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Elaeocarpaceae

Elaeocarpus L.

Photos
  1. Elaeocarpus affinis Merr., PJS 9 c (1914) Bot. 372; EPFP 3 (1923) 14; --Type: Wenzel 788 (US*, iso), Leyte: Leyte prov., Masaganap near Jaro, in forests, altitude c. 600m, May-Jul 1914. LEYTE. Forests, alt. c. 600m.
  2. Elaeocarpus angustifolius Blume, Bijdr. 7 (1827) 120. --Elaeocarpus trichopetalus Merr. & Quisumb., J. Arnold Arb. 32 (1951) 169; --Type: PNH 10099 Sulit (A*,   holo; L 2 sheets*, iso), Mindanao: Bukidnon prov., Mt Kitanglad, mossy forest, altitude 1850m. --Elaeocarpus wenzelii Merr., PJS 9 c (1914) Bot. 371; EPFP 3 (1923) 20; --Type: Wenzel 365 (US*, iso), Leyte: Leyte prov., Dagami, in forests, altitude c. 60m, 13 Jul 1913. NE India to Malesia, Australia and New Caledonia. LEYTE, SAMAR, MINDANAO: Bukidnon (Mt Kitanglad). Lowland to montane forests, alt. 100-1850m.
  3. Elaeocarpus argenteus Merr., Publ. Govt. Lab. Philipp. no. 29 (1905) 26; EPFP 3 (1923) 14; --Type: Elmer 6237 (US*, iso), Luzon: Benguet prov., Mt Sto Tomas, May 1904. --Elaeocarpus parvilimbus Merr., J. Arnold Arb. 32 (1951) 168; --Type: BS 42239 Ramos (A*, holo; iso in A*, SING*!), Luzon: Rizal prov., Mt Lumutan. April 1923. Photos
  4. Elaeocarpus bontocensis Merr., PJS 13 c (1918) Bot. 27; EPFP 3 (1923) 14; --Type: Vanoverbergh 3892 (not seen), Luzon: Mountain Province, Bauco, in forests, altitude c. 1700m, Jan 1914. LUZON: Mountain Province. Forests, altitude 1700m.
  5. Elaeocarpus burebidensis Elmer., LPB 4 (1911) 1180; --Merr.,  EPFP 3 (1923) 15; --Type: Elmer 11837 (BISH*, L*, NY*, US*, iso), Mindanao: Davao, Mt Apo, Todaya approach, Mt Burebid, dry wooded ridge at 4000ft, Sept 1909. MINDANAO: Davao (Mt Apo). Forested ridges, altitude 1200m.
  6. Elaeocarpus calomala (Blanco) Merr., PJS 10 c (1915) Bot. 43; EPFP 3 (1923) 15; --Blanco, Fl. Filip. (1837) 439; --Type: Not extant, ‘Illustrative specimen’: Merr. Sp. Blancoanae 276, leg. M Ramos (US*), Luzon: Rizal prov., Antipolo, altitude 105m, 23 Feb 1914. --Elaeocarpus isotrichus (Turcz.) Fern.-Villar, Novis. App. (1880) 31; --A DC in Elmer, LPB 2 (1909) 638; --Monocera isotricha Turcz., Bull. Soc. Nat. Mosc. 19, 2 (1846) 494; --Type: Cuming 836 (not seen, cited by Merrill, 1910), Luzon: without definite locality. --Elaeocarpus philippinensis Warb. in Perkins Frag. Fl. Philip. (1904) 101; --A DC in Elmer, LPB 2 (1909) 636; --Type: Cuming s.n., (not seen, cited by Merrill, 1910), Luzon: without definite locality. --Elaeocarpus oblongus Fern.-Villar, Novis. App. (1880) 31; --Vidal, Sinopsis Atlas (1883) 17, t. 21, Fig. A; Rev. Pl. Vasc. Filip. (1886) 72; --Perkins, Frag. Fl. Philip. (1904) 100; --Merr., PJS 1 (1906) Suppl. 90; --A DC in Elmer, LPB 2 (1909) 636, non Gaertn. --Elaeocarpus bataanensis Merr., PJS 10 c (1915) Bot. 41; --Type: FB 21908 Medina (US*, iso), Luzon: Bataan prov., forests back of Limay, on forested slopes, altitude c. 300m, 2 Jul 1914.
  7. Elaeocarpus culminicola Warb., Bot. Jahrb. 16 (1893) 23.
