banner small

Elaeocarpaceae

Elaeocarpus L.

Photos
  1. Elaeocarpus affinis Merr., PJS 9 c (1914) Bot. 372; EPFP 3 (1923) 14; --Type: Wenzel 788 (US, iso), Leyte: Leyte prov., Masaganap near Jaro, in forests, c. 600m, May/Jul-1914. LEYTE. Forests, alt. c. 600m.
  2. Elaeocarpus angustifolius Blume, Bijdr. 7 (1827) 120. --Elaeocarpus trichopetalus Merr. & Quisumb., J. Arnold Arb. 32 (1951) 169; --Type: PNH 10099 Sulit (A,   holo; L 2 sheets, iso), Mindanao: Bukidnon prov., Mt Kitanglad, mossy forest, 1850m. --Elaeocarpus wenzelii Merr., PJS 9 c (1914) Bot. 371; EPFP 3 (1923) 20; --Type: Wenzel 365 (US, iso), Leyte: Leyte prov., Dagami, in forests, c. 60m, 13-Jul-1913. NE India to Malesia, Australia and New Caledonia. LEYTE, SAMAR, MINDANAO: Bukidnon (Mt Kitanglad). Lowland to montane forests, alt. 100-1850m.
  3. Elaeocarpus argenteus Merr., Publ. Govt. Lab. Philipp. no. 29 (1905) 26; EPFP 3 (1923) 14; --Type: Elmer 6237 (US, iso), Luzon: Benguet prov., Mt Sto Tomas, May 1904. --Elaeocarpus parvilimbus Merr., J. Arnold Arb. 32 (1951) 168; --Type: BS 42239 Ramos (A, holo; A, SING, iso), Luzon: Rizal prov., Mt Lumutan. Apr-1923. Photos
  4. Elaeocarpus bontocensis Merr., PJS 13 c (1918) Bot. 27; EPFP 3 (1923) 14; --Type: Vanoverbergh 3892, Luzon: Mountain Province, Bauco, in forests, c. 1700m, Jan-1914. LUZON: Mountain Province. Forests, 1700m.
  5. Elaeocarpus burebidensis Elmer., LPB 4 (1911) 1180; --Merr.,  EPFP 3 (1923) 15; --Type: Elmer 11837 (BISH, L, NY, US, iso), Mindanao: Davao, Mt Apo, Todaya approach, Mt Burebid, dry wooded ridge at 4000ft, Sep-1909. MINDANAO: Davao (Mt Apo). Forested ridges, 1200m.
  6. Elaeocarpus calomala (Blanco) Merr., PJS 10 c (1915) Bot. 43; EPFP 3 (1923) 15; --Blanco, Fl. Filip. (1837) 439; --Type: Not extant, ‘Illustrative specimen’: Merr. Sp. Blancoanae 276, leg. M Ramos (US), Luzon: Rizal prov., Antipolo, 105m, 23-Feb-1914. --Elaeocarpus isotrichus (Turcz.) Fern.-Villar, Novis. App. (1880) 31; --A.DC. in Elmer, LPB 2 (1909) 638; --Monocera isotricha Turcz., Bull. Soc. Nat. Mosc. 19, 2 (1846) 494; --Type: Cuming 836 (not seen, cited by Merrill, 1910), Luzon: without definite locality. --Elaeocarpus philippinensis Warb. in Perkins Frag. Fl. Philip. (1904) 101; --A.DC. in Elmer, LPB 2 (1909) 636; --Type: Cuming s.n., (not seen, cited by Merrill, 1910), Luzon: without definite locality. --Elaeocarpus oblongus Fern.-Villar, Novis. App. (1880) 31; --Vidal, Sinopsis Atlas (1883) 17, t. 21, Fig. A; Rev. Pl. Vasc. Filip. (1886) 72; --Perkins, Frag. Fl. Philip. (1904) 100; --Merr., PJS 1 (1906) Suppl. 90; --A.DC. in Elmer, LPB 2 (1909) 636, non Gaertn. --Elaeocarpus bataanensis Merr., PJS 10 c (1915) Bot. 41; --Type: FB 21908 Medina (US, iso), Luzon: Bataan prov., forests back of Limay, on forested slopes, c. 300m, 2-Jul-1914.
  7. Elaeocarpus culminicola Warb., Bot. Jahrb. 16 (1893) 23.
