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Ericaceae

including Epacridaceae

Costera J.J.Sm.

No Photos
  1. Costera lanaensis (Merr.) Airy Shaw & J.J.Sm., Bull. Jard. Bot. Buitenz. 3, 13 (1935) 421; --Sleumer, Blumea 11 (1961) 3; --Iaera lanaensis (Merr.) H.F.Copel., PJS 47 (1932) 83; --Vaccinium lanaense Merr., PJS 3 c (1908) Bot. 161; EPFP 3 (1923) 250; --Type: Clemens 431(PNH, holo, lost). --Vaccinium medinilloides Elmer, LPB 3 (1911) 1097. Moluccas (Morotai; Talaud Is.), Philippines. NEGROS: Negros Oriental prov. (Cuernos Mtns), PANAY: Capiz prov. (Mt Bulilao), MINDANAO: Zamboanga (Sax River, San Ramon), Lanao del Norte prov. (Camp Keithley, type locality), Davao: (Mt Apo, type locality of Vaccinium medinilloides Elmer). Midmontane forest, humid river gullies, etc., epiphytic on branches, 700-1300m.
  2. Costera loheri (Merr.) Airy Shaw & JJ.Sm., Bull. Jard. Bot. Buitenz. 3, 13 (1935) 421; --Iaera loheri (Merr.) H.F.Copel., PJS 47 (1932) 85. --Sleumer, Blumea 11 (1961) 3; --Vaccinium loheri Merr., PJS 7 c (1912) Bot. 323; EPFP 3 (1923) 250; --Type: Loher 6187 (PNH, holo, lost; US). LUZON: Rizal (Sierra Madre Range E of Montalban), PANAY: Iloilo (Miag-ao). In mossy forest, probably above 1000m.
  3. Costera lucida (Merr.) Airy Shaw & J.J.Sm., Bull. Jard. Bot. Buitenz. 3, 13 (1935) 421; --Iaera lucida (Merr.) H.F.Copel., PJS 47 (1932) 85; --Diplycosia lucida Merr., PJS 11 c (1916) Bot. 28; --Type: BS 23504 (PNH, holo, lost; US), Luzon: Sorsogon prov., Mt Kilibong. --Vaccinium costeroides Merr., EPFP 3 (1923) 248. LUZON: Sorsogon. Epiphytic in the mossy forest at c. 1000m. Fl. & fr. Aug. Known only from the type collection.

Dimorphanthera (Drude) F.Muell. ex J.J.Sm.

Photos
  1. Dimorphanthera apoana (Merr.) Schltr., Bot. Jahrb. 55 (1918) 185; --Merr., EPFP 3 (1923) 252; --Copel., PJS 47 (1932) 89; --Sleumer, Nova Guinea, Bot. no. 7 (1961) 83; --Vaccinium apoanum Merr., Publ. Govt. Lab. Philipp. 29 (1905) 39; --Type: Copeland 1105 (PNH, holo, lost; K, NY, US, iso). Photos
    • var. apoana. --Vaccinium calelanum Elmer, LPB 3 (1911) 1096. PANAY: Antique (Mt Madiaas), NEGROS: Negros Occidental (Mt Silay; Mt Kanlaon), MINDANAO: Davao (Mt Apo; Mt Calelan; Mt McKinley), Bukidnon (Mt Candoon; Mt Kitanglad; Mt Kinasalapi). In mossy forest, often on exposed ridges or in summit vegetation, 1200-2450m. Fl. Jan-Dec. 
    • var. mindanaensis (Merr.) P.F.Stevens, J. Arnold Arb. 58 (1977) 439; --Dimorphanthera mindanaensis Merr., PJS 20 (1922) 418; EPFP 3 (1923) 252; --Copel., PJS 47 (1932) 92; --Sleumer, Nova Guinea, Bot. no. 7 (1961) 82; --Type: FB 4708 Mearns & Hutchinson (PNH, lecto, lost), Mindanao: Misamis Occidental prov., Mt Malindang, MINDANAO: Misamis Occidental (Mt Malindang), Bukidnon (Mt Lipa). Mossy forest, c. 1700m. Twice found.

Diplycosia Blume

Last edited by Pieter B. Pelser, 18 September 2017

Photos
  1. Diplycosia apoensis Elmer, LPB 3 (1911) 1101, as ‘apoense’; --Merr., EPFP 3 (1923) 246; --Copel., PJS 47 (1932) 79; --Sleumer, Reinwardtia 4 (1957) 139; --Type: Elmer 11676b (PNH, holo, lost; BISH, BO, E, FI, K, L, MO, NY, US, iso), Mindanao: Davao, Mt Calelan. Endemic to the Philippines. MINDANAO: Davao (Mt Calelan), Bukidnon (Mt Lipa). In dwarfed mossy forest, on exposed peaks and ridges, c. 2500m. Fl. Sep.
  2. Diplycosia bartolomei Ferreras & Argent, Edinburgh Journal of Botany 68 (2011) 39; --Type: U. Ferreras & N. Bartolome UF475 (PNH, holo; E, L, iso), Luzon: Aurora Province, Central Sierra Madre Mountains, Mt. Mingan. Endemic to the Philippines. AURORA. Photos
  3. Diplycosia coi Argent, PJS 142 (2013) 52; --Type: Argent 19972480 (PNH, holo; E, L, iso), cult. from Argent P30: Mindoro: Mindoro Oriental, Mt. Halcon, 25-Jan-2012. Endemic to the Philippines. MINDORO.
  4. Diplycosia heterophylla Blume Bijdr. (1826) 858; --Type: from Java.
    • var. latifolia (Blume) Sleumer, Reinwardtia 4 (1957) 156; --Diplycosia latifolia Blume, Bijdr. (1826) 858. --Diplycosia baclayensis Elmer, LPB 3 (1911) 1100; --Merr., EPFP 3 (1923) 246; --Copel., PJS 47 (1932) 78; --Type: Elmer 11595 (PNH, holo, lost; E, K, L, MO, NY, iso). S Thailand, Sumatra, Peninsular Malaysia, Banka, Java, Borneo (Sarawak, C& E, SE part), Philippines. MINDANAO: Davao (Mt Apo). Free standing treelet or shrub to 6m, or an epiphyte in montane forest, especially in mossy forest, 1000-2500m.
  5. Diplycosa kitangladensis P.W.Fritsch, Nat. Hist. Bull. Siam Soc. 62 (2017) 22; --Type: D.S. Penneys 2257 (PNH, holo; CAS, CMUH, E, US, iso), Mindanao: Bukidnon, Mt Kitanglad Range Natural Park, Lantapan, Brgy Songco, 23-Apr-2014, 2510m. Endemic to the Philippines. MINDANAO: Bukidnon. Conservation status: Critically Endangered (Fritsch & Amoroso 2017).
  6. Diplycosia loheri Merr., PJS 27 (1925) 44; EPFP 4 (1925) 253; --Copel., PJS 47 (1932) 67; --Sleumer, Reinwardtia 4 (1957) 131; --Type: Loher 13693 (PNH, holo, lost; E, fragment of type; US). Endemic to the Philippines. LUZON: Nueva Vizcaya (Caraballo Mtns). Note by L.L.Co: Only known from the type collection.
  7. Diplycosia luzonica (A.Gray) Merr., PJS 2 c (1907) Bot. 293; EPFP 3 (1923) 247; --Copel., PJS 47 (1932) 71. --Sleumer, Reinwardtia 4 (1957) 143; --Gaultheria luzonica A.Gray, Proc. Amer. Acad. Arts Sc. 5 (1861) 324; --Type: USS Pacif. Expl. Exp. (GH, holo; US, iso). --Diplycosia scandens Merr., PJS 1 (1906) Suppl. 219; EPFP 3 (1923) 247; --Type: Merrill 4597 (cited as ‘4579’) (PNH, holo, lost; BO, K, L, NY, US, iso), Luzon: Mountain Province, Mt Data. --Diplycosia fasciculiflora Merr., PJS 10 c (1915) Bot. 52; EPFP 3 (1923) 246; --Type: BS 37811 Ramos & Edaño (PNH, holo, lost), Luzon: Ifugao prov., Mt Polis. Endemic to the Philippines.
    • var. luzonica. LUZON: Abra, Kalinga (Balbalasang), Mountain Province (Mt Data; Mt Pukis), Ifugao (Mt Polis), Benguet (Mt Simacoco; Mt Pulag; Mt Pulogloco; Mt Baudan), Laguna (Mt Banahaw), CATANDUANES (Mt Mariguidon), BILIRAN (Mt Suiro), NEGROS: Negros Occidental (Kanlaon Volcano), Negros Oriental (Cuernos Mtns), MINDANAO: Misamis Occidental (Mt Malindang). Both in mossy forest or forest edge, and on dry exposed grassland-slope, 1100-2700m. Fl. & fr. Jan-Dec. Photos
    • var. calelanensis (Elmer) Sleumer, Reinwardtia 4 (1957) 144; --Diplycosia calelanensis Elmer, LPB 3 (1911) 1103; --Merr., EPFP 3 (1923) 246; --Copel., PJS 47 (1932) 74; --Type: Elmer 11681 (US, holo; A, BISH, BO, CAL, E, FI, K, L, MO, NY, US, iso), Mindanao: Davao, Mt Calelan, epiphytic shrub on stunted trees or shrubs on the summit, 8500ft, Sep-1909. --Diplycosia glabra Merr., PJS 14 (1919) 442; EPFP 3 (1923) 247; --Type: BS 31784 J.K. Santos (PNH, holo, lost; A, BO, CAL, K, L, iso), Luzon: Benguet prov., Pauai. --Diplycosia opaca C.B.Rob., PJS 6 c (1911) Bot. 355; --Merr., EPFP 3 (1923) 247; --Copel., PJS 47 (1932) 77; --Type: BS 9385 Robinson (PNH, holo, lost; US, iso), Luzon: Quezon prov., Infanta, Mt Binuang, Aug-1909. LUZON: Benguet (Mt Sto Tomas; Mt Ugo; Pauai), Rizal (Montalban; Pinauisan), Quezon (Mt Binuang), Sorsogon, PALAWAN (Mt Mantalingahan), MINDANAO: Davao (Mt Calelan); South Cotabato (Mt Matutum). In forest, especially in dwarfed mossy forest on exposed peaks and ridges, 900-2500m.
    • var. merrittii (Merr.) Sleumer, Reinwardtia 4 (1957) 144; --Diplycosia merrittii Merr., PJS 2 c (1907) Bot. 293; EPFP 3 (1923) 247; --Type: Merrill 5670 (PNH, holo, lost; K, L, NY, US, iso), Mindoro: Mindoro Oriental prov., Mt Halcon, Nov-1906. MINDORO: Mindoro Oriental (Mt Halcon), MINDANAO: Bukidnon (Mt Lipa); reported from PALAWAN: Mt Victoria by Copeland.
    • var. pubens Sleumer, Reinwardtia 4 (1957) 143; --Type: BS 40397 Ramos & Edaño (L, holo; A, BO, K, SING, US, iso), Luzon: Benguet prov., Mt Pulogloco, Sep-1921. LUZON: Benguet (Mt Pulag; Mt Pulogloco).
  8. Diplycosia parvifolia Merr. PJS 5 c (1910) Bot. 211; EPFP 3 (1923) 247; --Copel., PJS 47 (1932) 81; --Sleumer, Reinwardtia 4 (1957) 137; --Type: Merrill 6995 (PNH, holo, lost; K, L, iso), Negros: Negros Occidental prov., Mt Kanlaon, c. 1500m. Endemic to the Philippines. LUZON: Ifugao (Mt Polis); NEGROS: Negros Occidental (Mt Kanlaon). In mossy forest, 1500-1855m.
  9. Diplycosia platyphylla P.W.Fritsch, PhytoKeys 69 (2016) 31; --Type: D.S. Penneys 2302 (PNH, holo; CAS-1196249, CMUH-00010804, iso), Mindanao: North Cotabato Prov., Mt Apo, Barangay Ilomavis, Matingao River Watershed, 29-Apr-2014. Endemic to Philippines. MINDANAO: North Cotabato. Only known from type. Conservation status: Critically Endangered (Fritsch & Amoroso, 2016). Photos
  10. Diplycosia trinervia Elmer, LPB 3 (1911) 1102; --Merr., EPFP 3 (1923) 247; --Copel., PJS 47 (1932) 68; --Sleumer, Reinwardtia 4 (1957) 142; --Type: Elmer 11676a (US, lecto; E, K, L, NY, PNH, isolecto), Mindanao: Davao del Sur prov., Todaya, Mt Apo, elfinwoods at the summit of Mt Calelan, Sep-1909. --Diplycosia luzonica Elmer, LPB 3 (1911) 1105, nom. illeg., auct. non (A.Gray.) Merr. Endemic to the Philippines. MINDANAO: Davao (Mt Calelan). In dwarfed mossy forest, c. 2500m. Fl. Sep. Only known from the type (Fritsch & Amoroso 2017).
  11. Diplycosia urdanetensis Elmer, LPB 7 (1915) 2628; --Merr., EPFP 3 (1923) 247; --Diplycosia trinervia var. urdanetensis (Elmer) Sleumer, Reinwardtia 4 (1957) 143; --Type: Elmer 13800 (PNH, lecto, lost?; A, BISH, BO, CAL, E, GH, HBG, K, L, MO, NA, NY, P, U, UC, US, isolecto), Mindanao: Agusan del Norte prov., Mt Urdaneta. Mossy forest, epiphytic on mossy trunk, c. 2000m, Sep-1912. MINDANAO: Agusan del Norte. Mossy forest, c. 2000m. Fl. Sep. Known only from the type collection (Fritsch & Amoroso 2017).