  8. Elaeocarpus cumingii Turcz., Bull. Soc. Nat. Mosc. 19, 2 (1846) 491; --Fern.-Villar, Novis. App. (1880) 31; --Vidal, Phan. Cuming. Philip. (1885) 99; Rev. Pl. Vasc. Filip. (1886) 72; A DC in Elmer, LPB 2 (1909) 637; --Merr., EPFP 3 (1923) 15; --Type: Cuming 807. --Elaeocarpus nitidus Turcz., Bull. Soc. Nat. Mosc. 19, 2 (1846) 491; --Fern.-Villar, Novis. App. (1880) 31, non Jack; --Type: Cuming 893 (L*, iso), Luzon: Albay prov. --Elaeocarpus oblongus Turcz., Bull. Soc. Nat. Mosc. 31, 1 (1858) 235, non Gaertn. --Elaeocarpus versicolor Elmer, LPB 4 (1911) 1178; --Type: Elmer 12761(BISH*, GH*, NA*, US*, iso), Palawan: Palawan prov., Puerto Princesa, Mt Pulgar (=Thumb Peak), Mar 1911. Philippines, Sulawesi. LUZON: Nueva Vizcaya, Bataan, Laguna, Quezon, Camarines, Albay, Sorsogon, PALAWAN, CEBU, SAMAR, BASILAN, MINDANAO. Forests at low altitudes. Photos
  9. Elaeocarpus curranii Merr., PJS 4 c (1909) Bot. 288; EPFP 3 (1923) 16; --Type: FB 4 Ware (US*, iso). LUZON: Ilocos Norte (Pauquin), Cagayan, Zambales, Bataan, Pampanga, Quezon, Camarines. Low altitude forests.
  10. Elaeocarpus dinagatensis Merr., J. Arnold Arb. 32 (1951) 162; --Type: BS 35117 Ramos & Pascasio (A, holo?; iso in K, US*), Bucas Grande, June 1919. DINAGAT; BUCAS GRANDE (type locality). Low altitude. Both islands have predominantly ultramafic geology. Fr. May-June
  11. Elaeocarpus floribundus Blume, Bijdr. (1825) 120; --Merr., J. Arnold Arb. 32 (1951) 162. India, Myanmar, Thailand, Indochina, Peninsular Malaysia, Java, Borneo, SW Philippines. PALAWAN: (Taytay; Aborlan). Forests at low altitudes.
  12. Elaeocarpus forbesii Merr., PJS 12 c (1917) Bot. 282; EPFP 3 (1923) 16. LUZON: Benguet, MINDORO: Mindoro Oriental (Subaan River headwaters inland from San Teodoro, Coode 5663 with Ridsdale). Forested ravines and ridges, altitude 1100- 1500m.
  13. Elaeocarpus foxworthyi Merr., PJS 3 c (1908) Bot. 244; EPFP 3 (1923) 16; --Type: BS 858 Foxworthy. PALAWAN. Forests along streams at low altitudes.
  14. Elaeocarpus fulvus Elmer, LPB 4 (1911) 1179; --Merr., EPFP 3 (1923) 16; --Type: Elmer 10825. MINDANAO: Davao. Primary forests, altitude c. 1100m.
  15. Elaeocarpus gammillii Knuth, Repert. Sp. Nov. 50 (1841) 83; --Type: Gamill 300 (A*, iso), Guimaras. Feb. 1904. GUIMARAS
  16. Elaeocarpus gigantifolius Elmer, LPB 4 (1911) 1182; --Merr., EPFP 3 (1923) 17; --Type: Elmer 11184 (PNH, holo, lost; iso in K, L), Mindanao: Davao, Mt Apo, Todaya. LEYTE; MINDANAO: Davao (Mt Apo). Known with certainty from the type only. More material is needed to decide whether the MINDANAO and LEYTE specimens are all part of the same but variable species and whether subspecific status is justified for those from LEYTE (Coode & Weibel, 1994). Steep slopes in forest, 750m. 
  17. Elaeocarpus glaber Blume, Bijdr. (1825) 122; --Merr., J. Arnold Arb. 32 (1951) 164. Extra-Philippine range? MINDANAO: Zamboanga prov., Sax River.
  18. Elaeocarpus grandiflorus Sm, Rees Cyclop. 12 (1809) no. 5; --Merr., EPFP 3 (1923) 17. Myanmar, Indochina, Java, Philippines. BUSUANGA. Lowland forests along streams.
  19. Elaeocarpus halconensis Merr., PJS 26 (1925) 470; EPFP 4 (1925) 249; J. Arnold Arb. 32 (1951) 165; --Type: BS 40583 Ramos & Edaño, Mindoro: Mindoro Oriental prov., Mt Halcon.
  20. Elaeocarpus ilocanus Merr., PJS 14 (1919) 418; EPFP 3 (1923) 17; --Type: BS 33271 Ramos. LUZON: Ilocos Norte. Forested slopes, altitude c. 1000m
