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Elaeocarpaceae

Elaeocarpus L.

Edited by Mark Coode, James V. LaFrankie & Pieter B. Pelser, 22 August 2016

Photos
  1. Elaeocarpus angustifolius Blume, Bijdr. 7 (1827) 120; --Type from Java. --Elaeocarpus wenzelii Merr., PJS 9 c (1914) Bot. 371; EPFP 3 (1923) 20; --Type: Wenzel 365 (PNH, holo, lost?; G, L, US, iso), Leyte: Leyte prov., Dagami, in forests, c. 60m, 13-Jul-1913. --Elaeocarpus sphaericus (Gaertn.) K Schum. in Engl. & Prantl, Nat. Pflanzenfam. 3, 6 (1890) 5, nom. illeg.; --Corner,  Gard. Bull. Sing. 10 (1939) 326. NE India to Malesia, Australia and New Caledonia, Philippines. LEYTE, LUZON, PALAWAN, PANAY, SAMAR, MINDANAO: Bukidnon (Mt Kitanglad). Lowland to montane forests, 100-1850m.
  2. Elaeocarpus argenteus Merr., Publ. Govt. Lab. Philipp. no. 29 (1905) 26; EPFP 3 (1923) 14; --Type: Elmer 6237 (US, iso), Luzon: Benguet prov., Mt Sto Tomas, May-1904. --Elaeocarpus parvilimbus Merr., J. Arnold Arb. 32 (1951) 168; --Type: BS 42239 Ramos (A, holo; A, SING, iso), Luzon: Rizal prov., Mt Lumutan. Apr-1923. --Elaeocarpus affinis Merr., PJS 9 c (1914) Bot. 372; EPFP 3 (1923) 14; --Type: Wenzel 788 (US, iso), Leyte: Leyte prov., Masaganap near Jaro, in forests, c. 600m, May/Jul-1914. Photos
  3. Elaeocarpus brunnescens R.Knuth, Repert. Spec. Nov. Regni Veg. 44 (1938) 127. PALAWAN.
  4. Elaeocarpus burebidensis Elmer., LPB 4 (1911) 1180; --Merr.,  EPFP 3 (1923) 15; --Type: Elmer 11837 (BISH, E, G, K, L, NY, US, iso), Mindanao: Davao, Mt Apo, Todaya approach, Mt Burebid, dry wooded ridge at 4000ft, Sep-1909. MINDANAO: Davao (Mt Apo). Forested ridges, 1200m. Known only from the type (fide Mark Coode, 5-Aug-16).
  5. Elaeocarpus calomala (Blanco) Merr., PJS 10 c (1915) Bot. 43; EPFP 3 (1923) 15; --Blanco, Fl. Filip. (1837) 439; --Type: Not extant, ‘Illustrative specimen’: Merr. Sp. Blancoanae 276, leg. M Ramos (US), Luzon: Rizal prov., Antipolo, 105m, 23-Feb-1914. --Elaeocarpus isotrichus (Turcz.) Fern.-Villar, Novis. App. (1880) 31; --DC. in Elmer, LPB 2 (1909) 638; --Monocera isotricha Turcz., Bull. Soc. Nat. Mosc. 19, 2 (1846) 494; --Type: Cuming 836 (not seen, cited by Merrill, 1910), Luzon: without definite locality. --Elaeocarpus philippinensis Warb. in Perkins Frag. Fl. Philip. (1904) 101; --DC. in Elmer, LPB 2 (1909) 636; --Type: Cuming s.n., (not seen, cited by Merrill, 1910), Luzon: without definite locality. --Elaeocarpus oblongus Fern.-Villar, Novis. App. (1880) 31; --Vidal, Sinopsis Atlas (1883) 17, t. 21, Fig. A; Rev. Pl. Vasc. Filip. (1886) 72; --Perkins, Frag. Fl. Philip. (1904) 100; --Merr., PJS 1 (1906) Suppl. 90; --DC. in Elmer, LPB 2 (1909) 636, non Gaertn. --Elaeocarpus bataanensis Merr., PJS 10 c (1915) Bot. 41; --Type: FB 21908 Medina (US, iso), Luzon: Bataan prov., forests back of Limay, on forested slopes, c. 300m, 2-Jul-1914.