  8. Elaeocarpus cumingii Turcz., Bull. Soc. Nat. Mosc. 19, 2 (1846) 491; --Fern.-Villar, Novis. App. (1880) 31; --Vidal, Phan. Cuming. Philip. (1885) 99; Rev. Pl. Vasc. Filip. (1886) 72; A.DC. in Elmer, LPB 2 (1909) 637; --Merr., EPFP 3 (1923) 15; --Type: Cuming 807. --Elaeocarpus nitidus Turcz., Bull. Soc. Nat. Mosc. 19, 2 (1846) 491; --Fern.-Villar, Novis. App. (1880) 31, non Jack; --Type: Cuming 893 (L, iso), Luzon: Albay prov. --Elaeocarpus oblongus Turcz., Bull. Soc. Nat. Mosc. 31, 1 (1858) 235, non Gaertn. --Elaeocarpus versicolor Elmer, LPB 4 (1911) 1178; --Type: Elmer 12761(BISH, GH, NA, US, iso), Palawan: Palawan prov., Puerto Princesa, Mt Pulgar (=Thumb Peak), Mar 1911. Philippines, Sulawesi. LUZON: Nueva Vizcaya, Bataan, Laguna, Quezon, Camarines, Albay, Sorsogon, PALAWAN, CEBU, SAMAR, BASILAN, MINDANAO. Forests at low altitudes. Photos
  9. Elaeocarpus curranii Merr., PJS 4 c (1909) Bot. 288; EPFP 3 (1923) 16; --Type: FB 4 Ware (US, iso). LUZON: Ilocos Norte (Pauquin), Cagayan, Zambales, Bataan, Pampanga, Quezon, Camarines. Low elevation forests.
  10. Elaeocarpus dinagatensis Merr., J. Arnold Arb. 32 (1951) 162; --Type: BS 35117 Ramos & Pascasio (A, holo?; K, US, iso), Bucas Grande, Jun-1919. DINAGAT; BUCAS GRANDE (type locality). Low altitude. Note by L.L.Co: Both islands have predominantly ultramafic geology. Fr. May-June
  11. Elaeocarpus floribundus Blume, Bijdr. (1825) 120; --Merr., J. Arnold Arb. 32 (1951) 162. India, Myanmar, Thailand, Indochina, Peninsular Malaysia, Java, Borneo, SW Philippines. PALAWAN: (Taytay; Aborlan). Forests at low altitudes.
  12. Elaeocarpus forbesii Merr., PJS 12 c (1917) Bot. 282; EPFP 3 (1923) 16. LUZON: Benguet, MINDORO: Mindoro Oriental (Subaan River headwaters inland from San Teodoro, Coode 5663 with Ridsdale). Forested ravines and ridges, 1100-1500m.
  13. Elaeocarpus foxworthyi Merr., PJS 3 c (1908) Bot. 244; EPFP 3 (1923) 16; --Type: BS 858 Foxworthy. PALAWAN. Forests along streams at low altitudes.
  14. Elaeocarpus fulvus Elmer, LPB 4 (1911) 1179; --Merr., EPFP 3 (1923) 16; --Type: Elmer 10825. MINDANAO: Davao. Primary forests, c. 1100m.
  15. Elaeocarpus gammillii Knuth, Repert. Sp. Nov. 50 (1841) 83; --Type: Gamill 300 (A, iso), Guimaras. Feb-1904. GUIMARAS
  16. Elaeocarpus gigantifolius Elmer, LPB 4 (1911) 1182; --Merr., EPFP 3 (1923) 17; --Type: Elmer 11184 (PNH, holo, lost; K, L, iso), Mindanao: Davao, Mt Apo, Todaya. LEYTE; MINDANAO: Davao (Mt Apo). Note by L.L.Co: Known with certainty from the type only. More material is needed to decide whether the MINDANAO and LEYTE specimens are all part of the same but variable species and whether subspecific status is justified for those from LEYTE (Coode & Weibel, 1994). Steep slopes in forest, 750m. 
  17. Elaeocarpus glaber Blume, Bijdr. (1825) 122; --Merr., J. Arnold Arb. 32 (1951) 164. MINDANAO: Zamboanga prov., Sax River.
  18. Elaeocarpus grandiflorus Sm, Rees Cyclop. 12 (1809) no. 5; --Merr., EPFP 3 (1923) 17. Myanmar, Indochina, Java, Philippines. BUSUANGA. Lowland forests along streams.
  19. Elaeocarpus halconensis Merr., PJS 26 (1925) 470; EPFP 4 (1925) 249; J. Arnold Arb. 32 (1951) 165; --Type: BS 40583 Ramos & Edaño, Mindoro: Mindoro Oriental prov., Mt Halcon.
  20. Elaeocarpus ilocanus Merr., PJS 14 (1919) 418; EPFP 3 (1923) 17; --Type: BS 33271 Ramos. LUZON: Ilocos Norte. Forested slopes, c. 1000m