  21. Elaeocarpus luzonicus Merr., PJS 4 c (1909) Bot. 17; EPFP 3 (1923) 17; --Type: FB 10488 (US*, iso), Luzon: Camarines Sur prov., Mt Isarog LUZON: Camarines Sur, Sorsogon, MINDORO, PANAY, CAMIGUIN. Forested slopes, alt. c. 1000m.
  22. Elaeocarpus macranthus Merr., Publ. Govt. Lab. (Philipp.) 35 (1906) 38; EPFP 3 (1923) 17; --Type: Merrill 2298. --Elaeocarpus apoensis Elmer., LPB 4 (1911) 1185; --Merr., EPFP 3 (1923) 14; --Type: Elmer 10524. LUZON: Benguet, Rizal, Laguna, Quezon, Camarines, PANAY, LEYTE, MINDANAO: Davao (Mt Apo). Primary forests, altitude 200-1250m.
  23. Elaeocarpus mandiae Coode, Kew Bull. 56 (2001) 896. MINDORO: Mindoro Oriental (Mt Halcon).
  24. Elaeocarpus merrittii Merr., PJS 2 c (1907) 280; EPFP 3 (1923) 17; --Type: Merrill 5582. --Elaeocarpus urophyllus Merr. & Quisumb., J. Arnold Arb. 32 (1951) 171; --Type: PNH 6832, Sulit. Samar:   Eastern Samar prov., Mt Calbiga near Wright. LUZON: Quirino (Mt Alzapan, BS 45617 Ramos & Edano), MINDORO: Mindoro Oriental (Mt Halcon), SAMAR: Eastern Samar (Mt Calbiga). Primary forests, alt. (300-) 900-1500m.
  25. Elaeocarpus mindanaensis Merr., PJS 3 c (1908) Bot. 150; EPFP 3 (1923) 18; --Type: Clemens 910  MINDANAO: Lanao, Bukidnon. Forests, altitude c. 750m; according to Merrill (1923) allied to the Bornean E. gambir Becc.
  26. Elaeocarpus mindorensis Knuth, Repert. Sp. Nov. 50 (1941) 86. --Type: BS 39814 Ramos. MINDORO: Mindoro Occidental (Paluan). Only known from the type collection.
  27. Elaeocarpus mollis Merr., PJS 9 c (1914) Bot. 373; EPFP 3 (1923) 18; --Type: Wenzel 698 (A*, US*, iso). Leyte: Leyte prov., Buenavista near Jaro, 20 April 1914. LEYTE. Primary forests, altitude 60-500m.
  28. Elaeocarpus monocerus Cav., Icon. 6 (1801) 1, t. 501; DC, Prodr. 1 (1824) 519; --Fern.-Villar,   Novis. App. (1880) 31; --Merr., PJS 1 (1906) Suppl. 90; --A DC in Elmer, LPB 2 (1909) 637; --Merr.,   PJS 13 c (1918) Bot. 29; EPFP 3 (1923) 18; --Type:  Elaeocarpus megacarpus Elmer, LPB 7 (1915) 2627; --Type: Elmer 14059 (BISH*, L*, MO*, NY*, US*, iso).   Mindanao: Agusan del Norte prov., Cabadbaran, Mt Urdaneta (=Hilong-hilong), Oct 1912. LUZON: Ilocos Sur, Cagayan (Aparri), Isabela, Bataan, Rizal, Laguna (Mt Makiling; San Antonio), Quezon, Camarines, SAMAR: Northern Samar (Palapag), MINDANAO: Zamboanga, Agusan del Norte. Primary forests, ascending to 1000m.
  29. Elaeocarpus multiflorus (Turcz.) Fern.-Villar, Novis. App. Fl. Filip. 31 (1880) 31; --Merr., EPFP 3   (1923) 17; --Type: Cuming 873 Elaeocarpus leytensis Merr., PJS 8 c (1913) Bot. 383; EPFP 3 (1923) 17; --Type: Wenzel 212. Sulawesi (?) fide Koorders, Philippines. LUZON: Quezon, Camarines, Albay, Sorsogon, MINDORO, SIBUYAN, LEYTE, CAMIGUIN, MINDANAO. Low and medium altitude primary forests.
  30. Elaeocarpus nervosus Elmer, LPB 4 (1911) 1176; --Merr.,  EPFP 3 (1923) 18; --Type: Elmer 11566  LUZON: Mountain Province, Benguet, MINDANAO: Davao. Forests, alt. 1350-1600m. According to Merrill (1923), closely allied to E. cumingii.
  31. Elaeocarpus octopetalus Merr., PJS 3 c (1908) Bot. 149; EPFP 3 (1923) 18; J. Arnold Arb. 32   (1951) 167, emend., “petalis sepalisque 4, non 8.”  --Type: Clemens 1148  MINDANAO: Lanao del Sur (Camp Keithley). Forests, altitude c. 750m.