  6. Elaeocarpus culminicola Warb., Bot. Jahrb. 16 (1893) 23. --Elaeocarpus pendulus Merr., Publ. Govt. Lab. Philipp. no. 29 (1905) 27; EPFP 3 (1923) 18; --Type: Elmer 5801 (NY, US), Luzon: Benguet prov., Mt Sto Tomas, Mar-1904. --Elaeocarpus microphyllus Elmer., LPB 4 (1911) 1189 ; --Merr., EPFP 3 (1923) 17; --Type: Elmer 11860 (BISH, BM, E, G, K, L, NY, U, US), Mindanao: Davao, Mt Apo, Todaya, “in rich moist humus-covered soil of dense forests at 4500ft of Mt Calelan”, Sep-1909. --Elaeocarpus maquilingensis Elmer, LPB 8 (1919) 3080; --Type: Elmer 18039 (A, US, MO), Luzon: Mt Makiling, Jul-1917. LUZON: Mountain Province, Benguet, Nueva Vizcaya, Pangasinan, Zambales, Bataan, Laguna, MINDORO, LEYTE, MINDANAO: Bukidnon, Davao (Mt Apo). Montane and ridge-top forests, 1000-2200m.
  7. Elaeocarpus cumingii Turcz., Bull. Soc. Nat. Mosc. 19, 2 (1846) 491; --Fern.-Villar, Novis. App. (1880) 31; --Vidal, Phan. Cuming. Philip. (1885) 99; Rev. Pl. Vasc. Filip. (1886) 72; DC. in Elmer, LPB 2 (1909) 637; --Merr., EPFP 3 (1923) 15; --Type: Cuming 807 (BM, E, G, K, iso). --Elaeocarpus nitidus Turcz., Bull. Soc. Nat. Mosc. 19, 2 (1846) 491; --Fern.-Villar, Novis. App. (1880) 31, non Jack; --Type: Cuming 893 (BM, E, G, K, L, iso), Luzon: Albay prov. --Elaeocarpus oblongus Turcz., Bull. Soc. Nat. Mosc. 31, 1 (1858) 235, non Gaertn. --Elaeocarpus versicolor Elmer, LPB 4 (1911) 1178; --Type: Elmer 12761 (BISH, BM, E, G, GH, K, NA, US, iso), Palawan: Palawan prov., Puerto Princesa, Mt Pulgar (=Thumb Peak), Mar-1911. Philippines, Sulawesi. LUZON: Nueva Vizcaya, Bataan, Laguna, Quezon, Camarines, Albay, Sorsogon, PALAWAN, CEBU, SAMAR, BASILAN, MINDANAO. Forests at low elevation. Photos
  8. Elaeocarpus curranii Merr., PJS 4 c (1909) Bot. 288; EPFP 3 (1923) 16; --Type: FB 4 Ware (BM, K, US, iso). LUZON: Ilocos Norte (Pauquin), Cagayan, Zambales, Bataan, Pampanga, Quezon, Camarines. Low elevation forests.
  9. Elaeocarpus dinagatensis Merr., J. Arnold Arb. 32 (1951) 162; --Type: BS 35117 Ramos & Pascasio (A, holo?; K, US, iso), Bucas Grande, Jun-1919. DINAGAT, BUCAS GRANDE. Low elevation. Note by L.L.Co: Both islands have predominantly ultramafic geology. Fr. May-June
  10. Elaeocarpus floribundus Blume, Bijdr. (1825) 120; --Merr., J. Arnold Arb. 32 (1951) 162. India, Myanmar, Thailand, Indochina, Peninsular Malaysia, Java, Borneo, SW Philippines. PALAWAN: (Taytay; Aborlan). Forests at low elevation.
  11. Elaeocarpus forbesii Merr., PJS 12 c (1917) Bot. 282; EPFP 3 (1923) 16; --Type: Leano FB 24726 (PNH, holo, lost; K, iso), Luzon: Benguet Prov., Baguio, 27-Sep-1915. Endemic to Philippines. LUZON: Benguet, MINDORO: Mindoro Oriental (Subaan River headwaters inland from San Teodoro, Coode 5663 with Ridsdale). Forested ravines and ridges, 1100-1500m. Photos
  12. Elaeocarpus foxworthyi Merr., PJS 3 c (1908) Bot. 244; EPFP 3 (1923) 16; --Type: BS 858 Foxworthy. PALAWAN. Forests along streams at low elevation.