  21. Elaeocarpus luzonicus Merr., PJS 4 c (1909) Bot. 17; EPFP 3 (1923) 17; --Type: FB 10488 (US, iso), Luzon: Camarines Sur prov., Mt Isarog LUZON: Camarines Sur, Sorsogon, MINDORO, PANAY, CAMIGUIN. Forested slopes, c. 1000m.
  22. Elaeocarpus macranthus Merr., Publ. Govt. Lab. (Philipp.) 35 (1906) 38; EPFP 3 (1923) 17; --Type: Merrill 2298. --Elaeocarpus apoensis Elmer., LPB 4 (1911) 1185; --Merr., EPFP 3 (1923) 14; --Type: Elmer 10524. LUZON: Benguet, Rizal, Laguna, Quezon, Camarines, PANAY, LEYTE, MINDANAO: Davao (Mt Apo). Primary forests, 200-1250m.
  23. Elaeocarpus mandiae Coode, Kew Bull. 56 (2001) 896. MINDORO: Mindoro Oriental (Mt Halcon).
  24. Elaeocarpus merrittii Merr., PJS 2 c (1907) 280; EPFP 3 (1923) 17; --Type: Merrill 5582. --Elaeocarpus urophyllus Merr. & Quisumb., J. Arnold Arb. 32 (1951) 171; --Type: PNH 6832, Sulit, Samar: Eastern Samar prov., Mt Calbiga near Wright. LUZON: Quirino (Mt Alzapan, BS 45617 Ramos & Edano), MINDORO: Mindoro Oriental (Mt Halcon), SAMAR: Eastern Samar (Mt Calbiga). Primary forests, (300-)900-1500m.
  25. Elaeocarpus mindanaensis Merr., PJS 3 c (1908) Bot. 150; EPFP 3 (1923) 18; --Type: Clemens 910. MINDANAO: Lanao, Bukidnon. Forests, c. 750m. Note by L.L.Co: according to Merrill (1923) allied to the Bornean E. gambir Becc.
  26. Elaeocarpus mindorensis Knuth, Repert. Sp. Nov. 50 (1941) 86. --Type: BS 39814 Ramos. MINDORO: Mindoro Occidental (Paluan). Note by L.L.Co: Only known from the type collection.
  27. Elaeocarpus mollis Merr., PJS 9 c (1914) Bot. 373; EPFP 3 (1923) 18; --Type: Wenzel 698 (A, US, iso), Leyte: Leyte prov., Buenavista near Jaro, 20-Apr-1914. LEYTE. Primary forests, 60-500m.
  28. Elaeocarpus monocerus Cav., Icon. 6 (1801) 1, t. 501; DC., Prodr. 1 (1824) 519; --Fern.-Villar,   Novis. App. (1880) 31; --Merr., PJS 1 (1906) Suppl. 90; --A.DC. in Elmer, LPB 2 (1909) 637; --Merr.,   PJS 13 c (1918) Bot. 29; EPFP 3 (1923) 18. --Elaeocarpus megacarpus Elmer, LPB 7 (1915) 2627; --Type: Elmer 14059 (BISH, L, MO, NY, US, iso), Mindanao: Agusan del Norte prov., Cabadbaran, Mt Urdaneta (=Hilong-hilong), Oct-1912. LUZON: Ilocos Sur, Cagayan (Aparri), Isabela, Bataan, Rizal, Laguna (Mt Makiling; San Antonio), Quezon, Camarines, SAMAR: Northern Samar (Palapag), MINDANAO: Zamboanga, Agusan del Norte. Primary forests, ascending to 1000m.
  29. Elaeocarpus multiflorus (Turcz.) Fern.-Villar, Novis. App. Fl. Filip. 31 (1880) 31; --Merr., EPFP 3 (1923) 17; --Type: Cuming 873. --Elaeocarpus leytensis Merr., PJS 8 c (1913) Bot. 383; EPFP 3 (1923) 17; --Type: Wenzel 212. Sulawesi (?) fide Koorders, Philippines. LUZON: Quezon, Camarines, Albay, Sorsogon, MINDORO, SIBUYAN, LEYTE, CAMIGUIN, MINDANAO. Low and medium elevation primary forests.
  30. Elaeocarpus nervosus Elmer, LPB 4 (1911) 1176; --Merr.,  EPFP 3 (1923) 18; --Type: Elmer 11566. LUZON: Mountain Province, Benguet, MINDANAO: Davao. Forests, 1350-1600m. According to Merrill (1923), closely allied to E. cumingii.
  31. Elaeocarpus octopetalus Merr., PJS 3 c (1908) Bot. 149; EPFP 3 (1923) 18; J. Arnold Arb. 32 (1951) 167, emend., “petalis sepalisque 4, non 8.”; --Type: Clemens 1148. MINDANAO: Lanao del Sur (Camp Keithley). Forests, c. 750m.