  32. Elaeocarpus palimlimensis Knuth, Repert. Sp. Nov. 49 (1940) 68; --Merr., J. Arnold Arb. 32 (1951)   167; --Type: BS 33377 Ramos (A*, US*, iso). Luzon: Ilocos Norte prov., Mt Palimlim,  Aug 1918. LUZON: Ilocos Norte. Only known from the type collection.
  33. Elaeocarpus pedunculatus Wall., List (1832) no. 2678, nomen nudum,  ex Mast. in JD Hook., Fl.   Brit. Ind. (1874) 408, descr.  Elaeocarpus candollei Turcz., Bull. Soc. Nat. Mosc. 192 ; Elmer, LPB 4 (1911) 1185; --Merr., EPFP 3 (1923)   15; --Type: Elmer 12974 (L* 2 sheets, NY* 2 sheets, US*, iso). Palawan: Puerto Princesa, Mt   Pulgar (=Thumb Peak), April 1911. Elaeocarpus nitidulus Knuth, Fedde Repert. 49 (1940) 67; --Type: BS 41234 Ramos (A*, iso). Busuanga   Island, Sept. 1922.  Sumatra, Peninsular Malaysia, Banka, Billiton, Borneo, C Philippines. BUSUANGA, PALAWAN, PANAY, CEBU. Forests, altitude 1000-1300m.
  34. Elaeocarpus samari Weibel, Candollea 35 (1980) 524. --Type: PNH 6398 Sulit (L*, holo; iso in A). Samar: Eastern Samar prov., Wright, Mt Calbiga. April-May 1948. SAMAR.
  35. Elaeocarpus sphaericus (Gaertn.) K Schum. in Engl. & Prantl, Nat. Pflanzenfam. 3, 6 (1890) 5; --Corner,  Gard. Bull. Sing. 10 (1939) 326. --Elaeocarpus subglobosus Merr., PJS 4 c (1909) Bot. 290 ; EPFP 3 (1923) 19. --Elaeocarpus baclayanensis Elmer, LPB 4 (1911) 1188. --Elaeocarpus dolichopetalus Merr., PJS 9 c (1914) Bot. 372; EPFP 3 (1923) 16; --Type: Wenzel 897 (A*, US*, iso), Leyte: Leyte prov., Buenavista near Jaro, in forests, altitude c. 500m, 13 Jun 1914. Elaeocarpus ramiflorus Merr., PJS 10 c (1915) Bot. 43; EPFP 3 (1923) 19; --Type: BS 13115 Ramos (US*,   iso). Luzon: Laguna prov., San Antonio, trail between San Antonio and Piape, in forests, Jun 1912. Hainan. Philippines. LUZON: Mountain Province, Benguet, Laguna, Camarines Sur (Mt Isarog), Sorsogon, PANAY: Capiz (Libacao, BS 35438 Martelino & Edaño), LEYTE, MINDANAO: Davao (Mt Apo). Primary forests, altitude 400-1600m.
  36. Elaeocarpus submonoceras Miq., Fl. Ind. Bat., suppl. 1 (1860) 407.
  37. Elaeocarpus surigaensis Merr., PJS 13 c (1918) Bot. 28; EPFP 3 (1923) 19; --Type: FB 26000 Ponce (US*, iso), Mindanao: Surigao del Norte prov., near Mabini, in level rich soil at the edge of the forest, altitude c. 50m, 11 May 1916. MINDANAO: Surigao del Norte. Lowland forests.
  38. Elaeocarpus venosus C Robinson, Bull. Torr. Bot. Club 35 (1908) 71, 75; --Merr.,  EPFP 3 (1923) 20; --Type: RS Williams 2002. LUZON: Abra, Mountain Province, Benguet. Mossy forests on the higher mountains, altitude 1800-2500m.
  39. Elaeocarpus verruculosus A DC in Elmer, LPB 2 (1909) 635; EPFP 3 (1923) 20; --Syntypes: Elmer 8645 (Elmer 8734). LUZON: Benguet. Thickets or forests, altitude c. 1500m.
  40. Elaeocarpus verticillatus Elmer., LPB 4 (1911) 1181; --Merr., EPFP 3 (1923) 20; --Type: Elmer 12104 (PNH, holo, lost; iso: L* 2 sheets, MO*, NY*, US*), Sibuyan: Mt Giting-giting, Magallanes (=Magdiwang), along wooded streams at 750ft, March 1910.  Syntypes: Elmer 12427 (L*, MO*, NY*, US*), Sibuyan: Mt Giting-giting, Magallanes (=Magdiwang), Pauala River, alt. 2000ft, May 1910. Photos

