Nauhan; Mt Halcon). Low and medium elevation forests.
  61. Piper toppingii C.DC. in Elmer, LPB 3 (1910) 783; --Merr., EPFP 2 (1923) 16; --Quisumb., PJS 43 (1930) 30; --Type: Elmer 8375 (MO, NY, US, syn), Luzon: Benguet prov., Baguio, Mar-1907; --Elmer 5850 (NY, US, syn), Luzon: Benguet prov., Baguio, Mar-1904. --Piper chlorocarpum C.DC., PJS 11 c (1916) Bot. 221; --Merr., EPFP 2 (1923) 16; --Type: BS 16638 Ramos (US, iso), Luzon: Laguna prov., San Antonio, Sep/Oct-1912. LUZON: Ilocos Norte (Bangui to Claveria; Mt Palimlim), Mountain Province (Mt Pukis; Mt Masapilid), Ifugao (Mt Polis), Benguet (Mt Pulag; Mt Tonglon = Mt Sto Tomas; Baguio), Nueva Vizcaya (Compote), Rizal (Mt Susong Dalaga; Mt Lumutan; San Isidro), Laguna (Mt Makiling; Mt San Cristobal; San Antonio), MINDORO: Mindoro Occidental (Mt Calavite; Paluan). Medium elevation to high-montane forests, ascending to 2100m.
  62. Piper urdanetanum C.DC. in Elmer, LPB 6 (1914) 2293; --Merr., EPFP 2 (1923) 16; --Quisumb., PJS 43 (1930) 32; --Type: Elmer 13713 (BISH, NA, NY, US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta = Mt Hilonghilong, Sep-1912. --Piper wenzelii C.DC., PJS 11 c (1916) Bot. 213; --Merr., EPFP 2 (1923) 17; --Type: C.A. Wenzel 628 (US, iso), Leyte: Leyte prov., 30-Mar-1914. --Piper hirtirache C.DC., PJS 11 c (1916) Bot. 213; --Merr., EPFP 2 (1923) 17; --Type: C.A. Wenzel 1168, Leyte: Leyte prov., Jaro, Buenavista. LEYTE: Leyte (Jaro, Buenavista), SAMAR (Ambalate; Catubig River), MINDANAO: Agusan del Norte (Mt Urdaneta). Low and medium elevation forests.
  63. Piper varibracteum C.DC. in Elmer, LPB 3 (1910) 764; --Merr., EPFP 2 (1923) 16; --Quisumb., PJS 43 (1930) 78; --Type: Elmer 11998 (BISH, MO, NY, US, iso), Mindanao: Davao del Sur prov., Mt Apo, ‘scandent upon trees and overtopping dense thickets along the sea coast on moist sandy soil’, Oct-1909. --Piper striatum C.DC. in Elmer, LPB 3 (1910) 776; --Merr., EPFP 2 (1923) 15, nom. illeg., auct. non Vahl; --Type: Elmer 11764 (BISH, MO, NY, US, iso), Mindanao: Davao del Sur prov., Mt Apo, epiphytic in light woods, c. 3000ft, Sep-1909. MINDANAO: Davao del Sur (Mt Apo). Sea-level to medium elevation forests. Near Piper curtifolium.

Non-native species:

  1. Piper aduncum L., Sp. Pl. (1753) 29. Widely naturalized, invasive (Pelser & Barcelona, pers. obs. 2017). Photos
  2. Piper umbellatum L., Sp. Pl. 1 (1753) 30. --Piper subpeltatum Willd., Sp. Pl. 1(1) (1797) 166–167 1797; --Piper subumbellatum L. var. subpeltatum (Willd.) C.DC. in Donn.-Sm. Enum 6: 39; --Merr., EPFP 2 (1923) 16; --Quisumb., PJS 43 (1930) 17. --Piper umbellatum L. var. glabrum C.DC., Bull. Herb. Boiss. 16 (1898) 494; --Merr., EPFP 2 (1923) 16. Photos

Doubtfully occurring in the Philippines:

  1. Piper longum L., Sp. Pl. (1753) 29. See Quisumbing (1930).

Doubtful species:

  1. Piper jagori C.DC., Prodr. 161 (1859) 358, as ‘jayeri’; --Merr., EPFP 2 (1923) 10. --Piper rufinerve Opiz in C.Presl, Rel. Haenk. 1 (1828) 159; --Merr., EPFP 2 (1923) 15. --Piper taumanum C.DC. in Perkins Frag. Fl. Philip. (1905) 159; --Merr., EPFP 2 (1923) 15.

 Zippelia Blume = Piper


Literature

DENR Administrative Order. 2017. Updated national list of threatened Philippine plants and their categories. DAO 2017-11

Gardner, RO. 2007. Piper (Piperaceae) in the Philippine Islands: the climbing species. Blumea 51: 569-586

Mathieu, G. Peperomia.net

Quisumbing, E. 1930. Philippine Piperaceae. Philippine Journal of Science 43: 1-246 + 24 pl.

Suwanphakdee, C, DA Simpson, TR Hodkinson & P Chantaranothai. 2016. Taxonomic notes on the genus Piper (Piperaceae). Nordic Journal of Botany.

Acknowledgements

Ravan Schneider kindly provided some photographs for this webpage.


Copyright © 2011, Co's Digital Flora of the Philippines

Last updated 7 September 2017