  32. Elaeocarpus palimlimensis Knuth, Repert. Sp. Nov. 49 (1940) 68; --Merr., J. Arnold Arb. 32 (1951) 167; --Type: BS 33377 Ramos (A, US, iso), Luzon: Ilocos Norte prov., Mt Palimlim,  Aug-1918. LUZON: Ilocos Norte. Note by L.L.Co: Only known from the type collection.
  33. Elaeocarpus pedunculatus Wall., List (1832) no. 2678, nomen nudum,  ex Mast. in JD Hook., Fl. Brit. Ind. (1874) 408, descr. --Elaeocarpus candollei Turcz., Bull. Soc. Nat. Mosc. 192 ; Elmer, LPB 4 (1911) 1185; --Merr., EPFP 3 (1923) 15; --Type: Elmer 12974 (L 2 sheets, NY 2 sheets, US, iso), Palawan: Puerto Princesa, Mt   Pulgar (=Thumb Peak), Apr-1911. --Elaeocarpus nitidulus Knuth, Fedde Repert. 49 (1940) 67; --Type: BS 41234 Ramos (A, iso), Busuanga Island, Sep-1922.  Sumatra, Peninsular Malaysia, Banka, Billiton, Borneo, C Philippines. BUSUANGA, PALAWAN, PANAY, CEBU. Forests, 1000-1300m.
  34. Elaeocarpus samari Weibel, Candollea 35 (1980) 524. --Type: PNH 6398 Sulit (L, holo; A, iso), Samar: Eastern Samar prov., Wright, Mt Calbiga. Apr/May-1948. SAMAR.
  35. Elaeocarpus sphaericus (Gaertn.) K Schum. in Engl. & Prantl, Nat. Pflanzenfam. 3, 6 (1890) 5; --Corner,  Gard. Bull. Sing. 10 (1939) 326. --Elaeocarpus subglobosus Merr., PJS 4 c (1909) Bot. 290 ; EPFP 3 (1923) 19. --Elaeocarpus baclayanensis Elmer, LPB 4 (1911) 1188. --Elaeocarpus dolichopetalus Merr., PJS 9 c (1914) Bot. 372; EPFP 3 (1923) 16; --Type: Wenzel 897 (A, US, iso), Leyte: Leyte prov., Buenavista near Jaro, in forests, c. 500m, 13-Jun-1914. --Elaeocarpus ramiflorus Merr., PJS 10 c (1915) Bot. 43; EPFP 3 (1923) 19; --Type: BS 13115 Ramos (US, iso), Luzon: Laguna prov., San Antonio, trail between San Antonio and Piape, in forests, Jun-1912. Hainan. Philippines. LUZON: Mountain Province, Benguet, Laguna, Camarines Sur (Mt Isarog), Sorsogon, PANAY: Capiz (Libacao, BS 35438 Martelino & Edaño), LEYTE, MINDANAO: Davao (Mt Apo). Primary forests, 400-1600m.
  36. Elaeocarpus submonoceras Miq., Fl. Ind. Bat., suppl. 1 (1860) 407.
  37. Elaeocarpus surigaensis Merr., PJS 13 c (1918) Bot. 28; EPFP 3 (1923) 19; --Type: FB 26000 Ponce (US, iso), Mindanao: Surigao del Norte prov., near Mabini, in level rich soil at the edge of the forest, c. 50m, 11-May-1916. MINDANAO: Surigao del Norte. Lowland forests.
  38. Elaeocarpus venosus C.B.Rob., Bull. Torr. Bot. Club 35 (1908) 71, 75; --Merr.,  EPFP 3 (1923) 20; --Type: RS Williams 2002. LUZON: Abra, Mountain Province, Benguet. Mossy forests on the higher mountains, 1800-2500m.
  39. Elaeocarpus verruculosus A.DC. in Elmer, LPB 2 (1909) 635; EPFP 3 (1923) 20; --Syntypes: Elmer 8645 (Elmer 8734). LUZON: Benguet. Thickets or forests, c. 1500m.
  40. Elaeocarpus verticillatus Elmer., LPB 4 (1911) 1181; --Merr., EPFP 3 (1923) 20; --Type: Elmer 12104 (PNH, holo, lost; L 2 sheets, MO, NY, US, iso), Sibuyan: Mt Giting-giting, Magallanes (=Magdiwang), along wooded streams at 750ft, Mar-1910; --Syntypes: Elmer 12427 (L, MO, NY, US), Sibuyan: Mt Giting-giting, Magallanes (=Magdiwang), Pauala River, 2000ft, May-1910. Photos