Gaultheria Kalm ex L.

Photos
  1. Gaultheria borneensis Stapf, Trans. Linn. Soc. II, Bot. 4 (1894) 190; --Merr., EPFP 3 (1923) 246; --Sleumer, Reinwardtia 4 (1957) 163; --Type: from Borneo. --Gaultheria itoana Hayata, Mat. Fl. Formosa (1908) 150. --Gaultheria benguetensis H.F.Copel., PJS 47 (1932) 58; --Type: Merrill 4796 (PNH, holo, lost; K, NY, US, iso), Luzon: Benguet prov., Pauai to Baguio, Oct/Nov-1905. Borneo (Sabah: Mt Kinabalu), Philippines. LUZON: Mountain Province (Mt Data), Benguet (Halsema Road; Pauai; Mt Pulag). Prostrate, not rarely cushion-like shrublet in ± open places along pine forest edges, also on talus along road, 1800-2400m. Fl. & fr. Jan-Dec. Photos
  2. Gaultheria leucocarpa Blume, Bijdr. (1826) 856. Photos
    • var. leucocarpa forma cumingiana (S.Vidal) Sleumer, Reinwardtia 4 (1957) 179; --Gaultheria cumingiana S.Vidal, Phan. Cuming. Philip. (1885) 184; --Merr., EPFP 3 (1923) 246, pro parte; --Copel., PJS 47 (1932) 61; --Type: Cuming 934 (cited as ‘932’, MA, holo; BM, K, L, MEL, iso), Luzon: Albay prov., Mt Mayon, 1841. Upper Myanmar, SW and S China, N Vietnam, Thailand, Taiwan, Peninsular Malaysia, Sumatra, E Java, Philippines. LUZON: Kalinga (trail to Balbalasang), Benguet (Mt Sto Tomas; Baguio; Baguio-Ambuklao Road; Mt Pulag; Buguias; Abatan; Bucao; Halsema Road), Mountain Province (Mt Data; Besao, Agawa; Datakan), Ifugao (Mt Polis), Zambales (Mt Pinatubo), Laguna (Mt Banahaw; Mt San Cristobal), Albay (Mt Mayon). Usually in open, exposed sites in the montane zone (common in the pine belt of the Cordillera Highlands), c. 900-2500m. On steep slopes, exposed outcrops along roadcuts, crater margins, gravelly ground, volcanic or occasionally peaty and generally poor soil, occasionally epiphytic. Fl. Jan-Dec.
    • var. psilocarpa (H.F.Copel.) Sleumer, Reinwardtia 4 (1957) 183; --Gaultheria psilocarpa H.F.Copel., PJS 47 (1932) 62; --Type: BS 38903 Ramos & Edaño (PNH, holo, lost; A, BM, BRI, L, iso), Mindanao: Bukidnon prov., Mt Candoon. --Gaultheria cumingiana auct. non S.Vidal; --Merr., EPFP 3 (1923) 246, pro parte; LUZON: Laguna (Mt Mahayhay, probably within Mt Banahaw complex), MINDORO: Mindoro Oriental (Mt Halcon), NEGROS: Negros Occidental (Kanlaon Volcano), MINDANAO: Bukidnon (Mt Candoon), Davao (Mt Calelan). 1800-2800m.

 Iaera H.F.Copel. = Costera

Rhododendron L.