Doubtful:

  1. Elaeocarpus stipularis Blume, Bijdr. (1825) 120 Elaeocarpus fissistipulus Miq., Fl. Ind. Bat. 13 (1858-59) 210; --Merr.,  EPFP 3 (1923) 16.  Based on a Llanos specimen in de Candolle’s herbarium not seen by Merrill,

Sloanea L.

Photos
  1. Sloanea javanica (Miq.) Szyszyl. ex K Schum. in Engl. & Prantl, Nat. Pflanzenfam. 36 (1890) 5; --Merr., EPFP 3  (1923) 21. Peninsular Malaysia, Java, Philippines, ?Sulawesi. LUZON: Cagayan, Isabela, Laguna, MINDORO, MINDANAO: Lanao. Low altitude primary forests. Photos
  2. Sloanea sigun (Blume) K. Schum. in Engl. & Prantl, Nat. Pflanzenfam. 3, 6 (1890) 5; --Coode, Kew Bull. 38 (1983) 389. India (Assam), Thailand, Sumatra, Peninsular Malaysia, Java, Borneo (Sabah) and the Philippines (widespread). Lowland primary forests. 

Literature

Coode, MJE. 1983. A conspectus of Sloanea (Elaeocarpaceae) of the Old World. Kew Bulletin 38: 347-427.

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Last updated September 22, 2013