  13. Elaeocarpus fulvus Elmer, LPB 4 (1911) 1179; --Merr., EPFP 3 (1923) 16; --Type: Elmer 10825 (PNH, holo, lost; BISH, BP, E, G, K, L, U, iso). Endemic to Philippines. MINDANAO: Davao. Primary forests, c. 1100m. Known only from the type (fide Mark Coode, 5-Aug-16). Conservation status: DD (Coode, 2010).
  14. Elaeocarpus gammillii Knuth, Repert. Sp. Nov. 50 (1841) 83; --Type: Gamill 300 (A, K, iso), Guimaras. Feb-1904. GUIMARAS
  15. Elaeocarpus gigantifolius Elmer, LPB 4 (1911) 1182; --Merr., EPFP 3 (1923) 17; --Type: Elmer 11184 (PNH, holo, lost; K, L, iso), Mindanao: Davao, Mt Apo, Todaya. LEYTE; MINDANAO: Davao (Mt Apo). Note by L.L.Co: Known with certainty from the type only. More material is needed to decide whether the MINDANAO and LEYTE specimens are all part of the same but variable species and whether subspecific status is justified for those from LEYTE (Coode & Weibel, 1994). Steep slopes in forest, 750m.
  16. Elaeocarpus gitingensis Elmer., LPB 4 (1911) 1177; --Merr., EPFP 3 (1923) 17; --Syntypes: Elmer 12050a (BISH, BM, E, K, L, MO, NY, US), Sibuyan: Magallanes (=Magdiwang), Mt Giting-giting, hill forests at 500ft, Mar-1910; Elmer 12470 (A, BISH, GH, MO, US), Sibuyan: Magallanes (=Magdiwang), Mt Giting-giting, May-1910. SIBUYAN (Mt Giting-giting). Lowland forest, c. 225m.
  17. Elaeocarpus grandiflorus Sm, Rees Cyclop. 12 (1809) no. 5; --Merr., EPFP 3 (1923) 17. Myanmar, Indochina, Java, Philippines. BUSUANGA. Lowland forests along streams.
  18. Elaeocarpus halconensis Merr., PJS 26 (1925) 470; EPFP 4 (1925) 249; J. Arnold Arb. 32 (1951) 165; --Type: BS 40583 Ramos & Edaño (PNH, holo, lost; A, K, L, iso), Mindoro: Mindoro Oriental prov., Mt Halcon. Endemic to Philippines.
  19. Elaeocarpus ilocanus Merr., PJS 14 (1919) 418; EPFP 3 (1923) 17; --Type: BS 33271 Ramos (K, L, iso). LUZON: Ilocos Norte. Forested slopes, c. 1000m. Known only from the type (fide Mark Coode, 5-Aug-16).
  20. Elaeocarpus lagunensis Knuth, Repert. Sp. Nov. 50 (1941) 85; --Type: BS 15126 Ramos (B, holo, lost; BM, G, K, L, iso), Luzon: Laguna Prov., San Antonio, Jun-1912. Endemic to Philippines. Photos
  21. Elaeocarpus macranthus Merr., Publ. Govt. Lab. (Philipp.) 35 (1906) 38; EPFP 3 (1923) 17; --Type: Merrill 2298 (K, iso). --Elaeocarpus apoensis Elmer., LPB 4 (1911) 1185; --Merr., EPFP 3 (1923) 14; --Type: Elmer 10524 (BO, E, G, K, L, iso). LUZON: Benguet, Rizal, Laguna, Quezon, Camarines, PANAY, LEYTE, MINDANAO: Davao (Mt Apo). Primary forests, 200-1250m. Photos
  22. Elaeocarpus mandiae Coode, Kew Bull. 56 (2001) 896; --Type: Argent 29069 (E, holo; K, PNH, iso), Mindoro: Mt. Halcon, 14-Mar-1997. Endemic to Philippines. MINDORO: Mindoro Oriental.