Last edited by Pieter B. Pelser, 18 August 2017

Photos
  1. Rhododendron acrophilum Merr. & Quisumb., PJS 82 (1953) 333; Sleumer, Reinwardtia 5 (1960) 130; --Type: PNH 34 Edaño (A, holo; BM, K, L, PNH, US, iso), Palawan: Mt Mantalingahan. Mossy forest, c. 1725m, 13-May-1947. PALAWAN: Mt Mantalingahan. Known only from the type collection. Fl. May. Conservation status: Critically Endangered (DAO 2017-11).
  2. Rhododendron apoanum Stein, Gartenflora 34 (1885) 194; --Merr., EPFP (1923) 243; --Copel., PJS 40 (1929) 146; --Sleumer, Reinwardtia 5 (1960) 99; --Type: Schadenberg s.n. (B, holo, lost), Mindanao: Davao, Mt Apo, 1882. MINDANAO: Davao (Mt Apo and adjacent higher peaks), Bukidnon (Mt Lipa; Mt Kitanglad), Agusan del Norte (Mt Hilong-hilong). Mossy forest on peaks and ridges, 1380-2930m, apparently not rare on Mt Apo. Fl. Mar, May-Oct.
  3. Rhododendron bagobonum H.F.Copel., PJS 40 (1929) 151; Sleumer, Reinwardtia 5 (1960) 154; --Type: M.S. Clemens s.n. (UC, holo), Mindanao: Davao, Mt Apo, Jun-1924. --Rhododendron cuneifolium Stapf var. subspathulatum Ridl., J. St. Br. R. As. Soc. 33 (1900) 23. --Rhododendron galioides J.J.Sm., Bull. Jard. Bot. Btzg. 3, 13 (1935) 445. Borneo (Sarawak; C and E Borneo), Philippines. PALAWAN (Mt Mantalingahan), CAMIGUIN (Mt Hibok-hibok), MINDANAO: Davao (Mt Apo). Probably an epiphyte, other local ecological notes not clearly stated. Those given by Sleumer are largely based on Borneo: in primary mossy forest on mountain ridges, heath forest, mostly open sunny places, 900-2135m. Fl. Jan-Dec.
  4. Rhododendron brachygynum H.F.Copel., PJS 40 (1929) 137; --Sleumer, Reinwardtia 5 (1960) 219; --Type: FB 28346 Cenabre & de la Cruz (PNH, holo, lost), Cebu: Sudlon. CEBU: Sudlon. Known only from the type collection.
  5. Rhododendron edanoi Merr. & Quisumb., PJS 83 (1953) 333; --Type: PNH 153 Edaño (A, holo; BM, K, L, PNH, US, iso), Palawan: Mt Mantalingahan. Mossy forest, 1725m, 13-May-1947. --Rhododendron pneumonanthum Sleumer, Reinwardtia 5 (1960) 132.
    • ssp. edanoi. PALAWAN: Cleopatra Needle; Thumb Peak; Mt Mantalingahan. Fl. May. Conservation status: Vulnerable (DAO 2017-11).
  6. Rhododendron jasminiflorum Hook., Bot. Mag. (1850) t. 4524.
    • var. copelandii (Merr.) Sleumer, Reinwardtia 5 (1960) 130; --Rhododendron copelandii Merr., Publ. Govt. Lab. No. 29 (1905) 42; --Copel., PJS 40 (1929) 158; --Type: E.B. Copeland 1439 (PNH, holo, lost; L, lecto; A, K, NY, P, US, isolecto), Mindanao: Davao, Mt Apo summit thickets, Oct-1904. MINDANAO: Davao (Mt Apo). High-montane thickets, 2350-3100m. Fl. Mar-Nov. Conservation status: Vulnerable (DAO 2017-11). Photos
  7. Rhododendron javanicum (Blume) Benn. in Benn. & Br., Pl. Jav. Rar. (1838) 85. Photos
    • ssp. palawanense Argent, Gard. Bull. Sing. 56 (2004) 90; --Type: Argent & Romero Acc. 19922770 (PNH, holo; E, iso), Palawan: Mt Mantalingahan, cultivated specimen collected 16-Jan-1998. PALAWAN (Cleopatra Needle; Thumb Peak; ?Victoria Peak; Mt Mantalingahan). Conservation status: Critically Endangered (DAO 2017-11).
    • ssp. schadenbergii (Warb.) Argent, Rhododendrons of Subgenus Vireya (2006) 247; --Rhododendron javanicum var. schadenbergii (Warb.) Sleumer, Reinwardtia 5 (1960) 195; -- Rhododendron schadenbergii Warb. in Perkins, Fragm. Fl. Philip. 3 (1905) 172; --Merr., EPFP 3 (1923) 245, pro parte; --Copel., PJS 40 (1929) 170; --Type: Schadenberg s.n. (B, holo, lost), Luzon: Abra prov., 1300m. --Rhododendron spectabile Merr., Publ. Govt. Lab. Philipp. no. 29 (1905) 42; --Merr., EPFP 3 (1923) 245; --Copel., PJS 40 (1929) 168; --Type: E.B. Copeland 1438 (PNH, lecto, lost; A, E, fragm. ex PNH, K, US, iso), Mindanao: Davao, Mt Apo, Oct-1904. --Rhododendron clementis Merr., PJS 3 c (1908) Bot. 160, 381; EPFP 3 (1923) 243; --Copel., PJS 40 (1929) 167; --Type: Clemens 732 (PNH, holo, lost; US, iso), Mindanao: Lanao del Sur prov., Camp Keithley, Oct-1906. LUZON: Apayao (Mt Magnas), Kalinga (Tinglayan), Ifugao (Mt Polis), Abra, Mountain Province, Benguet, PALAWAN (Mt Mantalingahan), CAMIGUIN (Hibok-hibok Volcano), JOLO, MINDANAO: Zamboanga (Sax River); Lanao del Sur (Camp Keithley, along Lake Lanao), Bukidnon (Kaatoan), Davao (Mt Apo; Mt Calelan), South Cotabato (Mt Matutum). Low to high montane primary and secondary forests, also near volcanic craters, narrow ridge crests, steep slopes, in rather dry grounds, between rocks, along streams, etc., 600-2900m. Mostly epiphytic, but on Mt Apo also reported as a terrestrial shrub. Conservation status: Endangered (DAO 2017-11).
  8. Rhododendron kochii Stein, Gartenflora 34 (1885) 193; --Merr., EPFP 3 (1923) 243; --Copel., PJS 40 (1929) 160; --Sleumer, Reinwardtia 5 (1960) 212; --Type: Schadenberg s.n. (B, holo, lost); Elmer 11435 (L, neo; A, BM, BO, E, FI, GH, K, NY, U, iso), Mindanao: Davao, Mt Apo, Todaya, Aug-1909. LUZON: Mountain Province, Bataan (Mt Mariveles), Laguna (Mt San Cristobal; Mt Banahaw), MINDORO: Mindoro Oriental (Mt Halcon; Ilong Peak), NEGROS: Negros Occidental (Mt Kanlaon), MINDANAO: Davao (Mt Apo; Mt McKinley), Lanao del Sur (Camp Keithley along Lake Lanao), Misamis Occidental (Mt Malindang). Copeland also mentions BS 692 Foxworthy from PALAWAN (Mt Victoria) as possibly conspecific but because material is merely in fruit, cannot be positively identified. Nonetheless, this is the only Rhododendron seen by him from PALAWAN. On ridges on mossy forest, 1000-2300m. Fl. Jan-Dec. Conservation status: Vulnerable (DAO 2017-11). Photos
  9. Rhododendron leytense Merr., PJS 10 c (1915) 55; EPFP 3 (1923) 244; --Copel., PJS 40 (1929) 165; --Sleumer, Reinwardtia 5 (1960) 218; --Type: BS 15252 Ramos (PNH, holo, lost; BM; E, fragm. ex PNH; K, L, P, US, iso), Leyte: Mt Ibuni (back of Dagami). --Rhododendron flaviflorum Elmer ex Merr., EPFP 3 (1923) 244.
    • var. leytense. LUZON: Sorsogon (Mt Bulusan), LEYTE: Mt Ibuni (back of Dagami), Mt Lobi, CAMIGUIN. Mossy forest, probably above 1000m.
    • var. loheri (H.F.Copel.) Sleumer, Reinwardtia 5 (1960) 218; --Rhododendron loheri H.F.Copel., PJS 40 (1929) 166; --Type: Loher 14769 (PNH, holo, lost; E fragm. ex PNH, UC), Luzon: Rizal prov., Guiniuisan. LUZON: Rizal (Guiniuisan). Known only from the type collection.
  10. Rhododendron loboense H.F.Copel., PJS 40 (1929) 137; Sleumer, Reinwardtia 5 (1960) 130; --Type: FB 28045 Mabesa (PNH, holo, lost; A, E, fragm.), Luzon: Batangas prov., Lobo Mtns. LUZON: Batangas (Lobo Mtns), MINDORO: Mindoro Oriental (Mt Halcon), CAMIGUIN (Mt Hibok-hibok). Fl. Apr. Conservation status: Vulnerable (DAO 2017-11).
  11. Rhododendron madulidii Argent, New Plantsman 5(4) (1998) 204. PALAWAN: Mt Mantalingahan, near Brookes Point. Conservation status: Endangered (DAO 2017-11). Photos
  12. Rhododendron mendumiae Argent, Gard. Bull. Sing. 56 (2004) 82. PALAWAN: Cleopatra Needle. Conservation status: Critically Endangered (DAO 2017-11).
  13. Rhododendron mindanaense Merr., Publ. Govt. Lab. Philipp. no. 29 (1905) 41; EPFP 3 (1923) 244; --Copel., PJS 40 (1929) 164; Sleumer, Reinwardtia 5 (1960) 219; --Type: E.B. Copeland 1042 (PNH, holo, lost); De Vore & Hoover 73 (PNH, para, lost; A, E fragm. ex PNH). MINDANAO: Mt Apo. Mossy dwarfed shrub vegetation near summit, 2700-2895m. Fl. Jul-Aug. Photo
  14. Rhododendron nortoniae Merr., PJS 1 (1906) Suppl. 220; EPFP 3 (1923) 244; --Copel., PJS 40 (1929) 149; --Sleumer, Reinwardtia 5 (1960) 103; --Type: Clemens 500 (PNH, holo, lost; E fragm., sterile), Mindanao: Lanao del Sur prov., Camp Keithley, 800m. --Rhododendron catanduanense Merr., in Copel., PJS 40 (1929) 149; --Type: BS 30346 Ramos (PNH, holo, lost; E fragm., sterile, US, iso), Catanduanes: Mt Mariguidon, 270m, Nov/Dec-1917. CATANDUANES (Mt Mariguidon, 270m), NEGROS: Negros Oriental (Cuernos Mtns), MINDANAO: Lanao del Sur (Camp Keithley, 800m). Photos
  15. Rhododendron quadrasianum S.Vidal, Rev. Pl. Vasc. Filip. (1886) 170; --Copel., PJS 40 (1929) 140; –Merr., EPFP 3 (1923) 244; --Sleumer, Reinwardtia 5 (1960) 65; --Type: Vidal (K, holo), Luzon: Albay prov., Mt Mayon. Photos
    • var. quadrasianum. LUZON: Camarines Sur (Mt Isarog), Albay (Mt Mayon), MINDORO. Rocky open slopes, 1330-2100m, abundant. Fl. & fr. May-Jun.