Doubtful:

  1. Elaeocarpus stipularis Blume, Bijdr. (1825) 120. --Elaeocarpus fissistipulus Miq., Fl. Ind. Bat. 13 (1858-59) 210; --Merr.,  EPFP 3 (1923) 16. Note by L.L.Co: Based on a Llanos specimen in de Candolle’s herbarium not seen by Merrill,

Sloanea L.

Photos
  1. Sloanea javanica (Miq.) Szyszyl. ex K Schum. in Engl. & Prantl, Nat. Pflanzenfam. 36 (1890) 5; --Merr., EPFP 3  (1923) 21. Peninsular Malaysia, Java, Philippines, ?Sulawesi. LUZON: Cagayan, Isabela, Laguna, MINDORO, MINDANAO: Lanao. Low elevation primary forests. Photos
  2. Sloanea sigun (Blume) K. Schum. in Engl. & Prantl, Nat. Pflanzenfam. 3, 6 (1890) 5; --Coode, Kew Bull. 38 (1983) 389. India (Assam), Thailand, Sumatra, Peninsular Malaysia, Java, Borneo (Sabah) and the Philippines (widespread). Lowland primary forests. 

Literature

Coode, MJE. 1983. A conspectus of Sloanea (Elaeocarpaceae) of the Old World. Kew Bulletin 38: 347-427.

Coode, MJE & R Weibel. 1994. Elaeocarpus for Flora Malesiana: notes, new taxa and combinations in sect. Elaeocarpus: 1. Kew Bulletin 49: 235-259.

Coode, MJE. 2001. Elaeocarpus for Flora Malesiana: the E. stipularis complex, E. nitidus group & E. barbulatus. Kew Bulletin 56: 513-565.

Coode, MJE. 2001. Elaeocarpus for Flora Malesiana: the verticellatae subgroup of the Monocera group and a new Philippine species. Kew Bulletin 56: 885-901.

Coode, MJE. 2007. Elaeocarpus for Flora Malesiana: E. forbesii, a Philippine species with unusual features, and some possible 'associates'. Kew Bulletin 62: 321-328.

Coode, MJE. 2010. Elaeocarpus for Flora Malesiana: new taxa and understanding in the Ganitrus group. Kew Bulletin 65: 355-399.

Copyright © 2011, Co's Digital Flora of the Philippines

Last updated May 4, 2014