  23. Elaeocarpus merrittii Merr., PJS 2 c (1907) 280; EPFP 3 (1923) 17; --Elaeocarpus bellus R.Knuth, Repert. Spec. Nov. Regni Veg. 49 (1940) 67; --Type: Merrill 5582 (G, K, iso). LUZON: Quirino (Mt Alzapan, BS 45617 Ramos & Edano), MINDORO: Mindoro Oriental (Mt Halcon). Primary forests, 900-1500m.
  24. Elaeocarpus mindoroensis Knuth, Repert. Sp. Nov. 50 (1941) 86. --Type: BS 39814 Ramos (K, L, iso). MINDORO: Mindoro Occidental (Paluan), PANAY.
  25. Elaeocarpus mollis Merr., PJS 9 c (1914) Bot. 373; EPFP 3 (1923) 18; --Type: Wenzel 698 (A, BM, G, US, iso), Leyte: Leyte prov., Buenavista near Jaro, 20-Apr-1914. BILIRAN, LEYTE, SAMAR. Primary forests, 60-500m.
  26. Elaeocarpus monocera Cav., Icon. 6 (1801) 1, t. 501; DC., Prodr. 1 (1824) 519; --Fern.-Villar,   Novis. App. (1880) 31; --Merr., PJS 1 (1906) Suppl. 90; --DC. in Elmer, LPB 2 (1909) 637; --Merr.,   PJS 13 c (1918) Bot. 29; EPFP 3 (1923) 18. --Elaeocarpus megacarpus Elmer, LPB 7 (1915) 2627; --Type: Elmer 14059 (BISH, E, G, K, L, MO, NY, US, iso), Mindanao: Agusan del Norte prov., Cabadbaran, Mt Urdaneta, Oct-1912. LUZON: Ilocos Sur, Cagayan (Aparri), Isabela, Bataan, Rizal, Laguna (Mt Makiling; San Antonio), Quezon, Camarines, SAMAR: Northern Samar (Palapag), MINDANAO: Zamboanga, Agusan del Norte. Primary forests, ascending to 1000m.
  27. Elaeocarpus multiflorus (Turcz.) Fern.-Villar, Novis. App. Fl. Filip. 31 (1880) 31; --Merr., EPFP 3 (1923) 17; --Type: Cuming 873 (E, G, K, L, iso). --Elaeocarpus leytensis Merr., PJS 8 c (1913) Bot. 383; EPFP 3 (1923) 17; --Type: Wenzel 212 (E, G, iso). Sulawesi (?) fide Koorders, Philippines. LUZON: Quezon, Camarines, Albay, Sorsogon, MINDORO, SIBUYAN, LEYTE, CAMIGUIN, MINDANAO. Low and medium elevation primary forests.
  28. Elaeocarpus murudensis Merr., Sarawak Mus. J. 3 (1928) 527. Borneo, Philippines. PALAWAN.
  29. Elaeocarpus nervosus Elmer, LPB 4 (1911) 1176; --Merr.,  EPFP 3 (1923) 18; --Type: Elmer 11566 (BM, E, K, L, iso). LUZON: Mountain Province, Benguet, MINDANAO: Davao. Forests, 1350-1600m.
  30. Elaeocarpus nitidus Jack, Malayan Misc. 1 (1820) 41.
  31. Elaeocarpus obtusus Blume, Catalogus (1823) 78.
  32. Elaeocarpus octopetalus Merr., PJS 3 c (1908) Bot. 149; EPFP 3 (1923) 18; J. Arnold Arb. 32 (1951) 167, emend., “petalis sepalisque 4, non 8.”; --Type: Clemens 1148. Moluccas, Sulawesi, Philippines. MINDANAO: Lanao del Sur (Camp Keithley). Forests, c. 750m. Photos
  33. Elaeocarpus pedunculatus Wall., List (1832) no. 2678, nomen nudum,  ex Mast. in JD Hook., Fl. Brit. Ind. (1874) 408, descr. --Elaeocarpus candollei Turcz., Bull. Soc. Nat. Mosc. 192 ; Elmer, LPB 4 (1911) 1185; --Merr., EPFP 3 (1923) 15; --Type: Elmer 12974 (L 2 sheets, NY 2 sheets, E, G, K, US, iso), Palawan: Puerto Princesa, Mt   Pulgar (=Thumb Peak), Apr-1911. --Elaeocarpus nitidulus Knuth, Fedde Repert. 49 (1940) 67; --Type: BS 41234 Ramos (A, K, L, iso), Busuanga Island, Sep-1922.  Sumatra, Peninsular Malaysia, Banka, Billiton, Borneo, C Philippines. BUSUANGA, PALAWAN, PANAY, CEBU. Forests, 1000-1300m. Photos