    • var. davaoense (H.F.Copel.) Sleumer, Reinwardtia 5 (1960) 65; --Rhododendron quadrasianum forma davaoense H.F.Copel., PJS 40 (1929) 143; --Type: Williams 2543 (PNH, holo, lost; GH, K, L, NY, US, iso), Mindanao: Davao, Mt Apo, 31-Mar-1905. --Rhododendron quadrasianum forma negrosense H.F.Copel., PJS 40 (1929) 142; --Type: Merrill Philip. Pl. 247 (PNH, holo, lost; FI, L, U, US, iso), Negros: Negros Occidental prov., Mt Kanlaon, Apr-1910. LUZON: Laguna-Quezon (Mt Banahaw; Mt Mahayhay), Sorsogon (Mt Bagaua), LEYTE, NEGROS: Negros Occidental (Mt Kanlaon; Mt Mapara), Negros Oriental (Cuernos Mtns), MINDANAO: Davao (Mt Apo), South Cotabato (Mt Matutum). Dense mossy forest on summits, 1600-2440m. Fl. & fr. Jan-Dec.
    • var. intermedium Merr., PJS 3 c (1908) Bot. 382; EPFP 3 (1923) 244; --Copel., PJS 40 (1929) 143; --Sleumer, Reinwardtia 5 (1960) 66; --Type: FB 8063 Curran & Merritt (PNH, holo, lost; US, iso), Luzon: Zambales prov., Nov/Dec-1907. LUZON: Abra (Mt Paraga), Mountain Province (Mt Sisipitan), Benguet, Zambales (Mt Tapulao; Mt Pinatubo). Open exposed ridges, 2000-2400m.
    • var. malindangense (Merr.) H.F.Copel., PJS 40 (1929) 142; --Sleumer, Reinwardtia 5 (1960) 66; --Rhododendron malindangense Merr., PJS 3 c (1908) Bot. 256; EPFP 3 (1923) 244; --Type: FB 4705 Mearns & Hutchinson (PNH, holo, lost; BO, K, NY, US, iso), Mindanao: Misamis Occidental prov., Mt Malindang, May-1906. CAMIGUIN (Mt Hibok-hibok), MINDANAO: Misamis Occidental (Mt Malindang), Bukidnon (Mt Candoon; Mt Kitanglad), Agusan del Norte (Mt Hilong-hilong); Mossy forest, 1000-1600m. Fl. & fr. Mar-Sep.
    • var. marivelesense (H.F.Copel.) Sleumer, Reinwardtia 5 (1960) 66; --Rhododendron quadrasianum forma marivelesense H.F.Copel., PJS 40 (1929) 141; --Type: Merrill 3215 (PNH, holo, lost; BM, K, L 2 sheets, NY, P, US, iso), Luzon: Bataan prov., Mt Mariveles, Lamao River, Oct-1903. LUZON: Bataan (Mt Mariveles), Laguna (Mt Makiling; Mt San Cristobal), Batangas (Mt Agas), MINDORO: Mindoro Oriental (Mt Halcon), LEYTE (Wenzel 930). Mossy forest, 1200-2500m. Fl. & fr. Mar-Nov.
    • var. rosmarinifolium (S.Vidal) H.F.Copel., PJS 40 (1929) 144; --Merr., EPFP 3 (1923) 245; --Sleumer, Reinwardtia 5 (1960) 65; --Rhododendron rosmarinifolium S.Vidal, Rev. Pl. Vasc. Filip. (1886) 172; --Merr., EPFP 3 (1923) 245; --Type: Vidal 1530 (K, holo; A fragm., FI, L 2 sheets, iso), Luzon: Mountain Province, Bontoc. --Rhododendron quadrasianum var. intermedium Merr., PJS 3 c (1908) Bot. 382, pro parte; EPFP 3 (1923) 244, pro parte; --Sleumer, Reinwardtia 5 (1960) 65. --Rhododendron quadrasianum forma halconense H.F.Copel., PJS 40 (1929) 141; --Type: Merrill 5736 (PNH, holo, lost; K, L, NY, P, US, iso), Mindoro: Mindoro Oriental prov., Mt Halcon, Nov-1906. --Rhododendron quadrasianum forma banahaoense H.F.Copel., PJS 40 (1929) 145; --Type: Cuming 804 (PNH, holo, lost; BM, CGE, FI, K, L, MEL, iso), Luzon: Laguna prov., Mt Banahaw. --Rhododendron quadrasianum forma pulogense H.F.Copel., PJS 40 (1929) 144; --Type: BS 44944 Ramos & Edaño (PNH, holo, lost; A, BRI, L, NY, UC, iso), Luzon: Benguet prov., Mt Pulag, Feb/Mar-1925. LUZON: Benguet (Mt Sto Tomas; Baguio; La Trinidad; Pauai; Mt Pulag), Mountain Province (Bauko; Mt Data; Bontoc; Bontoc-Banaue Road; Mt Caua), Ifugao (Mt Polis); Zambales Range (Mt Pinatubo); Laguna-Quezon prov. (Mt Banahaw), MINDORO: Mindoro Oriental (Mt Halcon; Mt Dulangan; Ilong Peak), BILIRAN (Mt Suiro), NEGROS. Mossy (oak) forest, also in secondary growth and open places, locally common, 1300-2400m. Fl. & fr. Jan-Dec.
  16. Rhododendron reynosoi Argent, Gard. Bull. Sing. 56 (2004) 84. PALAWAN: Cleopatra Needle. Conservation status: Critically Endangered (DAO 2017-11).
  17. Rhododendron rousei Argent & Madulid, New Plantsman 5 (1998) 25; --Type: Argent 8984 & E. Reynoso. SIBUYAN: Mt Giting-giting. 
  18. Rhododendron x sarcodes Argent & Madulid, New Plantsman 2 (1995) 156. MINDANAO: North Cotabato (Mt Apo), 1800m.
  19. Rhododendron subsessile Rendle, J. Bot. 34 (1896) 357; --Merr., EPFP 3 (1923) 245; --Copel., PJS 40 (1929) 174; --Sleumer, Reinwardtia 5 (1960) 228; --Type: Whitehead s.n. (BM, holo), Luzon: Mountain Province, Mt Data. --Rhododendron subsessile var. baucoense H.F.Copel., PJS 40 (1929) 175. LUZON: Abra (Mt Paraga), Mountain Province (Mt Data; Bauko; Mt Sisipitan), Ifugao (Mt Polis; Banaue), Benguet (Pauai; Halsema Road; Mt Malaya; Mt Pulag; Mt Baudan; Bucao; Baguio; Mt Sto Tomas). Rather common in the pine region and the pine-oak association below the mossy forest, rare in the mossy forest (Mt Pulag), again on exposed grass-covered slopes above the moss forest, often in clumps or masses, 1300-2400m. Fl. & fr. Jan-Dec. Conservation status: Vulnerable (DAO 2017-11). Photos
  20. Rhododendron taxifolium Merr., PJS 30 (1926) 419; --Copel., PJS 40 (1929) 156; -- Sleumer, Reinwardtia 5 (1960) 141; --Type: BS 44880 Ramos & Edaño (PNH, holo, lost; A, BM, E type fragm., K, P), Luzon: Benguet prov., Mt Pulag. LUZON: Benguet (Mt Pulag), 2700m. Epiphytic shrub on trees in the mossy forest. Fl. Feb-Mar. Conservation status: Critically Endangered (DAO 2017-11). Photos
  21. Rhododendron vidalii Rolfe, J. Bot. 24 (1886) 348; --Merr., EPFP 3 (1923) 245; --Copel., PJS 40 (1929) 152; --Sleumer, Reinwardtia 5 (1960) 151; --Type: Vidal 1529 (K, holo; FI, L), Luzon: Mountain Province, Bontoc. --Rhododendron curranii Merr., PJS 3 c (1908) Bot. 255; EPFP 3 (1923) 243; --Type: FB 8061 Curran & Merritt (PNH, lecto, lost; US). Photos
    • ssp. brachystemon Argent, Folia Malesiana 4 (2003) 101. MINDORO: Mindoro Oriental (Mt Halcon).
    • ssp. vidalii. LUZON: Mountain Province (Bontoc; Bontoc-Banaue Road; Mt Caua; Mt Data; Bauko, Sabangan), Benguet (Loo; Bandschan; Tabbak), Abra, Cagayan (Mt Cagua), Isabela (Bayabat), Zambales (Mt Tapulao), Bataan (Mt Mariveles), Laguna (Mt Makiling), Quezon (Mt Banahaw de Lucban), Batangas. (Mt Malarayat). Mostly in mossy forest, but also terrestrial on banks in rather barren places or grassland, epiphytic on large, moss covered trees or terrestrial on shrubby mountain side or exposed ridges, locally not rare, 1000-2100m. Fl. Jan-Dec. Conservation status: Vulnerable (DAO 2017-11).
  22. Rhododendron whiteheadii Rendle, J. Bot. 34 (1896) 356; --Merr., EPFP 3 (1923) 245; --Copel., PJS 40 (1929) 154; --Sleumer, Blumea 21 (1973) 357--Type: Whitehead s.n. (BM, holo), Luzon: Ifugao prov., Mt Polis. LUZON: Ifugao (Mt Polis), Benguet (Mt Tabayoc; Mt Pulag), Zambales (Mt Tapulao). Photos
  23. Rhododendron wilkei Argent, Gard. Bull. Sing. 56 (2004) 88. PALAWAN: Thumb Peak. Conservation status: Vulnerable (DAO 2017-11).
  24. Rhododendron williamsii Merr. ex H.F.Copel., PJS 40 (1929) 137; --Sleumer, Reinwardtia 5 (1960) 190; --Type: Elmer 6519 (US, holo; K, NY, P), Luzon: Benguet prov., Baguio. --Rhododendron schadenbergii auct. non Warb.; --Merr., EPFP 3 (1923) 245, pro parte. LUZON: Mountain Province (Mt Data; Mt Pukis), Benguet (Baguio; Mt Sto Tomas; Tabio), Zambales (Mt Pinatubo). Along streams and ravines, 1500-2200m. Photos
  25. Rhododendron xanthopetalum Merr., Publ. Govt. Lab. Philipp. no. 29 (1905) 41; --Merr., EPFP 3 (1923) 246; --Copel., PJS 40 (1929) 169; --Type: Whitford 322 (PNH, holo, lost). LUZON: Nueva Vizcaya (Caraballo Sur Mtns), Bataan (Mt Mariveles), Rizal (Montalban), MINDORO: (Ibolo). In Bataan, an epiphyte in mossy forest, 1200m. Note by L.L.Co: An imperfectly known species, apparently rare. Conservation status: Endangered (DAO 2017-11).
  26. Rhododendron zollingeri J.J.Sm., Icon. Bog. 4 (1910) 73; --Sleumer, Reinwardtia 5 (1960) 109; --Type: from Java. Java, Bali, Lombok, Sulawesi, N Philippines. LUZON: Benguet (Loo, Mt Bandschan). Reported elsewhere in Malesia on bare open slopes or ridges, exposed summits, volcanic rocks or crater walls, in fern-, grass-, and scrub vegetation or open forest, in Casuarina forest, on dry places, on sandy, stony, or ashy soil, also near solfataras, common locally, (1200-)1800-3090m. Fl. & fr. predominantly in the dry season Apr-Oct.

Ined. name:

  1. Rhododendron brookeanum Low ex Lindl., J. Hort. Soc. 3 (1848) 82.
    • ssp. palawananense. Palawan: Mt Mantalingahan (Argent M270. Very close to var. kinabaluense but differs in the longer flower-tubes, the almost glabrous ovary (lacking silvery scales as well as only having a few scattered simple hairs) and a longer style with a smaller stigma. Flowers orange with yellow star, highlighted with dark pink at the edge. Anthers purple.

Styphelia Sm.

Photos
  1. Styphelia malayana (Jack) Sprengel, Syst. 4 (1827) Cur. Post. 67, as ‘malaica’; --Sleumer, Fl. Males. ser. 1, (1964) 426. S Indochina, S Myanmar, S Thailand, Sumatra (W Coast), Peninsular Malaysia, Banka, Billiton and Riouw Archipelago, Anambas, Borneo incl. Karimata Archipelago, Philippines. MINDANAO: Davao Oriental prov., Mt Hamiguitan (Amoroso & Aspiras s.n., PUH). Ultramafic scrub vegetation, c. 1200m. Photos
  2. Styphelia suaveolens (Hook.f.) Warb. in Sarasin, Reisen. 2 (1905) 329; --Sleumer, Blumea 12 (1963) 148; --Sleumer, Fl. Males. ser. 1, (1964) 428. --Styphelia philippinensis Merr., PJS 20 (1922) 419; EPFP 3 (1923) 252. Borneo, Sulawesi, Timor, New Guinea, Solomon Isls, Australia (S Queensland; New South Wales; Victoria; Tasmania). LUZON: Benguet (between Suyoc and Pauai; Banguinoc, 2500m; Mt Singakalsa, 2300-2700m; Timbac Ridge), NEGROS (Kanlaon Volcano, 1950-2300m), MINDANAO: Davao (Mt Apo, dominant from 2600-2820m and Lake Linao, 2300m). Exposed ridges on the higher mountains, 2000-2820m. Photos

Vaccinium L.