  34. Elaeocarpus ramiflorus Merr., PJS 10 c (1915) Bot. 43; EPFP 3 (1923) 19; --Type: BS 15115 Ramos (PNH, holo, lost; G, K, L, US, iso), Luzon: Laguna prov., San Antonio, trail between San Antonio and Piape, in forests, Jun-1912. Endemic to Philippines. LUZON: Benguet, Laguna, Mountain Province, Sorsogon, MINDORO?, PANAY?. Conservation status: DD (Coode, 2010).
  35. Elaeocarpus samari Weibel, Candollea 35 (1980) 524. --Type: PNH 6398 Sulit (L, holo; A, K, iso), Samar: Eastern Samar prov., Wright, Mt Calbiga. Apr/May-1948. SAMAR. Known only from the type (fide Mark Coode, 5-Aug-16).
  36. Elaeocarpus stipularis Blume, Bijdr. Fl. Ned. Ind. 3 (1825) 121. --Elaeocarpus fissistipulus Miq., Fl. Ind. Bat. 13 (1858-59) 210; --Merr.,  EPFP 3 (1923) 16.
  37. Elaeocarpus submonoceras Miq., Fl. Ind. Bat., suppl. 1 (1860) 407.
  38. Elaeocarpus surigaensis Merr., PJS 13 c (1918) Bot. 28; EPFP 3 (1923) 19; --Type: FB 26000 Ponce (PNH, holo, lost; K, US, iso), Mindanao: Surigao del Norte prov., near Mabini, in level rich soil at the edge of the forest, c. 50m, 11-May-1916. Endemic to Philippines. MINDANAO: Surigao del Norte. Lowland forests. Known only from the type (fide Mark Coode, 5-Aug-16).
  39. Elaeocarpus trichopetalus Merr. & Quisumb., J. Arnold Arb. 32 (1951) 169; --Type: PNH 10099 Sulit (A, holo; G, K, L 2 sheets, iso), Mindanao: Bukidnon prov., Mt Kitanglad, mossy forest, 1850m. Moluccas, Sulawesi, Philippines. MINDANAO: Bukiknon. Close to Elaeocarpus angustifolius (Mark Coode, 5-Aug-16).
  40. Elaeocarpus triflorus Merr., J. Arnold Arb. 32 (1951) 170; --Type: Escritor BS 20796 (A, holo; K, iso), Luzon: Quezon prov., Guinayangan, Mar/Apr-1913. Endemic to Philippines. LUZON: Aurora (Mt Dingalan), Quezon (Guinayangan).
  41. Elaeocarpus urophyllus Merr. & Quisumb., J. Arnold Arb. 32 (1951) 171; --Type: PNH 6832, Sulit, Samar: Eastern Samar prov., Mt Calbiga near Wright. SAMAR.
  42. Elaeocarpus venosus C.B.Rob., Bull. Torr. Bot. Club 35 (1908) 71, 75; --Merr.,  EPFP 3 (1923) 20; --Type: RS Williams 2002 (BO, K, NY, iso). LUZON: Abra, Mountain Province, Benguet. Mossy forests on the higher mountains, 1800-2500m.
  43. Elaeocarpus verruculosus DC. in Elmer, LPB 2 (1909) 635; EPFP 3 (1923) 20; --Syntypes: Elmer 8645 (BM, E, G, K, L, isosyn), Elmer 8734 (G, isosyn). LUZON: Benguet. Thickets or forests, c. 1500m.
  44. Elaeocarpus verticillatus Elmer., LPB 4 (1911) 1181; --Merr., EPFP 3 (1923) 20; --Type: Elmer 12104 (PNH, holo, lost; K, lecto; E, G, K, L 2 sheets, MO, NY, U, US, iso), Sibuyan: Mt Giting-giting, Magallanes (=Magdiwang), along wooded streams at 750ft, Mar-1910; --Syntypes: Elmer 12427 (PNH, holo, lost; G, K, L, MO, NY, U, US), Sibuyan: Mt Giting-giting, Magallanes (=Magdiwang), Pauala River, 2000ft, May-1910. Endemic to Philippines. Photos