Last edited by Pieter B. Pelser, 18 August 2017

Photos
  1. Vaccinium agusanense Elmer, LPB 7 (1915) 2630; --Merr., EPFP 3 (1923) 247; --Copel., PJS  42 (1930) 551; --Sleumer, Blumea 11 (1961) 49; --Type: Elmer 13765 (PNH, holo, lost; BISH, BM, BO, FI, GH, K, L, NY, US, iso), Mindanao: Agusan del Norte prov., Mt Urdaneta, epiphytic shrub upon a mossy wind-swept peak of the Maaay group at 5750ft (1752m), Sep-1912. MINDANAO: Agusan del Norte; Lanao del Sur, Bukidnon (Kabaritan; Mt Kitanglad). In semi-open mountain forest, or in mossy forest, 1100-1800m. Fr. Feb, Sep.
  2. Vaccinium alvarezii Merr., PJS 4 c (1909) Bot. 304; EPFP 3 (1923) 247; --Copel., PJS 42 (1930) 562; --Sleumer, Blumea 11 (1961) 87; --Type: FB 18466 (PNH, holo, lost), Luzon: Cagayan prov., Dalisay River. --Vaccinium turbinatum Merr., PJS 10 c (1915) Bot. 54; EPFP 3 (1923) 252; --Type: BS 15068 Ramos (US, holo; K, iso), Luzon: Laguna prov., San Antonio near Paete.
    • var. alvarezii. LUZON: Sierra Madre Range in Cagayan (Mt Cagua; Dalisay River), Rizal (Mt Irig = Irid; Angilog = ?Angilo or Angelo Mtns; Montalban), Laguna (Siniloan; San Antonio). Primary forest, (100-)300 to at most 1470m (Sleumer’s maximum citation of ‘2000m’ is improbable as the highest point among aforecited localities is 1470m on Mt Irid summit).
    • var. moisense H.F.Copel., PJS 42 (1930) 564; --Sleumer, Blumea 11 (1961) 88; --Type: BS 47303 Ramos & Edaño (UC, holo; BM, L, NY, P, SING), Luzon: Isabela prov., Mt Moises, epiphytic, 900m, Mar-1926. LUZON: Isabela (Mt Moises), epiphytic at 900m. Known only from the type collection.
  3. Vaccinium bancanum Miq., Fl. Ind. Bat. Suppl. 1 (1860) 587; --Type: from Banka. --Vaccinium suluense H.F.Copel., PJS 42 (1930) 568; --Type: Clemens 9337 (PNH, ?lost), Sulu.
    • var. bancanum. Sumatra, Lingga, Banka, Billiton, Anambas Isl, Borneo, Philippines. MINDANAO: Bukidnon (Mt Maraniag, FB 30277 Rojas, only collected in fruit and cited by Copeland as Vaccinium suluense, ?lost). SULU. On a hill (possibly with secondary forest, 10m) in Sulu, c. 730m in Bukidnon. Sleumer (1961, 1967) did not accredit this species to the Philippines, though Vaccinium suluense is included in the synonymy of Vaccinium bancanum.
  4. Vaccinium banksii Merr., Publ. Govt. Lab. Philipp. no. 35 (1906) 54; EPFP 3 (1923) 248; --Copel., PJS 42 (1930) 590. --Sleumer, Blumea 11 (1961) 81; --Type: Banks s.n. (PNH, holo, lost); --Type: Merrill 223 (US, FI, G, PNH, lost, neo). NEGROS: Negros Occidental (Mt Kanlaon). On ridges in the mossy forest, 1300-2000m.
  5. Vaccinium barandanum S.Vidal, Rev. Pl. Vasc. Filip. (1886) 169; --Merr., EPFP 3 (1923) 248; --Copel., PJS 42 (1930) 556; --Sleumer, Blumea 11 (1961) 48; --Type: Vidal 1532 (K, holo; FI, L, iso), Luzon: Lepanto (most likely in present-day W Mountain Province). Photos
    • var. barandanum. LUZON: Abra (Tue, near Sallapadan), Ilocos Sur (Cervantes), Mountain Province (W part, portion of the former Lepanto subprov; Mt Sisipitan; Mt Data; ?Tadian; Bucao; Mt Malaya), Benguet (Pauai and vicinity; Mt Pulag; La Trinidad to Tabio; Baguio; Mt Sto Tomas), Nueva Vizcaya (Caraballo Mtns); Zambales (Mt Pinatubo), Rizal (Sierra Madre Range towards Quezon and Bulacan borders; Montalban-Paningtingan area; Balacbac). In damp shaded ravines and in mossy forest, also in pine-oak association, reported to be tree-like on cliffs, 1300-2600m, locally not rare. Fl. & fr. Jan-Dec.
    • var. cagayanense H.F.Copel., PJS 42 (1930) 557; --Sleumer, Blumea 11 (1961) 49; --Type: BS 47300 Ramos & Edaño (UC, holo; A, BM, BO, L, NY, P, SING 2 sheets, iso), Luzon: Isabela prov., Mt Moises, Mar-1926. LUZON: Ifugao (Monhoyohoy), Benguet (Mt Pulogloco), Cagayan (Mt Cagua; Mt Dos Cuernos), Isabela (Mt Moises), Quirino-Aurora border (Mt Alzapan), Laguna (Luisiana).
    • var. hutchinsonii (Merr.) H.F.Copel., PJS 42 (1930) 558; --Sleumer, Blumea 11 (1961) 49; --Vaccinium hutchinsonii Merr., PJS 2 c (1907) Bot. 294; EPFP 3 (1923) 249; --Type: Merrill 5524 (PNH, holo, lost; NY, US, iso), Mindoro: Mindoro Oriental prov., Mt Halcon, Nov-1906. MINDORO: Mindoro Oriental (Mt Halcon). Epiphytic in mossy ridge forest, 1600-2000m. Fl. Nov. 
    • Possible undescribed variety: fide Sleumer, Blumea 11 (1961) 49; --Voucher: BS 75850 Edaño (BO, NY, PUH). LUZON: Camarines Sur (Kamugong River, 485 m). A plant corresponding to var. hutchinsonii in leaf characters, but with densely shortly grey-pubescent rhachis, pedicel, calyx and disk, appears to be an undescribed variety. However, can not yet be named as no flowers are present.
  6. Vaccinium benguetense S.Vidal, Rev. Pl. Vasc. Filip. (1886) 168; --Merr., EPFP 3 (1923) 248; --Copel., PJS 42 (1930) 566; --Sleumer, Blumea 11 (1961) 104; --Type: Vidal 1534 (K, lecto; FI, L, MA, isolecto); Vidal 1515 (syn), Luzon: Benguet prov. LUZON: Ilocos Norte (Mt Piao), Ilocos Sur (most probably E part), Abra (most probably E part), Mountain Province (Mt Data; Bauko; Mankayan; Mt Masapilid), Benguet (Buguias; Mt Pulag; Baguio), Zambales (Mt Pinatubo; Mt Tapulao), PANAY: Iloilo (Miag-ao, Mt Pulacan). Forested slopes and in ravines, 500-1600m. Fl. Jan-Dec.
  7. Vaccinium brachytrichum Sleumer, Blumea 11 (1961) 78; --Type: Merrill 9526 (not ‘BS 9526 Merrill’; Sleumer 1961) (L, holo; BM, BO, GH, K, NY, P, SING, US, iso), Palawan: Palawan prov., Mt Capoas, Apr-1913. --Vaccinium foxworthyi H.F.Copel., PJS 42 (1930) 580, pro parte. PALAWAN: Mt Capoas, 1000m. Fl. & fr. Apr. Only known from the type collection, unless conspecific with Elmer 13202, in fruit, from nearby Mt Pulgar. The latter is similar to the type but with narrower leaves. Conservation status: Vulnerable (DAO 2017-11).
  8. Vaccinium camiguinense Merr., PJS 7 c (1912) Bot. 321; EPFP 3 (1923) 248; --Copel., PJS 42 (1930) 578; --Sleumer, Blumea 11 (1961) 72; --Type: BS 14622 Ramos (PNH, holo, lost). CAMIGUIN (Mt Mahinog). Mossy forest near summit, c. 1000m. Fl. & fr. Mar-Apr.
  9. Vaccinium caudatum Warb. in Perkins, Fragm. Fl. Philipp. (1905) 173; --Merr., EPFP 3 (1923) 248; --Copel., PJS 42 (1930) 565; --Sleumer, Blumea 11 (1961) 104; Blumea 12 (1963) 137; --Type: Cuming 905 (B, lecto, lost; BM, FI, G, K, L), Luzon: Albay prov.; Warburg 13753 (B, syn, lost; E, syn), Luzon: Laguna prov., Siniloan. LUZON: Cagayan (Pinagtipunan River), Rizal (Montalban; Balacbac), Laguna (Siniloan; S. Bakong, Famy; Paete, San Antonio), Albay, MINDORO (Mag-asawang Tubig), LEYTE (Jaro), PANAY: Aklan (Libacao; Mt Salibongbong), Iloilo (Ulian River), MINDANAO: Surigao del Norte (Placer). Damp primary forests at low elevation, mostly below 500m. Fl. Jan-Dec. Photos
  10. Vaccinium cumingianum S.Vidal, Rev. Pl. Vasc. Filip. (1886) 167; --Merr., EPFP 3 (1923) 248; --Copel., PJS 42 (1930) 584; --Sleumer, Blumea 11 (1961) 82; --Type: Cuming 805 (PNH, lecto, lost; BM, G, K, L, P, iso), Luzon: Laguna prov., Mt Banahaw. Photos
    • var. cumingianum. LUZON: Rizal (Sierra Madre Range towards Quezon border; Montalban), Laguna-Quezon (Mt Banahaw; Mt San Cristobal), Quezon (Mt Banahaw de Lucban; Mt Calvario), Batangas (Lobo Mtns). A form with smaller leaves: Quezon-Aurora border (Umiray); Nueva Ecija (Mt Umingan = Mt Mingan), Rizal (Sierra Madre Range at Mabiluang). Montane forest, especially mossy forest, locally common on exposed ridges, (600-)1100-2000m. Fl. Jan-Dec.
    • var. igorotorum H.F.Copel., PJS 42 (1930) 586; --Sleumer, Blumea 11 (1961) 83; --Type: Elmer 5804 (US, holo; K, NY, P, iso), Luzon: Benguet prov., Mt Sto Tomas, Mar-1904. LUZON: Mountain Province (Bauko; Mt Sinapsapan; Mt Pukis), Benguet (Mt Nangaoto; Mt Singakalsa; Lusud-Bayabas trail, Mt Pulag; Mt Baudan; Pauai; Mt Sto Tomas; Baguio), La Union (Naguilian), Benguet-Nueva Vizcaya prov. border. 1500m to more than 2000m in the Cordillera Highlands of N LUZON.
    • var. marivelesense H.F.Copel., PJS 42 (1930) 586; --Sleumer, Blumea 11 (1961) 83; --Type: Whitford 459 (US, holo; K, NY, iso). LUZON: Zambales Range (general vicinity of Mt Pinatubo including Mt Abu); Bataan (Mt Mariveles).
    • var. pyriforme (Merr.) H.F.Copel., PJS 42 (1930) 586; --Sleumer, Blumea 11 (1961) 83; --Vaccinium pyriforme Merr., PJS 2 c (1907) Bot. 295; EPFP 3 (1923) 251; --Type: FB 4424 Merritt (PNH, holo, lost; K, iso), Mindoro: Mindoro Oriental prov., Mt Halcon. LUZON: Camarines Norte or Camarines Sur, Albay (Mt Mayon), MINDORO: Mindoro Oriental (Mt Halcon), PANAY: Capiz (Mt Bulilao). Epiphytic in forest on Mt Halcon, at least 1260-2700m on Mt Mayon.
    • Possible new variety: Voucher: L. Co 3800 (PUH, replicates to be distributed to A, L, US, K). LUZON: Ifugao (Mt Polis), high-montane forest, 1845 m. Closest to var. marivelesensis & var. pyriforme in having well-developed calyx lobes. Differing from the above in having a glabrous (non-ciliated calyx) and bigger leaves. Could be identical with BS 45660 Ramos & Edaño (Copeland 1930), probably an undescribed variety from north-central Luzon. Possibly lost at PNH in 1945 and hence not cited by Sleumer in his 1961 and 1966 revisions.
  11. Vaccinium elegans Elmer, LPB 3 (1911) 1903; --Merr., EPFP 3 (1923) 249; --Copel., PJS 42 (1930) 569; --Sleumer, Blumea 11 (1961) 69; --Type: Elmer 11683 (PNH, holo, lost; A, BM, FI, G, GH, K, L, NY, P, iso). MINDANAO: Davao (Mt Apo), Bukidnon (Mt Kitanglad), twice found. Mossy forest over ridges, 1980-2370m. Fl. Apr, Sep.
  12. Vaccinium epiphyticum Merr., PJS 7 c (1912) Bot. 322; EPFP 3 (1923) 249; --Copel., PJS 42 (1930) 555; --Sleumer, Blumea 11 (1961) 48; --Type: Merrill 8087 (PNH, holo, lost; BM, K, L, iso), Mindanao: Zamboanga (Sax River, Mtns above San Ramon), c. 900m. MINDANAO: Zamboanga. Known only from the type collection. Fl. Nov.
  13. Vaccinium gitingense Elmer, LPB 4 (1912) 1490; --Merr., EPFP 3 (1923) 249; --Copel., PJS 42 (1930) 553; --Sleumer, Blumea 11 (1961) 46; --Type: Elmer 12555 (PNH, holo, lost; A, BM, BO, DS, FI, G, GH, K, L, P, US, iso), Sibuyan: Mt Giting-giting. SIBUYAN, MINDANAO: Surigao del Norte (Tubungan-Dayan; Iron Deposit), DINAGAT. On hot, dry, gravelly river bottom (probably washed down from mountain forest, showing minor leaves) or along streams and on ridges at low elevation, ascending to 450m. Aforecited localities (especially Surigao and Dinagat) are predominantly ultramafic sites. Conservation status: Vulnerable (DAO 2017-11). Photos
  14. Vaccinium halconense Merr., PJS 2 c (1907) Bot. 294; EPFP 3 (1923) 249; --Copel., PJS 42 (1930) 578; --Sleumer, Blumea 11 (1961) 89; --Type: Merrill 5665 (PNH, holo, lost; K, NY, US, iso), Mindoro: Mindoro Oriental prov., Mt Halcon. LUZON: Zambales (Mt Tapulao; Mt Pinatubo), Bataan (Mt Mariveles), MINDORO: Mindoro Oriental (Mt Halcon; Ilong Peak), NEGROS: Negros Occidental (Mt Mapara), Negros Oriental (Cuernos Mtns); MINDANAO: Bukidnon (Tangkulan and vicinity), Davao (Mt McKinley), South Cotabato (Mt Matutum). In mossy forest, on exposed ridges, 1200-2135m. Fl. & fr. Jan-Dec. Photos
  15. Vaccinium indutum S.Vidal, Rev. Pl. Vasc. Filip. (1886) 169; --Merr., EPFP 3 (1923) 249; --Copel., PJS 42 (1930) 594; --Sleumer, Blumea 11 (1961) 35; --Type: Vidal 1831 (K, holo; L, MA, iso). --Vaccinium macgregorii Merr., PJS 10 c (1915) Bot. 53; EPFP 3 (1923) 250; --Type: BS 19846 McGregor (PNH, holo, lost; BM, K, P, US, iso), Luzon: Ifugao prov., Mt Polis. --Vaccinium sorsogonense Elmer, LPB 10 (1939) 3730; --Type: Elmer 17026 (PNH, holo, lost; BM, BO, FI, K, L, NY, UC, US, iso), Luzon: Sorsogon prov., Mt Bulusan. LUZON: Mountain Province (?W half; Malamey; Mt Caua), Ifugao (Mt Polis), Benguet (Mt Pulag), Sorsogon (Mt Bulusan). Rocky ground of densely wooded steep slopes, also in mossy forest, apparently generally above 1000m, rarely down to 600m. Fl. Feb-Aug. Photos
  16. Vaccinium irigaense Merr., PJS 10 c (1915) Bot. 52; EPFP 3 (1923) 249; --Copel., PJS 42 (1930) 576; --Sleumer, Blumea 11 (1961) 88; --Type: Philip. Pl. 1549 Ramos (PNH, holo, lost; BM, BO, BRI, G, GH, L, K, NY, P, SING, iso), Luzon: Camarines Sur prov., Mt Iriga. LUZON: Camarines Sur (Mt Iriga), Sorsogon (Mt Bulusan; Mt Poidol), CATANDUANES (Mt Mariguidon), BILIRAN (Mt Suiro). On ridges in mossy forest, 530-1000m.
  17. Vaccinium jagori Warb., in Perkins, Fragm. Fl. Philipp. (1905) 174; --Merr., EPFP 3 (1923) 250; --Copel., PJS 42 (1930) 592; --Sleumer, Blumea 11 (1961) 98; --Type: Jagor s.n. (B, holo, lost). --Vaccinium angustilimbum Merr., PJS 12 c (1917) 294; EPFP 3 (1923) 248; --Type: BS 26603 Ramos & Edaño (PNH, holo, lost; A, NY, US, iso), Luzon: Aurora prov., Mt Dingalan. LUZON: Abra, Mountain Province (Mt Malaya; Bauko), Benguet (Mt Nangaoto; Mt Natoo), Isabela, Aurora (Mt Dingalan), Zambales (Mt Tapulao = High Peak), Bataan (Mt Mariveles), Rizal (Sierra Madre Range at Montalban; Angilog (probably Angelo Mtns or Angilo); Mt Irid), Quezon (Mt Binuang), Camarines Sur (Mt Isarog). Exposed ridges in primary forest or in the mossy forest, (300-)1000-2000m. Photos
  18. Vaccinium luzoniense S.Vidal, Rev. Pl. Vasc. Filip. (1886) 168; --Merr., EPFP 3 (1923) 250; --Copel., PJS 42 (1930) 570; --Sleumer, Blumea 11 (1961) 77; --Type: Vidal 1535 (K, holo). LUZON: Mountain Province (W part; Mankayan), Ifugao-Mountain Province (South Inomin), Benguet (Loo; Pauai; Baguio; Mt Sto Tomas). Forested stream depressions and on ridges in the mossy forest, also on rocky banks by roadsides, 1500-2100m. Fl. & fr. Apr-Jul.
  19. Vaccinium microphyllum Reinw. ex Blume, Bijdr. (1826) 851; --Merr., EPFP 3 (1923) 250; --Copel., PJS 42 (1930) 600; --Sleumer, Blumea 11 (1961) 33; --Type: from the Moluccas. --Vaccinium mindorense Rendle, J. Bot. 34 (1896) 355. Philippines, Moluccas (Ternate; Tidore; Halmahera), Vanuatu (Espiritu Santo). LUZON: Laguna (Mt Banahaw), MINDORO: Mindoro Oriental (Mt Dulangan; Mt Halcon), NEGROS: Negros Occidental (Mt Kanlaon), PANAY: Antique (Mt Madia-as), MINDANAO: Davao (Mt Apo). Mossy exposed ridges and also epiphytic in mossy forest, locally common, 1220-2954m. Photos
  20. Vaccinium myrtoides (Blume) Miq., Fl. Ind. Bat. 2 (1859) 1062; --Merr., EPFP 3 (1923) 250; --Copel., PJS 42 (1930) 596; --Sleumer, Blumea 11 (1961) 51; Blumea 12 (1963) 127; --Thibaudia myrtoides Blume Bijdr. (1826) 861; --Type: from the Moluccas. --Vaccinium microphyllum auct. non Reinw. ex Blume; --Fern.-Vill., Novis. App. (1880) 121, pro parte. --Vaccinium varingiaefolium auct. non (Blume) Miq.; --S.Vidal, Sinopsis Atlas (1883) 30. --Vaccinium villarii S.Vidal, Rev. Pl. Vasc. Filip. (1886) 166. Philippines, N Sulawesi (Minahassa), Moluccas (Peak of Tidore). LUZON: Ilocos Sur (Cervantes trail), Mountain Province (Sagada; Mt Data; Bauko), Ifugao (Mt Polis), Benguet (Loo; Mt Pulag; Halsema Road; Pauai; Baguio); Laguna-Quezon (Mt San Cristobal; Mt Banahaw), Quezon, Albay (Mayon Volcano), MINDORO: Mindoro Oriental (Mt Halcon), MINDANAO: Davao (Mt Apo). On steep rocky, open and bare slopes, pine-mossy forest borders, mossy forest undergrowth, summit vegetation, open montane grassland, c. 1000-3000m. Locally abundant in Benguet and summit area of Mt Apo. Photos
  21. Vaccinium nitens Sleumer, Blumea 11 (1961) 97; --Type: PNH 17677 Sulit & Conklin (L, holo; BM, K, L, iso), Mindoro: Mindoro Oriental prov., Mt Yagaw, E slopes, steep rocky slope in virgin forest, 2500ft, 7-Jun-1953. MINDORO: Mindoro Oriental (Mt Yagaw). Steep rocky slope in virgin forest, 760m. Fl. Jul.
  22. Vaccinium oscarlopezianum Co, Edinburgh J. Bot. 59 (2002) 373; --Type: Co 3326 (PNH, holo; A, PUH, CAHUP, L, US, K, iso), Luzon: Isabela prov., San Mariano Mun., Brgy. Disulap, Dappig Creek, 16º56.5’N, 122º13’E. LUZON: Isabela (San Mariano), Aurora (Mtns NNW of Dingalan = Mingan Mtns). High-canopy epiphyte in fairly open sites like creeks and logging trails, 490-740m. Collected in flower in Feb and fruiting in Mar. Fairly common in the vicinity of the type locality. Thus far known from two sites that are c. 175(?) km apart. Photos
  23. Vaccinium palawanense Merr., PJS 3 c (1908) Bot. 373; EPFP 3 (1923) 251, pro parte; --Copel., PJS 42 (1930) 581; --Sleumer, Blumea 11 (1961) 73; --Type: BS 696 Foxworthy (PNH, holo, lost; BM, BO, K, L, NY 2 sheets, SING, US, iso), Palawan: Mt Victoria. Mar/Apr-1906.
    • var. palawanense. PALAWAN (Mt Victoria), MINDANAO: Davao (Mt Apo). On exposed ridges and steep cliffs in mossy forest, 1000-1820m. Conservation status: Vulnerable (DAO 2017-11).
    • var. foxworthii (H.F.Copel.) Sleumer, Blumea 11 (1961) 73; --Vaccinium foxworthii H.F.Copel., PJS 42 (1930) 580, pro parte, as ‘foxworthyi’; --Type: BS 649 Foxworthy (PNH, holo, lost; BM, BO, GH, L, NY, US, iso), Palawan: Mt Victoria, Mar/Apr-1906. PALAWAN (Mt Victoria). Once found together with var. palawanense at 1750m. Conservation status: Vulnerable (DAO 2017-11).
  24. Vaccinium perrigidum Elmer, LPB 3 (1911) 1094; --Merr., EPFP 3 (1923) 251; --Copel., PJS 42 (1930) 561; --Sleumer, Blumea 11 (1961) 86; --Type: Elmer 11686 (PNH, holo, lost; A, BM, BO, FI, GH, L, NY, US, iso), Mindanao: Davao, Mt Apo. MINDANAO: Agusan del Norte (Mt Urdaneta), Davao (Mt Apo; Mt McKinley). On rocky bluffs or ridges in the mossy forest, 1525-1675m.
  25. Vaccinium philippinense Warb. in Perkins, Fragm. Fl. Philipp. (1905) 174; --Merr., EPFP 3 (1923) 251; --Copel., PJS 42 (1930) 550; --Sleumer, Blumea 11 (1961) 49; --Type: Cuming 832 (cited by Sleumer as ‘822’) (B, holo, lost; A, BM, K, L, NY, iso), Luzon: Laguna prov., Mayayay (possibly Majayjay, a classical Cuming locality). LUZON: Mountain Province (W part formerly under the old Lepanto subprov.), Nueva Ecija (Mt Umingan = Mingan), Rizal (Mt Batay; Mt Susong Dalaga; Mt Irid; Angilog (possibly Angelo Mtns or Angilo towards Quezon prov.); Mt Lumutan; Montalban; San Isidro, Tanay), Laguna (San Antonio, Paete; Mt Makiling), Quezon-Laguna prov. (Mt San Cristobal and Mahayhay ‘Mayayay’). On forested slopes and ridges, 800-1200m. Fl. & fr. Apr-Dec. Photos
  26. Vaccinium platyphyllum Merr., PJS 12 c (1917) Bot. 294; EPFP 3 (1923) 251; --Copel., PJS 42 (1930) 573; --Sleumer, Blumea 11 (1961) 50; --Type: BS 26583 Ramos & Edaño (PNH, holo, lost; US, iso), Luzon: Aurora prov., Mt Dingalan. Aug/Sep-1916. --Vaccinium ilocanum Merr., PJS 14 (1919) 441; EPFP 3 (1923) 249; --Type: BS 33372 Ramos (PNH, holo, lost; A, BO, K, L, P, US, iso), Luzon: Ilocos Norte prov., Mt Palimlim, Aug-1918. --Vaccinium rizalense Merr., PJS 27 (1925) 43; EPFP 4 (1925) 253; --Type: Loher 14979 (PNH, holo, lost; UC, iso), Luzon: Rizal prov., Balacbac. LUZON: Ilocos Norte (Mt Palimlim), Aurora (Mt Dingalan), Rizal (Guiniuisan; Balacbac). Shrub, small tree or epiphytic in primary lowland to mossy forest, 200-1100m. Fl. Jun-Aug.
  27. Vaccinium pseudocaudatum Sleumer, Blumea 11 (1961) 103; --Type: FB 31527 Franco (NY, holo), Mindanao: Bukidnon prov., Maramag, Sitio Bagumbaan. Near creek along edge of grassland, 2200ft, 17-Dec-1932. MINDANAO: Bukidnon. Near creek along edge of grassland, 670m. Perhaps also growing at higher elevation. Fl. Feb, Dec.
  28. Vaccinium sylvaticum Elmer, LPB 3 (1911) 1095; --Merr., EPFP 3 (1923) 251; --Copel., PJS 42 (1930) 588; --Sleumer, Blumea 11 (1961) 98; --Type: Elmer 11819 (PNH, holo, lost; A, BM, BO, FI, G, GH, K, L, NY, US, iso), Mindanao: Davao, Mt Apo, Todaya, 1250m. --Vaccinium mearnsii Elmer, LPB 3 (1911) 1098; --Merr., EPFP 3 (1923) 250; --Type: Elmer 11251 (PNH, holo, lost; A, BM, BO, FI, G, GH, K, L, NY, US, iso), Mindanao: Davao, Mt Apo, Todaya, 1000m. MINDANAO: Davao (Mt Apo, Todaya). Forested ridges, 1000-1250m. Fl. & fr. Aug-Sep.
  29. Vaccinium tenuipes Merr., PJS 3 c (1908) Bot. 375; EPFP 3 (1923) 251; --Copel., PJS 42 (1930) 571; --Sleumer, Blumea 11 (1961) 77; Blumea 12 (1963) 130; --Type: Clark s.n. (PNH, lecto, lost); Elmer 10108 (US, syn and neolecto; BM, BO, FI, K, L); Merrill 6133 (PNH, syn, lost); FB 11485 Merritt (PNH, syn, lost); Elmer 9819 (PNH, syn, lost). LUZON: Cagayan (Cagua Volcano; Bauan-Mt Tabuan), Ifugao-Mountain Province (South Inomin), Benguet (Mt Pulag; Mt Pulogloco), Rizal (Sierra Madre Range at Angilog, possibly Angelo Mtns or Angilo; Montalban), MINDORO: Mindoro Oriental (Mt Halcon; Ibolo River), NEGROS: Negros Oriental (Cuernos Mtns), PANAY: Capiz (Jamindan). In primary forest, 600-1500m; on Mt Pulag in mossy forest at 2600m. Fl. Feb-Sep. Photos
  30. Vaccinium trichocarpum Sleumer, Blumea 11 (1961) 47; --Type: PNH 10778 Mendoza & Convocar (L, holo). MINDANAO: Agusan del Norte (Mt Hilong-hilong, peak of Mt Malampayan); Davao Oriental (Mt Mayo). In summit vegetation, 1300-1750m. Fl. & fr. Apr.
  31. Vaccinium vidalii Merr. & Rolfe, PJS 3 c (1908) Bot. 374; --Merr., EPFP 3 (1923) 252; --Copel., PJS 42 (1930) 582; --Sleumer, Blumea 11 (1961) 80. LUZON. On exposed ridges in the mossy forest, (300-)850-2000m.
  32. Vaccinium whitfordii Merr., PJS 2 c (1907) Bot. 295; EPFP 3 (1923) 252; --Copel., PJS 42 (1930) 594; --Sleumer, Blumea 11 (1961) 35; --Type: Whitford s.n. (PNH, lecto, lost; K, US), Negros: Negros Occidental prov., Mt Silay. LUZON: Ilocos Norte (Mt Nagapatan), Abra (Mt Paraga), Kalinga (trail to Balbalasang), Mountain Province (W part, formerly part of Lepanto subprov.; Mt Data; Mt Malaya), Benguet (Bucao; Pauai; Baguio), Ifugao (Mt Polis), Nueva Ecija (Mt Umingan = Mt Mingan); Aurora (Mt Alzapan, along Sierra Madre Mtn Range near Quirino prov. border), Rizal (Sierra Madre Mtns near Quezon border), Camarines Sur (Mt Isarog), MINDORO: Mindoro Oriental (Mt Halcon-Ilong Peak), NEGROS: Negros Occidental (Mt Silay; Mt Kanlaon), LEYTE. In forests, open heaths and thickets, 1000-2500m. Fl. & fr. Jan-Dec. Photos
  33. Vaccinium woodianum H.F.Copel., PJS 42 (1930) 591; --Sleumer, Blumea 11 (1961) 82; --Type: Merrill 5506 (PNH, holo, lost; NY flowers partly galled, US, iso), Mindoro: Mindoro Oriental prov., Mt Halcon, 2700m. MINDORO: Mindoro Oriental (Mt Halcon), 2700m. Known only from the type collection.