Doubtfully present:

  1. Elaeocarpus glaber Blume, Bijdr. (1825) 122; --Merr., J. Arnold Arb. 32 (1951) 164. MINDANAO: Zamboanga prov., Sax River. Note by Mark Coode (5-Aug-16): Williams 2134 seems better interpreted as a large-leaved Elaeocarpus cumingii.

Insufficiently known:

  1. Elaeocarpus baclayanensis Elmer, LPB 4 (1911) 1188; --Coode, Kew Bull. 65 (2010) 371; --Type: Elmer 10734 (G, lecto), Mindanao: Todaya (Mt Apo), May-1909. Insufficiently known within the Elaeocarpus angustifolius complex.
  2. Elaeocarpus bontocensis Merr., PJS 13 c (1918) Bot. 27; EPFP 3 (1923) 14; --Type: Vanoverbergh 3892, Luzon: Mountain Province, Bauco, in forests, c. 1700m, Jan-1914. LUZON: Mountain Province. Forests, 1700m.
  3. Elaeocarpus dolichopetalus Merr., PJS 9 c (1914) Bot. 372; EPFP 3 (1923) 16; --Type: Wenzel 897 (PNH, holo, lost; A, G, US, iso), Leyte: Leyte prov., Buenavista near Jaro, in forests, c. 500m, 13-Jun-1914. Insufficiently known within the Elaeocarpus angustifolius complex.
  4. Elaeocarpus luzonicus Merr., PJS 4 c (1909) Bot. 17; EPFP 3 (1923) 17; --Type: FB 10488 (US, iso), Luzon: Camarines Sur prov., Mt Isarog LUZON: Camarines Sur, Sorsogon, MINDORO, PANAY, CAMIGUIN. Forested slopes, c. 1000m.
  5. Elaeocarpus subglobosus Merr., PJS 4 c (1909) Bot. 290; EPFP 3 (1923) 19; --Syntypes: Williams 1036 (PNH, syn, lost; K, isosyn), Alvarez FB 18317 (PNH, syn, lost), Luzon: Benguet Prov., Baguio, 15-Oct-1904. Insufficiently known within the Elaeocarpus angustifolius complex.

Sloanea L.

Edited by Pieter B. Pelser & James V. LaFrankie, 22 August 2016

Photos
  1. Sloanea javanica (Miq.) Szyszyl. ex K Schum. in Engl. & Prantl, Nat. Pflanzenfam. 36 (1890) 5; --Merr., EPFP 3  (1923) 21. Peninsular Malaysia, Java, Philippines, ?Sulawesi. LUZON: Cagayan, Isabela, Laguna, MINDORO, MINDANAO: Lanao. Low elevation primary forests. Photos

Literature

Coode, MJE. 1983. A conspectus of Sloanea (Elaeocarpaceae) of the Old World. Kew Bulletin 38: 347-427.

Coode, MJE & R Weibel. 1994. Elaeocarpus for Flora Malesiana: notes, new taxa and combinations in sect. Elaeocarpus: 1. Kew Bulletin 49: 235-259.

Coode, MJE. 2001. Elaeocarpus for Flora Malesiana: the E. stipularis complex, E. nitidus group & E. barbulatus. Kew Bulletin 56: 513-565.

Coode, MJE. 2001. Elaeocarpus for Flora Malesiana: the verticellatae subgroup of the Monocera group and a new Philippine species. Kew Bulletin 56: 885-901.

Coode, MJE. 2007. Elaeocarpus for Flora Malesiana: E. forbesii, a Philippine species with unusual features, and some possible 'associates'. Kew Bulletin 62: 321-328.

Coode, MJE. 2010. Elaeocarpus for Flora Malesiana: new taxa and understanding in the Ganitrus group. Kew Bulletin 65: 355-399.

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Last updated 1 November 2016