Literature  

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Argent, G. 2011. Additions to the checklist of Philippine Ericaceae. Philippine Journal of Systematic Biology 5: 97-98

Argent, G. 2013. A new species of Diplycosia: D. coi Argent (Ericaceae) is described in honour of Leonardo Co. Philippine Journal of Science 142: 51-55.

Copeland, HF. 1929. Philippine Ericaceae, I. The species of Rhododendron. Philippine Journal of Science 40: 133-179, +16 pl.

Copeland, HF. 1930. Philippine Ericaceae, II The species of Vaccinium. Philippine Journal of Science 42: 537-607, +7 pl.

Copeland, HF. 1932. Philippine Ericaceae, III: A taxonomic revision, concluded. Philippine Journal of Science 47: 57-118, +8 pl.

DENR Administrative Order. 2017. Updated national list of threatened Philippine plants and their categories. DAO 2017-11

Fritsch, PW & VB Amoroso. 2016. Diplycosia platyphylla (Ericaceae), a new species from Mindanao, Philippines. PhytoKeys 69: 31-38

Fritsch, PW & VB Amoroso. 2017. Diplycosia kitangladensis sp. nov. from Mindanao, the Philippines, and a taxonomic reassessment of D. trinervia. Nat. Hist. Bull. Siam Soc. 62: 21-27

Sleumer, H. 1957. Florae Malesianae Praecursores XV. The genus Gaultheria in Malaysia. Reinwardtia 4: 163-188.

Sleumer, H. 1957. Florae Malesianae Praecursores XIV. A revision of the genus Diplycosia  (Ericaceae). Reinwardtia 4: 119-161.

Sleumer, H. 1960. Rhododendron in Malaysia. Reinwardtia 5: 45-231.

Sleumer, H. 1961. Florae Malesianae Praecursores XXVI. The genus Dimorphanthera F.v.M.,  Nova Guinea, Bot. no. 7: 73-102.

Sleumer, H. 1961. Florae Malesianae Precursores XXVIII. The genus Vaccinium in Malaysia. Blumea 11: 9-112


Sleumer, H. 1963. Florae Malesianae Precursores XXXV. Supplementary notes towards the knowledge of the Ericaceae in Malaysia. Blumea 12: 89-144


Sleumer, H. 1963. Epacridaceae mainly in Asia, Malaysia, and the Pacific. Blumea 12: 145-171

Sleumer, H. 1964. Florae Malesianae Precursores XXXIX. Supplementary notes towards the knowledge of the Ericaceae in Malesia II. Blumea 12: 339-347

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Sleumer, H. 1966. Ericaceae. Flora Malesiana series 1, 6: 469-914

Acknowledgements  

Jasper John Obico and Ravan Schneider kindly contributed some of the photographs on this webpage and George Argent provided some of the identifications.

Pelser, P.B., J.F. Barcelona & D.L. Nickrent (eds.). 2011 onwards. Co's Digital Flora of the Philippines. www.philippineplants.org

Copyright © 2011, Co's Digital Flora of the Philippines

Last updated 28 October 2017